taxonID	type	description	language	source
03B82C73FFB401233282FA11FAC4F690.taxon	diagnosis	Diagnosis. Body size 1162 – 1294 × 763 – 863. Rostral and anterior exobothridial setae phylliform, serrate; lamellar, interlamellar, posterior exobothridial and notogastral setae setiform, smooth. Bothridial setae setiform, slightly serrate unilaterally. Notogastral transverse bands complete. Epimeral setae 1 b, 3 a, 3 b, 4 a, 4 b setiform; others phylliform. Genital setae similar in length.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFB401233282FA11FAC4F690.taxon	description	Description. Measurements. Very large species. Body length 1228 (holotype: female), 1162 – 1294 (three paratypes: all females); body width 780 (holotype), 763 – 863 (four paratypes). Integument. Body color yellow-brownish to red-brownish. Surface of body foveolate; foveolae very small (less than 1, distinctly visible only under high magnification). Porose areas larger (up to 6) than foveolae, forming transverse bands on notogaster. Prodorsum. Rostrum widely rounded. Rostral (ro, 116 – 132) and anterior exobothridial (exa, 99) setae phylliform, dilated in medial part, weakly serrate. Lamellar (le), interlamellar (in) and posterior exobothridial (exp) setae similar in length (282 – 365), setiform, smooth. Bothridial setae (ss, 232 – 249) setiform, thickened, slightly serrate unilaterally. Postbothridial transverse band (S b) well visible. Notogaster. Anterior notogastral margin weakly convex medially. Nine transverse bands (S 1 – S 9) distinct, all complete. Sixteen pairs of notogastral setae long (282 – 332, posterior setae in some specimens up to 398), setiform, smooth. Gnathosoma. Subcapitulum with one pair of lateral tubercles. Subcapitulum longer than wide (298 – 332 × 282). Subcapitular setae similar in length (77 – 82), smooth; h, m 1, m 2 phylliform, a setiform. Three pairs of adoral setae present: or 1 (45 – 49) wide, lobe-formed, weakly concave distally, smooth; or 2 (69 – 73) setiform, thickened, with strong spine in distal part; or 3 (53 – 61) setiform, thickened, smooth. Palps (164) with setation 0 – 1 – 0 – 1 – 10 (+ ω). Distal three setae connected basally. Solenidion little longer than palptarsus, thick, not fused with acm. Chelicerae (323 – 328) with two cheliceral setae; chb (77 – 82), setiform, smooth, cha (10 – 12) thorn-like. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 3, 4) well visible. Epimeral setal formula: 3 – 1 – 3 – 4. Setae 1 b, 3 a, 3 b, 4 a, 4 b (83) setiform, with one to two barbs; others (66 – 83) phylliform, serrate. Anogenital region. Ten pairs of genital setae (g 1 – g 10, 66 – 83) setiform, with one to two barbs. One pair of anal (an, 66 – 99) and four pairs of adanal (ad 1, 215; ad 2 – ad 4, 149 – 182) setae setiform, smooth. Roman letters refer to normal setae (e to famulus), Greek letters to solenidia, d σ — seta and solenidion coupled. Single prime (') marks setae on anterior and double prime (") setae on posterior side of the given leg segment. Parentheses refer to a pair of setae.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFB401233282FA11FAC4F690.taxon	materials_examined	Type material. Collection data for holotype (female): locality 2 (soil); collection data for four paratypes (all females): locality 6 (soil). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; three paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFB401233282FA11FAC4F690.taxon	etymology	Etymology. The specific name “ monosetosa ” refers to the one seta on each anal plate.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFB401233282FA11FAC4F690.taxon	discussion	Remarks. The subgenus Lohmannia (Carolohmannia) comprises only the type species, Lohmannia (Carolohmannia) carolensis Norton, Metz & Sharma, 1978, which is distributed in U. S. A. (Norton et al. 1978). Lohmannia (Carolohmannia) monosetosa sp. nov. differs from the latter by the serrate bothridial setae (versus with long branches in L. (C.) carolensis), phylliform rostral setae (versus setiform in L. (C.) carolensis), setiform posterior exobothridial setae (versus phylliform in L. (C.) carolensis), smooth lamellar, interlamellar and notogastral setae (versus serrate in L. (C.) carolensis), complete notogastral transverse bands (versus interrupted medially in L. (C.) carolensis) and genital setae similar in length (versus heterotrichous in L. (C.) carolensis).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFB301393282FB0AFE83F372.taxon	diagnosis	Diagnosis. Body size 481 – 614 × 398 – 448. Rostral and lamellar setae setiform, slightly barbed. Interlamellar setae short, weakly thickened, smooth. Bothridial setae long, thick, weakly dilated in medial part, densely barbed. Spinae adnatae absent. Dorsal notogastral setae thorn-like, slightly barbed, differ in length: h 1, h 3> c 1, c 2> lp> la, lm, h 2. Parastigmatic tubercles Sa triangular, Sp rectangular. Epimeral tubercles E 2 a, E 2 p, Va and Vp present. Leg tibiae with nine setae.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFB301393282FB0AFE83F372.taxon	description	Description. Measurements. Body length 531 (holotype: female), 481 – 614 (three paratypes: one female and two males); body width 415 (holotype), 398 – 448 (three paratypes). Integument. Body color brown to black. Body covered with tuberculate cerotegument (diameter tubercles up to 4). Body surface smooth. Prodorsum. Rostrum narrowly rounded. Rostral (82 – 90) and lamellar (139 – 155) setae setiform, slightly barbed. Interlamellar setae short (16 – 20), weakly thickened, smooth, covered by several granules on their surface. Bothridial setae longest setae on prodorsum (217 – 233), thick, weakly dilated in medial part, densely barbed. Exobothridial setae (32 – 36) thin, slightly barbed. Three pairs of prodorsal tubercles (Da, Ba, Bp) developed, triangular, round-ended. Notogaster. Oval. Anterior notogastral margin convex medially. Spinae adnatae absent. Dorsal notogastral setae inserted in 2 sub-parallel rows, thorn-like, slightly barbed, differ in length: h 1, h 3 (86 – 98) longer than c 1, c 2 (53 – 61), lp (32 – 41), la, lm, h 2 (20 – 28). Posterior setae p 1 – p 3 (65 – 69) setiform, thickened, slightly barbed. Gnathosoma. Subcapitulum longer than wide (143 – 147 × 86 – 90). Subcapitular setae setiform; h (41) barbed, longer than a and m (both 32), slightly barbed. Two pairs of adoral setae (12) setiform, smooth. Palps (114 – 118) with setation 0 – 2 – 1 – 3 – 9 (+ ω). Solenidion shorter than half of palptarsus length, straight, thickened, blunt-ended, not attached to eupathidium. Chelicerae (98 – 106) with two setae, having flagellate tip; cha (61 – 65) barbed, longer than chb (36 – 41) with long cilia in middle part. Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal) weakly visible. Parastigmatic tubercles large: Sa elongate, triangular, blunt-ended; Sp rectangular. Epimeral tubercles E 2 a and E 2 p small, broadly triangular in ventral view. Ventrosejugal tubercles Va and Vp strongly developed, triangular to broadly rounded. Epimeral formula: 3 – 1 – 3 – 4. Setae setiform, slightly barbed; 1 b, 3 b (61 – 73) longer than 3 c, 4 a, 4 b, 4 c, 4 d (45 – 53) and 1 a, 1 c, 2 a, 3 a (28). Setae 3 b and 4 a inserted on Vp. Discidia (dis) well developed, triangular, blunt-ended. Anogenital region. Six pairs of genital (g 1, 53; g 2 – g 6, 32 – 36) and two pairs of anal setae (an 1, an 2, 32 – 36) setiform, smooth. One pair of aggenital (ag) and three pairs of adanal (ad 1 – ad 3) similar in length (41 – 45), setiform, slightly barbed. Adanal lyrifissures (iad) located anteriorly to ad 3. Ovipositor typical for Damaeidae (Ermilov 2010), short (89 × 69); length of lobes 28, length of cylindrical distal part 61. Each lobes with four setae; all thornlike, smooth, similar in length: ψ 1 ≈ τ 1 ≈ ψ 2 ≈ τ a ≈ τ b ≈ τ c (16). Six coronal setae short (6), thin, smooth. Legs. Monodactylous; claws smooth. Lengths of legs and their segments are given in Table 3. Formulae of leg setation and solenidia: I (1 – 7 – 3 – 9 – 21) [1 – 2 – 2], II (1 – 7 – 4 – 9 – 18) [1 – 1 – 2], III (2 – 4 – 3 – 9 – 18) [1 – 1 – 0], IV (1 – 4 – 2 – 9 – 15) [0 – 1 – 0]; homologies of setae and solenidia indicated in Table 4. Leg Trochanter Femur Genu Tibia Tarsus I v' d, (l), (v), v' ', bv' ' d σ, l' ', v' d, l' ', (v 1), v 2 ', (v 3), (v 4), (ft), (tc), (it), (p), (u), (a), s, (pv), (v 1), v 2 ', φ 1, φ 2 (pl), e, ω 1, ω 2 II v' d, (l), (v), v' ', bv' ' d σ, (l), v' d, l' ', (v 1), v 2 ', (v 3), (v 4), φ (ft), (tc), (it), (p), (u), (a), s, (pv), (v 1), v 2 ', ω 1, ω 2 III l', v' d, l', v', ev' d σ, l', v' d, (v 1), (v 2), (v 3), (v 4), φ (ft), (tc), (it), (p), (u), (a), s, (pv), (v 1), v 2 ' IV v' d, l', v', ev' d, v' d, (v 1), (v 2), (v 3), (v 4), φ ft' ', (tc), (p), (u), (a), s, (pv), (v 1), v 2 ' See Table 2 for explanations.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFB301393282FB0AFE83F372.taxon	materials_examined	Type material. Collection data for holotype (female): locality 6 (litter); collection data for three paratypes (one female and two males): locality 6 (litter). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; two paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFB301393282FB0AFE83F372.taxon	etymology	Etymology. The specific name “ heterotrichus ” refers to the heterotrichy of notogastral setae.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFB301393282FB0AFE83F372.taxon	discussion	Remarks. The genus Tectodmaeus comprises about 20 species (see different opinions on classification, for example: Wang & Cui 1994; Subías 2004), which are distributed in the Palaearctic and Oriental regions (data summarized by Subías 2004, updated 2014). In having the characteristic morphology of ventral side of the body, absence of spinae adnatae and the presence of notogastral heterotrichy, Tectodamaeus heterotrichus sp. nov. is most similar to Tectodamaeus brevisetus (Wang, 1994 in Wang & Cui 1994) from southern China. However, it clearly differs from the latter by the smaller body size (481 – 614 × 398 – 448 versus 733 – 766 × 479 – 499 in T. brevisetus), rectangular parastigmatic tubercles (versus thorn-like in T. brevisetus), more clear heterotrichy of notogastral setae (h 1, h 3> c 1, c 2> lp> la, lm, h 2 versus c 2> c 1, la, la, lm, lp, h 3> h 2> h 1 in T. brevisetus) and nine setae on all leg tibiae (versus I (7) – II (6) – III (5) – IV (5) in T. brevisetus).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA901343282FE64FD70F0D4.taxon	diagnosis	Diagnosis. Body size 282 – 348 × 166 – 232. Rostrum bidentate. Lateral sides of rostrum with three blunt teeth. Rostral region with one pair of oval foveolae. Tectopedial fields absent. Rostral setae geniculate, densely ciliate. Lamellar and interlamellar setae setiform, sparsely barbed. Bothridial setae spindleform. A pair of humeral tubercles present. Notogastral setae setiform, smooth, with short flagellate tip. Genital and aggenital setae slightly barbed; anal and adanal setae smooth.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA901343282FE64FD70F0D4.taxon	description	Description. Measurements. Body length 315 (holotype: male), 282 – 348 (five paratypes: one female and four males); body width 182 (holotype), 166 – 232 (five paratypes). Integument. Body color light brownish. Body surface smooth. Dorsal and lateral sides of prodorsum with foveolae, forming a polygonate pattern (length of foveolae up to 8). Prodorsum. Rostrum bidentate. Lateral sides of rostrum with three teeth: anterior tooth narrowly triangular, others large tubercle-like, round-ended. Rostral region with one pair of oval foveolae (fov, 6 – 12 × 4 – 8). Tectopedial fields absent. Rostral setae (24 – 36) geniculate, densely ciliate unilaterally. Lamellar setae (49 – 53) setiform, sparsely barbed, inserted on triangular tubercle rounded distally. Interlamellar setae (41 – 49) setiform, sparsely barbed, inserted in basal part of triangular interbothridial tubercles. Bothridial setae (94 – 112) spindleform, with long, thin tip, having barbs. Exobothridial setae (24 – 32) setiform, thin, smooth. A pair of triangular, roundly blunted postbothridial tubercles well visible. Notogaster. Anterior notogastral margin weakly convex medially. A pair of large, rounded humeral tubercles developed, directed towards the postbothridial tubercles. Ten pairs of notogastral setae setiform, smooth, with short flagellate tip: c, la, lm, lp, h 2, h 3 (73 – 90) longer than h 1, p 1, p 2, p 3 (53 – 69). Lyrifissures and opisthonotal gland openings indistinctly visible. Gnathosoma. Subcapitulum longer than wide (73 × 41 – 45). Subcapitular setae setiform; h (28) with long cilia unilaterally, little longer than slightly barbed m and smooth a (both 24). Adoral setae absent. Palps (45 – 49) with setation 0 – 2 – 0 – 2 – 8 (+ ω). Solenidion longer than half of palptarsus length, straight, thickened, pressed to the surface of palptarsus, not attached to eupathidium. Chelicerae (61) with one smooth setae cha (20). Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 4) well visible. Epimeral setal formula 3 – 1 – 3 – 3. All setae setiform, barbed; 1 a, 2 a, 3 a (20) shorter than others (32 – 41). Discidia triangular, blunt-ended. Anogenital region. Six pairs of genital (g 1, 20 – 28, g 2 – g 6, 16 – 20) and one pair of aggenital (26) setae setiform, slightly barbed. Two pairs of anal (24 – 32) and three pairs of adanal (ad 1, ad 2, 51 – 55; ad 3, 45 – 49) setae setiform, smooth. Distance ag – ag shorter than ad 3 – ad 3. Lyrifissures iad short. Legs. Monodactylous; claws smooth. Formulae of leg setation and solenidia: I (1 – 5 – 2 – 4 – 20) [1 – 2 – 2], II (1 – 5 – 2 – 4 – 14) [1 – 1 – 2], III (2 – 3 – 1 – 3 – 13) [1 – 1 – 0], IV (1 – 2 – 2 – 3 – 10) [0 – 1 – 0]; homologies of setae and solenidia indicated in Table 5.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA901343282FE64FD70F0D4.taxon	materials_examined	Type material. Collection data for holotype (male): locality 3 (soil); collection data for five paratypes (one female and four males): locality 6 (soil). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; two paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; three paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA901343282FE64FD70F0D4.taxon	etymology	Etymology. The specific name “ vietnamensis ” refers to the country of origin, Vietnam. See Table 2 for explanations.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA901343282FE64FD70F0D4.taxon	discussion	Remarks. The genus Allosuctobelba comprises 14 species, which are distributed in the Holarctic, Oriental and Neotropical regions (data summarized by Subías 2004, updated 2014). In having the spindle-form bothridial setae, absence of tectopedial fields, presence of humeral tubercles and notogastral setae with short flagellate tip, Allosuctobelba vietnamensis sp. nov. is similar to Allosuctobelba tricuspidata tricuspidata Aoki, 1984 (b) (see also Chinone 2003) from Japan. However, it clearly differs from the latter by the bidentate rostrum (versus tridentate in A. tricuspidata tricuspidata) and rostrum with lateral teeth (versus without teeth in A. tricuspidata tricuspidata).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA401333282FCD4FD22F684.taxon	diagnosis	Diagnosis. Body size 332 – 365 × 182 – 215. Body covered with granular cerotegument. Two pairs of subcostular ridges straight, longitudinally oriented. Rostral setae setiform, thin, slightly barbed; lamellar setae setiform, thickened, ciliate; interlamellar setae dilated medio-distally, with strong barbs. Bothridial setae clavate, with strong cilia. Anterior part of notogaster with two short ridges. Ten pairs of notogastral setae of medium size, dilated medio-distally, with strong, dense barbs. Monodactylous.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA401333282FCD4FD22F684.taxon	description	Description. Measurements. Body length 348 (holotype: female), 332 – 365 (five paratypes: three females and two males); body width 199 (holotype), 182 – 215 (five paratypes). Integument. Body color light brownish. Body covered by with granular cerotegument; granules rounded (their diameter up to 4). Body surface smooth. Prodorsum. Rostrum widely rounded. Costulae and two pairs of subcostular ridges (r 1, r 2) long, well developed, longitudinally oriented, parallel. Rostral setae (16 – 20) setiform, thin, slightly barbed. Lamellar setae (45 – 49) setiform, thickened, ciliate unilaterally, inserted on tubercle nearly to costular end. Interlamellar setae (12 – 16) dilated medio-distally, with strong, dense barbs. Bothridial setae (49 – 57) clavate, with strong cilia. Interbothridial tubercles rounded, located separately. Exobothridial setae (8 – 10) setiform, straight, thin, smooth. Pair of small, round-blunted postbothridial tubercles weakly developed. Notogaster. Anterior notogastral margin straight. Pair of large triangular humeral tubercles developed, directed towards the postbothridial tubercles. Anterior part of notogaster with two short longitudinal sclerotized ridges. Ten pairs of notogastral setae dilated medio-distally, with strong, dense barbs: p 1, p 2, p 3 (12 – 20) shorter than others (24 – 28). Lyrifissures and opisthonotal gland openings distinctly visible. Gnathosoma. Subcapitulum longer than wide (77 – 82 × 61). Subcapitular setae setiform, smooth; h (16) longer than a 12) and m (6). Adoral setae absent. Palps (49 – 53) with setation 0 – 2 – 1 – 1 – 8 (+ ω). Solenidion shorter than half of palptarsus length, straight, thickened, blunt-ended, pressed to surface of palptarsus, not attached to eupathidium. Chelicerae (77 – 82) with two setiform, slightly barbed setae; cha (20 – 22) longer than chb (16 – 18). Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 4) well visible. Epimeral setal formula 3 – 1 – 3 – 3. All setae setiform, thin, smooth; 1 a, 1 c, 2 a, 3 a, 4 a, 4 b, 4 c (12 – 16) shorter than 3 b, 3 c (16 – 24) and 1 b (26 – 32). Discidia triangular, widely rounded. Anogenital region. Six pairs of genital (g 1, 12, g 2 – g 6, 6 – 8), one pair of aggenital (4), two pairs of anal (8 – 10) and three pairs of adanal (ad 1, ad 2, 16 – 20; ad 3, 4 – 6) setae setiform, smooth. Lyrifissures iad in inverse apoanal position. Setae ad 3 inserted posteriorly to iad. See Table 2 for explanations.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA401333282FCD4FD22F684.taxon	materials_examined	Type material. Collection data for holotype (female): locality 1 (litter); collection data for five paratypes (three females and two males): locality 2 (litter). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; four paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA401333282FCD4FD22F684.taxon	etymology	Etymology. The specific name “ bidupensis ” refers to the Vietnamese park origin, Bi Dup — Nui Ba National Park.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA401333282FCD4FD22F684.taxon	discussion	Remarks. The genus Epieremulus comprises 10 species, having a cosmopolitan distribution except Antarctic region (data summarized by Subías 2004, updated 2014). In having the two short notogastral ridges, absence of transcostula and localization and morphology of ventral setae, Epieremulus bidupensis sp. nov. is most similar to Epieremulus granulatus granulatus (Balogh & Mahunka, 1979) from the Neotropical region. However, it clearly differs from the latter by the monodactylous legs (versus tridactylous in E. granulatus granulatus), setiform lamellar setae (versus thick in E. granulatus granulatus), smooth adanal setae (versus barbed in E. granulatus granulatus) and roundly blunted interbothridial tubercles (versus rectangular in E. granulatus granulatus).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA301323282FAF9FD4AF5BE.taxon	diagnosis	Diagnosis. Body size 614 – 863 × 232 – 365. Body covered with sparse granules. Interlamellar setae longer than rostral and lamellar setae. Bothridial setae with four to six long, strong branches (usually bifurcate distally) and one to four small spines unilaterally. All prodorsal and notogastral condyles present. Notogastral setae long, barbed. Longitudinal short ridge present laterally to setae c and on apodemes IV. Genital plates smooth. Adanal setae ad 3 in preanal position. Lyrifissures iad in paraanal position. Leg setae u thorn-like on all tarsi.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA301323282FAF9FD4AF5BE.taxon	description	Description. Measurements. Body length 796 (holotype: female), 796 – 863 (three paratypes: females), 614, 647 (two paratypes: males); body width 315 (holotype), 315 – 365 (three paratypes: females), 232 (two paratypes: males). Integument. Body color yellow-brownish. Body covered with granulate cerotegument; granules conical, sparsely located (up to 1). Body surface densely microfoveolate (diameter of foveolae less than 1) and with larger sparse foveolae (their diameter up to 6), but foveolae visible only at high magnification in dissected specimens. Lateral regions of body near pedotecta II tuberculate (diameter of tubercles up to 6). Prodorsum. Rostrum widely rounded. Rostral and lamellar setae (both 61 – 82) setiform, densely ciliate unilaterally. Interlamellar setae (77 – 123) setiform, straight, barbed, inserted dorso-medially to bothridial openings. Bothridial setae (98 – 123) with weakly developed head, having four to six long, strong branches (usually bifurcate distally) and one to four small spines unilaterally. Exobothridial setae (32 – 49) setiform, barbed. Medial prodorsal condyles triangular, rounded distally, located separately. Lateral prodorsal condyles elongate, triangular, rounded distally, contacting with lateral notogastral condyles. Notogaster. Anterior notogastral margin straight, weakly developed. Medial notogastral condyles rectangular, rounded distally, located separately. Lateral notogastral condyles triangular, rounded distally. Ten pairs of long, thickened, barbed notogastral setae: c, la (77 – 123), h 1, p 1 – p 3 (90 – 118), others (127 – 143). Longitudinal short ridge (r) present laterally to setae c (sometimes similar ridges also present laterally to la). Lyrifissures im and opisthonotal gland openings located postero-laterally to la. Gnathosoma. Subcapitulum longer than wide (155 – 180 × 106 – 123). Subcapitular setae setiform, smooth; m and h (both 41 – 49) longer than a (16 – 24). Adoral setae absent. Palps (86) with setation 0 – 2 – 1 – 3 – 8 (+ ω). Solenidion longer than half of palptarsus length, straight, thickened, pressed to the surface of palptarsus, distal part attached to one distal seta. Chelicerae (155 – 180) with two barbed setae; cha (41 – 53) longer than chb (20 – 28). Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 4) well visible (3 absent). Longitudinal short ridge present on each apodeme IV. Epimeral setal formula: 3 – 1 – 3 – 3. Setae 1 b, 3 b, 3 c, 4 a (45 – 53) longer than 4 c (36 – 41), 1 c (24 – 28), 1 a, 2 a, 3 a, 4 b (20 – 24). Setae 1 a, 2 a, 3 a, 4 b smooth, others slightly barbed. Setae 3 c inserted on pedotecta II; 4 c inserted on discidia. Discidia triangular, blunted. Anogenital region. Four pairs of genital (20 – 28) and one pair of aggenital (69 – 77) setae setiform, smooth. Two pairs of anal (69 – 77) and three pairs of adanal (ad 1, ad 2, 102 – 110; ad 3, 69 – 77) setae setiform, barbed. Adanal setae ad 3 in preanal position. Lyrifissures iad short, in paraanal position. Ovipositor elongate (212 – 245 × 53 – 61); length of lobes 73 – 86, length of cylindrical distal part 139 – 159. Each lobes with four thin, smooth setae: ψ 1 ≈ τ 1 (45 – 49) longer than ψ 2 ≈ τ a ≈ τ b ≈ τ c (28). Coronal setae absent. Legs. Monodactylous; claws smooth. Tarsi without teeth. Formulae of leg setation and solenidia: I (1 – 4 – 3 – 4 – 16) [1 – 2 – 2], II (1 – 4 – 3 – 3 – 15) [1 – 1 – 2], III (2 – 3 – 1 – 2 – 15) [1 – 1 – 0], IV (1 – 2 – 2 – 2 – 12) [0 – 1 – 0]; homologies of setae and solenidia indicated in Table 7. Leg setae u thorn-like (S-type) on all tarsi. See Table 2 for explanations.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA301323282FAF9FD4AF5BE.taxon	materials_examined	Type material. Collection data for holotype (female): locality 6 (soil); collection data for five paratypes (three females and two males): locality 6 (litter). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; four paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA301323282FAF9FD4AF5BE.taxon	etymology	Etymology. The specific name “ insolitus ” refers to the unusual morphology of bothridial setae.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFA301323282FAF9FD4AF5BE.taxon	discussion	Remarks. The genus Dolicheremaeus comprises 175 species, which are distributed in the Pantropical and Subtropical regions (data summarized by Subías 2004, updated 2014). In having the bothridial setae with long branches and presence of all prodorsal and notogastral condyles, Dolicheremaeus insolitus sp. nov. is similar to Dolicheremaeus pectinatus Balogh, 1970 from Sri Lanka. However, it clearly differs from the latter by the morphology of setae u on tarsi (S – S – S – S versus L – S – S – S in D. pectinatus), bifurcate branches on bothridial setae (versus not furcate in D. pectinatus) and presence of notogastral ridges laterally to setae c (versus absent in D. pectinatus).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF9E010E3282FF65FE26F5C4.taxon	diagnosis	Diagnosis. Body size 481 – 581 × 182 – 232. Tip of costulae simple. Rostral and lamellar setae setiform, indistinctly serrate. Interlamellar, notogastral and ano-adanal setae dilated in medial part, barbed. All medial and lateral condyles present, triangular. Epimeral setal formula: 2 – 0 – 2 – 3. Genital plates with some longitudinal stria. Adanal setae ad 3 in paraanal position. Lyrifissures iad in lateral position. Leg setae u setiform on all tarsi.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF9E010E3282FF65FE26F5C4.taxon	description	Description. Measurements. Body length 564 (holotype: female), 481, 581 (two paratypes: female and male); body width 215 (holotype), 182, 232 (two paratypes). Integument. Body color yellow-brownish. Body surface densely microfoveolate (diameter of foveolae less than 1) and with larger sparse foveolae (their diameter up to 4), but foveolae visible only at high magnification in dissected specimens. Genital plates with some longitudinal stria. Prodorsum. Rostrum widely rounded. Rostral (49 – 61) and lamellar (65 – 73) setae setiform, indistinctly serrate (visible under high magnification). Interlamellar setae (20 – 28) dilated in medial part, barbed, inserted dorsomedially to bothridial openings. Bothridial setae (57 – 65) with long stalk and small, barbed, distally rounded head. Exobothridial setae (6 – 8) setiform, smooth. Medial and lateral prodorsal condyles triangular, rounded distally, located separately. Notogaster. Anterior notogastral margin straight, well developed. Medial and lateral notogastral condyles triangular, rounded distally. Ten pairs of short (24 – 32), dilated in medial part, barbed notogastral setae present. Lyrifissures im and opisthonotal gland openings located postero-laterally to la. Gnathosoma. Subcapitulum longer than wide (102 – 110 × 90). Subcapitular setae setiform, indistinctly serrate; m (36 – 41) longer than a and h (both 28 – 32). Adoral setae absent. Palps (73) with setation 0 – 2 – 1 – 3 – 8 (+ ω). Solenidion longer than half of palptarsus length, straight, thickened, pressed to the surface of palptarsus, distal part attached to distal seta. Chelicerae (123 – 135) each with one barbed seta cha (41 – 45); chb and their alveoli absent. Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 3, 4) well visible. Anterior epimeral condyles triangular, posterior epimeral condyles elongate, rounded distally. Epimeral setal formula: 2 – 0 – 2 – 3. All setae setiform, thin, slightly barbed. Setae 1 a, 2 a, 3 a and their alveoli absent. Setae 4 a, 4 b, 4 c (16 – 20) shorter than others (28 – 36). Setae 3 c inserted on pedotecta II; 4 c inserted close to discidia. Discidia triangular, blunted. Anogenital region. Four pairs of genital (8 – 10) and one pair of aggenital (6 – 8) setae setiform, smooth. Two pairs of anal (16 – 20) and three pairs of adanal (20 – 28) setae dilated in medial part, barbed. Adanal setae ad 3 located in paraanal position. Lyrifissures iad short, located in lateral position, antero-laterally to setae ad 3. Legs. Monodactylous; claws smooth. Tarsi without teeth. Formulae of leg setation and solenidia: I (1 – 4 – 3 – 4 – 16) [1 – 2 – 2], II (1 – 4 – 3 – 3 – 15) [1 – 1 – 2], III (2 – 3 – 1 – 2 – 15) [1 – 1 – 0], IV (1 – 2 – 2 – 2 – 12) [0 – 1 – 0]; homologies of setae and solenidia indicated in Table 7. Leg setae u setiform (L-type) on all tarsi.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF9E010E3282FF65FE26F5C4.taxon	materials_examined	Type material. Collection data for holotype (female): locality 2 (litter); collection data for two paratypes (one female and one male): locality 2 (litter). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; one paratype is deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF9E010E3282FF65FE26F5C4.taxon	etymology	Etymology. The specific name is dedicated to our colleague, the soil zoologist, Dr. Bella R. Striganova (A. N. Severtsov Institute of Problems of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF9E010E3282FF65FE26F5C4.taxon	discussion	Remarks. The genus Fissicepheus comprises 18 species and two subspecies, which are distributed in the Palaearctic and Oriental regions (data summarized by Subías 2004, updated 2014). In combination of interlamellar and notogastral setae dilated in medial part, presence of 10 pairs of notogastral setae and all prodorsal and notogastral condyles, Fissicepheus striganovae sp. nov. is similar to Fissicepheus coronarius coronarius Aoki, 1967 from the Palaearctic region and Taiwan. However, it clearly differs from the latter by the costulae with rounded tip (versus with dentate expansion in F. coronarius coronarius), ano-adanal setae dilated in medial part (versus simple in F. coronarius coronarius) and striate genital plates (versus smooth in F. coronarius coronarius).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF9A01013282FF65FC22F3BF.taxon	diagnosis	Diagnosis. Body size 298 – 332 × 199 – 249. Lamellar setae longer than rostral setae, all setiform, smooth. Interlamellar setae short, thickened, barbed. Bothridial setae setiform, thickened, ciliate unilaterally. Ten pairs of notogastral setae present (centrodorsal setae absent): c, h 1, p 1 – p 3 short, others of medium size (16 – 20); all barbed. Epimeral setae 1 c, 3 c and their alveoli absent; setae 1 a, 2 a represented by alveoli; other setae setiform, smooth. Genito-aggenital and ano-adanal setae simple, smooth.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF9A01013282FF65FC22F3BF.taxon	description	Description. Measurements. Body length 298 (holotype: female), 298 – 332 (seven paratypes: five females and two males); body width 199 (holotype), 199 – 249 (seven paratypes). Integument. Body color light brown. Body surface smooth. Dorso-lateral sides of notogaster with long striate longitudinal band. Prodorsum. Rostrum rounded. Lamellae well separated, protruding in medio-distal part. Translamella thin, distinct. Rostral (12 – 14) and lamellar (6 – 8) setae setiform, thin, smooth. Interlamellar setae (6 – 8) thickened, barbed. Bothridial setae (69 – 77) setiform, thickened, ciliate unilaterally; distal cilia form weakly visible head in some specimens. Exobothridial setae and their alveoli absent. Tutoria (tu) long, widely knife-like. Notogaster. Anterior notogastral margin well developed, convex medially. Lenticulus (len) large, inverse trapezoid, distinctly visible. Humeral regions with one small, weakly developed concavity. Ten pairs of notogastral setae present: c, h 1, p 1 – p 3 short (4 – 6), thickened, barbed; others of medium size (16 – 20), setiform, barbed. Gnathosoma. Subcapitulum longer than wide (77 – 82 × 61). Subcapitular setae setiform, smooth; a and m (both 12 – 16) longer than h (8 – 10). Two pairs of adoral setae (8 – 10) setiform, smooth. Palps (61) with setation 0 – 2 – 1 – 3 – 8 (+ ω). Solenidion little longer than half of palptarsus length, straight, thickened, not attached to eupathidium, inserted in distal part of palptarsus. Chelicerae (77 – 82) with two barbed setae; cha (24 – 28) longer than chb (16 – 18). Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 3, 4) well visible. Epimeral setal formula: 2 – 1 – 2 – 3. Setae 1 c, 3 c and their alveoli absent; setae 1 a, 2 a represented by alveoli; other setae (20 – 24) setiform, thin, smooth. Setae 4 a, 4 b inserted antero-medially to triangular tubercle. Discidia triangular. Anogenital region. Six pairs of genital (g 1 – g 4, 8 – 10; g 5, g 6, 6 – 8), one pair of aggenital (6 – 8), two pairs of anal (6 – 8) and three pairs of adanal (6 – 8) setae setiform, smooth. Lyrifissures iad long. Legs. Monodactylous; claws smooth. Formulae of leg setation and solenidia: I (1 – 5 – 2 – 4 – 18) [1 – 2 – 2], II (1 – 5 – 2 – 4 – 15) [1 – 1 – 2], III (2 – 3 – 1 – 3 – 15) [1 – 1 – 0], IV (1 – 2 – 2 – 3 – 12) [0 – 1 – 0]; homologies of setae and solenidia indicated in Table 8. See Table 2 for explanations.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF9A01013282FF65FC22F3BF.taxon	materials_examined	Type material. Collection data for holotype (female): locality 1 (litter); collection data for seven paratypes (five females and two males): locality 1 (litter). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; five paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; two paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF9A01013282FF65FC22F3BF.taxon	etymology	Etymology. The specific name is dedicated to our colleague, acarologist, Dr. Pablo Martinez (Universidad Nacional de Mar del Plata, Mar del Plata, Argentina).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF9A01013282FF65FC22F3BF.taxon	discussion	Remarks. The genus Tegeocranellus comprises 17 species, having a cosmopolitan distribution except Antarctic region (data summarized by Subías 2004, updated 2014). In the absence of centrodorsal notogastral setae and the morphology of notogastral setae (c, h 1, p 1 – p 3 short; others of medium size), Tegeocranellus martinezi sp. nov. is most similar to Tegeocranellus kethleyi Behan- Pelletier, 1997 from U. S. A. However, it clearly differs from the latter by the presence of notogastral striate bands (versus absent in T. kethleyi), pedotecta I without teeth (versus with tooth or teeth in T. kethleyi) and lamellar setae clearly shorter than rostral setae (versus similar in length in T. kethleyi).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF91011F3282FE33FB9BF0D4.taxon	diagnosis	Diagnosis. Body size 332 – 381 × 182 – 232. Body surface foveolate, epimeral region with reticulate pattern. Lamellae with lateral point anteriorly. Bothridial setae long, with well developed, dilated unilaterally, pointed, smooth head. Interlamellar setae short. Notogastral setae of medium size, setiform, thin. Notogastral porose areas and sacculi absent. Legs monodactylous.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF91011F3282FE33FB9BF0D4.taxon	description	Description. Measurements. Body length 365 (holotype: female), 332 – 381 (three paratypes: two females and one male); body width 199 (holotype), 182 – 232 (three paratypes). Integument. Body color brown. Body surface distinctly foveolate (diameter of foveolae up to 4). Epimeral region with weakly developed reticulate pattern. Prodorsum. Rostrum rounded. Longitudinal ridge present near rostrum. Lamellae with lateral point anteriorly. Rostral setae (61 – 69) setiform, barbed. Lamellar setae (28 – 36) setiform, smooth. Interlamellar setae short (8 – 12), setiform, with indistinct barbs. Bothridial setae (61 – 69) with long stalk and well developed, dilated unilaterally, pointed, smooth head. Exobothridial setae and their alveoli not found. Tutoria knife-like. Notogaster. Anterior notogastral margin convex medially. Dorsophragmata (D) large, rounded anteriorly. Pteromorphs with lateral point. Ten pairs of notogastral setae of medium size, setiform, smooth: p 1, p 2, p 3 (16 – 20) shorter than others (41 – 49). Notogastral porose areas and sacculi absent. Lyrifissures and opisthonotal gland openings weakly visible. Gnathosoma. Subcapitulum longer than wide (92 × 69). Subcapitular setae setiform, slightly barbed; m (28 – 32) longer than a and h (both 20 – 24). Two pairs of adoral setae (16) setiform, slightly barbed. Palps (65 – 69) with setation 0 – 2 – 1 – 3 – 9 (+ ω). Solenidion shorter than half of palptarsus length, straight, thickened, weakly dilated distally, attached to eupathidium. Chelicerae (98 – 102) with two barbed setae; cha (36) longer than chb (20). Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Genal teeth (gt) rectangular, with tapered tip. Apodemes (1, 2, sejugal, 3) well visible. Epimeral setal formula: 3 – 1 – 3 – 3. Setae 3 c, 4 c represented by alveoli. Setae 1 b (28 – 32) longer than 4 a (24 – 28), 3 a, 3 b (20 – 23) and 1 a, 1 c, 2 a, 4 b (4); all simple, smooth. Custodia and discidia not generated. Circumpedal carinae distinct. Anogenital region. Six pairs of genital (g 1 – g 4, 20 – 24; g 5, g 6, 16), one pair of aggenital (20 – 24), two pairs of anal (16 – 18) and three pairs of adanal (16) setae setiform, smooth. Adanal setae ad 3 inserted laterally to lyrifissures iad. Ovipositor elongate (114 – 130 × 32 – 36); length of lobes 49 – 57, length of cylindrical distal part 65 – 73. Each lobes with four thin, smooth setae: ψ 1 ≈ τ 1 (32) longer than ψ 2 ≈ τ a ≈ τ b ≈ τ c (8 – 12). Coronal setae (k) present (6 – 8), setiform, smooth. Legs. Monodactylous; claws smooth. Formulae of leg setation and solenidia: I (1 – 5 – 2 – 4 – 20) [1 – 2 – 2], II (1 – 5 – 2 – 4 – 15) [1 – 1 – 2], III (2 – 2 – 1 – 2 – 15) [1 – 1 – 0], IV (1 – 2 – 2 – 2 – 12) [0 – 1 – 0]; homologies of setae and solenidia indicated in Table 9. Setae l’ on femora III and tibiae III, IV absent.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF91011F3282FE33FB9BF0D4.taxon	materials_examined	Type material. Collection data for holotype (female): locality 2 (litter); collection data for three paratypes (two females and one male): locality 2 (litter). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; two paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia. See Table 2 for explanations.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF91011F3282FE33FB9BF0D4.taxon	etymology	Etymology. The specific name is dedicated to wife of the first author, Svetlana A. Ermilova, for her constant assistance in scientific research.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF91011F3282FE33FB9BF0D4.taxon	discussion	Remarks. The genus Anachipteria comprises 18 species, which are distributed in the Holarctic, Oriental and Neotropical regions (data summarized by Subías 2004, updated 2014). Anachipteria svetlanae sp. nov. is distinguishable from all known species of Anachipteria by the presence of densely foveolate of body surface (versus without foveolate surface), absence of porose areas and sacculi (versus porose areas or sacculi present) and monodactylous legs (versus tridactylous).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF8F011A3282FCC9FBAEF648.taxon	diagnosis	Diagnosis. Body size 564 – 647 × 348 – 448. Body surface microfoveolate. Prodorsal and adanal setae setiform. Ten pairs of notogastral setae of medium size, setiform, smooth. Three pairs of aggenital setae present. Anal plates with longitudinal ridge.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF8F011A3282FCC9FBAEF648.taxon	description	Description. Measurements. Body length 597 (holotype: female), 564 – 647 (three paratypes: two females and one male); body width 398 (holotype), 348 – 448 (three paratypes). Integument. Body color light brown. Body surface microfoveolate, visible under high magnification in dissected specimens (diameter of foveolae less than 1). Anal plates with longitudinal ridge. Prodorsum. Rostrum widely rounded. Lamellae about 1 / 2 of length of prodorsum. Rostral (73 – 82) and lamellar (106 – 114) setae setiform, barbed. Interlamellar setae (102 – 110) setiform, indistinctly barbed. Bothridial setae (127 – 139) setiform, densely ciliate unilaterally. Exobothridial setae (16 – 20) thin, slightly barbed. Tutoria long, with tooth distally, fused part reaching insertion of rostral setae. Sublamellar lines distinct. Sublamellar porose areas (Al) rounded. Notogaster. Anterior notogastral margin well developed, convex medially. Dorsophragmata of medium size, elongate. Ten pairs of notogastral setae of medium size (c 1, la, 65 – 77; others 45 – 49), setiform, smooth. Four pairs of sacculi elongate, sometimes S 2 triangular. Opisthonotal gland openings located laterally to lp. Gnathosoma. Subcapitulum longer than wide (155 × 110). Subcapitular setae setiform, slightly barbed; m (32 – 41) longer than a and h (24 – 32). Adoral setae not studied. Palps (73) with setation 0 – 2 – 1 – 3 – 9 (+ ω). Solenidion little longer than half of palptarsus length, straight, thickened, weakly dilated distally, attached to eupathidium. Chelicerae (196) with two barbed setae; cha (57) longer than chb (32). Trägårdh’s organ (not illustrated) distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 3) well visible. Epimeral setal formula: 3 – 1 – 3 – 3. All setae setiform, thin, smooth. Setae 1 b (28) longer than others (12 – 16). Discidia triangular, blunted. Circumpedal carinae distinct. Anogenital region. Five pairs of genital (g 1, 12 – 16; g 2 – g 5, 8 – 12), three pairs of aggenital (8 – 12), two pairs of anal (20 – 28) and three pairs of adanal (ad 1, ad 2, 41 – 45, ad 3, 20 – 28) setae setiform, smooth. Adanal setae ad 3 inserted anteriorly to lyrifissures iad. Ovipositor elongate (241 × 57); length of lobes 106, length of cylindrical distal part 135. Each lobe with four thin, smooth setae: ψ 1 ≈ τ 1 (45) longer than ψ 2 ≈ τ a ≈ τ b ≈ τ c (24). Coronal setae absent. Leg Trochanter Femur Genu Tibia Tarsus I v' d, (l), v' ', bv' ' (l), v', σ (l), (v), φ 1, φ 2 (ft), (tc), (it), (p), (u), (a), s, (pv), v', (pl), l' ', e, ω 1, ω 2 II v' d, l' 1, l' 2, v' ', bv' ' (l), v', σ (l), (v), φ (ft), (tc), (it), (p), (u), (a), s, (pv), ω 1, ω 2 III l', v' d, l', ev' l', σ l', (v), φ (ft), (tc), (it), (p), (u), (a), s, (pv) IV v' d, ev' d, l' l', (v), φ ft' ', (tc), (p), (u), (a), s, (pv) See Table 2 for explanations.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF8F011A3282FCC9FBAEF648.taxon	materials_examined	Type material. Collection data for holotype (female): locality 4 (soil); collection data for three paratypes (two females and one male): locality 4 (litter). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; two paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF8F011A3282FCC9FBAEF648.taxon	etymology	Etymology. The specific name “ bicarinatus ” refers to the longitudinal ridges on anal plates.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF8F011A3282FCC9FBAEF648.taxon	discussion	Remarks. The subgenus Indoribates (Indoribates) comprises 10 species, which are distributed in the Australian, Oriental and southern Palaearctic regions (data summarized by Subías 2004, updated 2014). In having the setiform prodorsal and adanal setae and microfoveolate / micropunctate body surface, Indoribates (Indoribates) bicarinatus sp. nov. is most similar to Indoribates (Indoribates) microsetosus Ermilov & Anichkin, 2011 (b) from Vietnam. However, it clearly differs from the latter by the presence of notogastral setae of medium size (versus microsetae in I. (I.) microsetosus), three pairs of aggenital setae (versus one pair in I. (I.) microsetosus) and longitudinal ridge on the each anal plate (versus absent in I. (I.) microsetosus).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF8A01113282FB39FCAEF743.taxon	diagnosis	Diagnosis. Body size 547 – 597 × 365 – 431. Body surface of notogaster and ventral side microfoveolate and densely striate. Rostral setae setiform, ciliate. Lamellar and interlamellar setae flagellate, smooth. Bothridial setae clavate, with barbed head. Notogastral setae smooth: c 1, c 2, da, la, dm, dp of medium size, setiform; lm, lp, h 1 – h 3, p 1 long, flagellate; p 2, p 3 short, thin. Setae lm located posteriorly to la. Epimeral setal formula: 3 – 1 – 3 – 2. Legs tridactylous; lateral claws with ventral tooth.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF8A01113282FB39FCAEF743.taxon	description	Description. Measurements. Body length 581 (holotype: female), 547 – 597 (three paratypes: one female and two males); body width 381 (holotype), 365 – 431 (three paratypes). Integument. Body color brown. Prodorsum covered with granulate cerotegument; granules conical (length up to 2). Body surface of notogaster and ventral side microfoveolate (diameter of foveolae less than 1) and densely striate. Prodorsum. Rostrum widely rounded. Lamellae about 1 / 2 of length of prodorsum. Rostral setae (94 – 104) setiform, ciliate unilaterally. Lamellar and interlamellar setae (both 215 – 233) flagellate, smooth. Bothridial setae (45 – 57) clavate, with head rounded distally, barbed. Exobothridial setae (20) thin, smooth. Tutoria long, with tooth distally, reaching insertion of rostral setae. Sublamellar lines short. Sublamellar porose areas oval (12 × 8). Notogaster. Anterior notogastral margin well developed, weakly convex medially. Dorsophragmata of medium size, elongate. Fourteen pairs of smooth notogastral setae of different length and morphology: c 1, c 2, da, la, dm, dp of medium size (45 – 53), setiform; lm (215 – 233), lp, h 1 – h 3, p 1 (262 – 365) long, flagellate; p 2, p 3 shortest, thin. Setae lm located posteriorly to la. Four pairs of sacculi (Sa, S 1, S 2, S 3) oval. Opisthonotal gland openings and lyrifissures visible only under high magnification in dissected specimens. Gnathosoma. Subcapitulum longer than wide (131 – 147 × 118 – 123). Subcapitular setae setiform; a (24 – 28) slightly barbed, m (32 – 36) barbed, h (24 – 28) smooth. Two pairs of adoral setae 14 – 16) setiform, barbed. Palps (86) with setation 0 – 2 – 1 – 3 – 9 (+ ω). Solenidion shorter than half of palptarsus length, straight, thickened, weakly dilated distally, attached to eupathidium. Chelicerae (147 – 168) with two ciliate setae; cha (41) longer than chb (32 – 36). Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 3) well visible. Epimeral setal formula: 3 – 1 – 3 – 2. All setae (20 – 28) setiform, thin, smooth. Setae 3 c inserted on pedotecta II; 4 c inserted on discidia. Setae 4 b and their alveoli absent. Discidia triangular, blunted. Circumpedal carinae distinct. Anogenital region. Five pairs of genital (g 1, 16 – 20; g 2 – g 5, 12 – 16), one pair of aggenital (20 – 28), two pairs of anal (20 – 28) and three pairs of adanal (20 – 28) setae setiform, smooth. Adanal setae ad 3 inserted anteriorly to lyrifissures iad. Legs. Tridactylous; lateral claws with ventral tooth. Formulae of leg setation and solenidia: I (1 – 5 – 3 – 4 – 20) [1 – 2 – 2], II (1 – 5 – 3 – 4 – 15) [1 – 1 – 2], III (2 – 3 – 1 – 3 – 15) [1 – 1 – 0], IV (1 – 2 – 2 – 3 – 12) [0 – 1 – 0]; homologies of setae and solenidia indicated in Table 10.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF8A01113282FB39FCAEF743.taxon	materials_examined	Type material. Collection data for holotype (female): locality 1 (litter); collection data for three paratypes (female and two males): locality 1 (litter). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; two paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF8A01113282FB39FCAEF743.taxon	etymology	Etymology. The specific name is dedicated to mother of the first author, Tatyana L. Ermilova, for her constant assistance in scientific research.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF8A01113282FB39FCAEF743.taxon	discussion	Remarks. The genus Peloribates comprises 85 species having a cosmopolitan distribution (data summarized by Subías 2004, updated 2014). In having the clavate bothridial setae and flagellate notogastral setae, Peloribates tatyanae sp. nov. is most similar to Peloribates fragilis Hammer, 1967 from New Zealand and Peloribates neonominatus Subías, 2004 (see Mahunka 1983) from Tanzania. However, it clearly differs from the both by the striate body surface (versus without stria in P. fragilis and P. neonominatus), notogastral setae c 1, c 2, da, la, dm, dp, p 2, p 3 of medium size or short (versus all very long, flagellate in P. fragilis and P. neonominatus). Also, in having the heterotrichy of notogastral setae (short or medium size, setiform and long, flagellate) and clavate bothridial setae, Peloribates tatyanae sp. nov. is similar to Topobates berndhauseri (Mahunka, 1978) from Mauritius. However, it clearly differs from the latter by the presence of five pairs of genital setae (versus four pairs in T. berndhauseri), absence of prolamellae (versus present in T. berndhauseri), striate body surface (versus without stria in T. berndhauseri), smooth, flagellate lamellar and interlamellar setae (versus barbed, setiform in T. berndhauseri) and short, setiform adanal setae ad 1 (versus long, flagellate in T. berndhauseri).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF81016C3282FA3FFEE8F79B.taxon	diagnosis	Diagnosis. Body size 332 – 348 × 207 – 232. Prodorsal setae long. Bothridial setae setiform, thickened, weakly dilated in medial part, ciliate. Four pairs of genital setae. Legs monodactylous.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF81016C3282FA3FFEE8F79B.taxon	description	Description. Measurements. Body length 332 (holotype: female), 348 (one paratype: male); body width 207 (holotype), 232 (one paratype). Integument. Body color brown. Body surface smooth. Pteromorphs with weakly developed radiate wrinkles. Prodorsum. Rostrum widely rounded. Lamellae distinct, reaching insertions of lamellar setae. Rostral setae (28 – 36) setiform, ciliate. Lamellar (49 – 53) and interlamellar (77) setae setiform, sparsely barbed. Bothridial setae (98 – 102) longest on prodorsum, setiform, thickened, weakly dilated in medial part, ciliate unilaterally. Exobothridial setae absent. Notogaster. Anterior notogastral margin convex medially. Notogastral setae represented by 10 pairs of alveoli. Four pairs of small sacculi well visible: (Sa and S 3 elongate, S 1 and S 2 oval). Opisthonotal gland openings located laterally to S 1. Gnathosoma. Subcapitulum longer than wide (77 × 69). Subcapitular setae a (12) and h (16) setiform, sparsely barbed; h represented by alveoli. Two pairs of adoral setae (6) setiform, barbed. Palps (45) with setation 0 – 2 – 1 – 3 – 9 (+ ω). Solenidion little longer than half of palptarsus length, straight, thickened, dilated distally, attached to eupathidium. Chelicerae (86) with two barbed setae; cha (28) longer than chb (20). Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 3) well visible. Epimeral setal formula: 3 – 1 – 3 – 3. All setae setiform; 1 a, 1 b, 2 a, 3 a, 3 c barbed, others smooth. Setae 1 b, 3 b, 3 c (12 – 14) longer than 1 a, 2 a, 3 a (8 – 10) and 1 c, 4 a, 4 b, 4 c (6 – 8). Setae 3 c inserted on pedotecta II; 4 c inserted on discidia. Discidia triangular, blunted. Circumpedal carinae distinct. Anogenital region. Four pairs of genital (8), one pair of aggenital (4), two pairs of anal (12 – 16) and three pairs of adanal (ad 1, ad 2, 16 – 20, ad 3, 12) setae setiform, smooth. Adanal setae ad 3 inserted anteriorly to lyrifissures iad. Legs. Monodactylous; claw of each leg smooth. Formulae of leg setation and solenidia: I (1 – 5 – 3 – 4 – 20) [1 – 2 – 2], II (1 – 5 – 3 – 4 – 15) [1 – 1 – 2], III (2 – 2 – 1 – 3 – 13) [1 – 1 – 0], IV (1 – 2 – 2 – 4 – 11) [0 – 0 – 0]; homologies of setae and solenidia indicated in Table 11. Setae l’ on femora III, (it) and on tarsi III and a’ on tarsi IV absent. See Table 2 for explanations.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF81016C3282FA3FFEE8F79B.taxon	materials_examined	Type material. Collection data for holotype (female): locality 4 (soil); collection data for one paratype (male): locality 1 (soil). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF81016C3282FA3FFEE8F79B.taxon	etymology	Etymology. The specific name “ monodactylus ” refers to the monodactylous legs.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FF81016C3282FA3FFEE8F79B.taxon	discussion	Remarks. The subgenus Neoribates (Neoribates) comprises more than 38 species having a cosmopolitan distribution except the Ethiopian and Antarctic regions (data summarized by Subías 2004, updated 2014). In having the monodactylous legs I, Neoribates (Neoribates) monodactylus sp. nov. is similar to Neoribates (Neoribates) spindleformis Ermilov & Anichkin, 2012 (d) from Vietnam. However, it clearly differs from the latter by the smaller body size (332 – 348 × 207 – 232 versus 481 – 498 × 332 – 348 in N. (N.) spindleformis), monodactylous legs II – IV (versus bidactylous in N. (N.) spindleformis) and setiform bothridial setae (versus spindle-form in N. (N.) spindleformis).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFFC016A3282F9FFFC92F353.taxon	diagnosis	Diagnosis. Body size 332 – 348 × 249 – 265. Anterior part of prodorsum with very large apophysis. Prodorsal setae long: rostral setae densely ciliate; lamellar and interlamellar setae barbed. Bothridial setae setiform, ciliate. Anterior notogastral margin not developed. Four pairs of rounded or oval porose areas present. Median pore developed, located between porose areas A 1. Adanal setae ad 3 inserted posteriorly to lyrifissures iad. Postanal porose area oval.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFFC016A3282F9FFFC92F353.taxon	description	Description. Measurements. Body length 332 (holotype: female), 332, 348 (two paratypes: one female and one male); body width 249 (holotype), 249, 265 (two paratypes). Integument. Body color yellowish-brown. Body surface smooth. Pteromorphs with weakly developed radiate wrinkles. Some stria located around of the genital and anal apertures. Prodorsum. Anterior part of prodorsum with very large apophysis (pap). Rostrum widely rounded. Rostral setae (41 – 45) setiform, densely ciliate unilaterally. Lamellar (53 – 61) and interlamellar (82 – 90) setae setiform, barbed. Bothridial setae (127 – 147) longest on prodorsum, setiform, with cilia unilaterally. Exobothridial setae absent. Sublamellar lines (S) distinct, typical for Allogalumna: long, curving backwards. Porose areas Ad oval, transverse, elongate (18 – 20 × 4 – 6). Notogaster. Anterior notogastral margin not developed. Dorsophragmata small. Notogastral setae represented by nine pairs of alveoli (tenth pair located on pteromorphs). Four pairs of porose areas of medium size, circular or oval (Aa, 14 – 20 × 12 – 16; A 1, 10 – 14 × 8 – 12; A 2, 8 – 18 × 10 – 12; A 3, 8 – 14 × 8 – 12), with distinct borders. Alveoli of setae la inserted posteriorly to Aa. Lyrifissures im located between lm and A 1. Opisthonotal gland openings located laterally to A 1. Median pore (mp) present in all specimens, located between porose areas A 1. Gnathosoma. Subcapitulum longer than wide (90 × 82). Subcapitular setae setiform; a (18 – 20) slightly barbed, m (10 – 12) and h (14) smooth. Two pairs of adoral setae (12) setiform, slightly barbed. Palps (69) with setation 0 – 2 – 1 – 3 – 9 (+ ω). Solenidion little longer than half of palptarsus length, straight, thickened, blunt-ended, attached to eupathidium. Chelicerae (102 – 106) with two barbed setae; cha (32) longer than chb (20). Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 3) well visible. Four pairs of thin, smooth epimeral setae observed ventrally; 3 b (16 – 20) longer than 1 a, 4 a, 4 b short (8 – 10). Discidia triangular, pointed. Circumpedal carinae distinct. Anogenital region. Six pairs of genital, one pair of aggenital, two pairs of anal and three pairs of adanal setae little differs in size (10 – 14), thin, smooth. Anterior edge of genital plates with three setae. Adanal setae ad 3 inserted posteriorly to lyrifissures iad. Postanal porose area (Ap) oval (l 2 – 14 × 8 – 10). Legs. Tridactylous; claws smooth. Formulae of leg setation and solenidia: I (1 – 4 – 3 – 4 – 20) [1 – 2 – 2], II (1 – 4 – 3 – 4 – 15) [1 – 1 – 2], III (1 – 2 – 1 – 3 – 15) [1 – 1 – 0], IV (1 – 2 – 2 – 3 – 12) [0 – 1 – 0]; homologies of setae and solenidia indicated in Table 12. Leg Trochanter Femur Genu Tibia Tarsus I v' d, (l), bv' ' (l), v', σ (l), (v), φ 1, φ 2 (ft), (tc), (it), (p), (u), (a), s, (pv), v', (pl), l' ', e, ω 1, ω 2 II v' d, (l), bv' ' (l), v', σ (l), (v), φ (ft), (tc), (it), (p), (u), (a), s, (pv), ω 1, ω 2 III v' d, ev' l', σ l', (v), φ (ft), (tc), (it), (p), (u), (a), s, (pv) IV v' d, ev' d, l' l', (v), φ ft' ', (tc), (p), (u), (a), s, (pv) See Table 2 for explanations.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFFC016A3282F9FFFC92F353.taxon	materials_examined	Type material. Collection data for holotype (female): locality 4 (soil); collection data for: locality 6 (soil). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; one paratype in deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFFC016A3282F9FFFC92F353.taxon	etymology	Etymology. The specific name “ paramachadoi ” refers to the similarity between the new species and the species Acrogalumna machadoi Balogh, 1960.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFFC016A3282F9FFFC92F353.taxon	discussion	Remarks. The genus Allogalumna comprises more than 30 species having a cosmopolitan distribution (data summarized by Subías 2004, updated 2014) collectively. In having the combination of small body size, long rostral, lamellar and interlamellar setae, setiform and ciliate bothridial setae, absence of anterior margin of notogaster and presence of four pairs of rounded porose areas, Allogalumna (Allogalumna) paramachadoi sp. nov. is similar to Acrogalumna machadoi Balogh, 1960 (see also Engelbrecht 1972) from the Ethiopian region and India. However, it clearly differs from the latter by the presence of ciliate rostral setae (versus slightly barbed in Aс. machadoi), prodorsal apophysis (versus absent in Aс. machadoi) and medial pore (versus absent or several pores present in Aс. machadoi) and localization of adanal setae ad 3 posteriorly to lyrifissures iad (versus anteriorly or antero-laterally in Aс. machadoi). In having the combination of setiform and ciliate bothridial setae, absence of anterior margin of notogaster and presence of four pairs of rounded porose areas, Allogalumna (Allogalumna) paramachadoi sp. nov. is similar to Allogalumna (Allogalumna) costata Mahunka, 1996 from Madagaskar. However, it clearly differs from the latter by the smaller body size (332 – 348 × 249 – 265 versus 542 – 598 × 403 – 445 in A. (A.) costata), presence of ciliate rostral setae (versus barbed in A. (A.) costata), long interlamellar setae (versus represented by alveoli in A. (A.) costata), prodorsal apophysis (versus absent in A. (A.) costata), medial pore (versus absent in A. (A.) costata), localization of adanal setae ad 3 posteriorly to lyrifissures iad (versus antero-laterally in A. (A.) costata) and smooth genital and anal plates (versus striate in A. (A.) costata).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFFA01613282FE46FCF3F6B6.taxon	diagnosis	Diagnosis. Body size 431 – 448 × 282 – 348. Prodorsal setae well developed, setiform, barbed. Bothridial setae setiform, ciliate. Lamellar lines straight, reaching insertions of rostral setae. Four pairs of porose areas present, circular or oval. Median pore absent. Medial part of genital plates with longitudinal stria. Postanal porose area oval.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFFA01613282FE46FCF3F6B6.taxon	description	Description. Measurements. Body length 431 (holotype: female), 431 – 448 (three paratypes: two females and one male); body width 298 (holotype), 282 – 348 (three paratypes). Integument. Body color yellowish-brown. Body surface smooth. Pteromorphs with well developed radiate wrinkles. Some stria located posteriorly to the genital aperture. Medial part of genital plates with longitudinal stria. Prodorsum. Rostrum widely rounded. Rostral (36 – 41), lamellar (57 – 61) and interlamellar (69 – 82) setae setiform, barbed. Bothridial setae (98 – 127) longest on prodorsum, setiform, with cilia unilaterally. Exobothridial setae absent. Lamellar (L) and sublamellar lines distinct. Lamellar lines straight, long, reaching insertions of rostral setae. Sublamellar lines typical for Galumna: long, curving backwards. Porose areas Ad oval, transverse elongate (20 – 24 × 8). Notogaster. Anterior notogastral margin developed. Dorsophragmata very small. Notogastral setae represented by ten pairs (additionally c x present) of alveoli (eleventh pair located on pteromorphs). Four pairs of porose areas (Aa, 26 – 36, A 1, 12 – 18, and sometimes A 3, 16 – 20, circular; A 2, 12 – 16 × 8 – 12 and A 3, 20 – 24 × 12 – 16, oval), with distinct borders. Alveoli of setae la located posteriorly to Aa. Lyrifissures im located between lm and A 1. Opisthonotal gland openings located laterally to A 1. Median pore absent in all specimens. Gnathosoma. Subcapitulum longer than wide (98 – 106 × 90 – 98). Subcapitular setae setiform, slightly barbed; a (18 – 20) longer than m and h (both 8 – 12). Two pairs of adoral setae (6 – 8) setiform, slightly barbed. Palps (82) with setation 0 – 2 – 1 – 3 – 9 (+ ω). Solenidion little longer than half of palptarsus length, straight, thickened, blunt-ended, attached to eupathidium. Chelicerae (118 – 127) with two barbed setae; cha (36 – 45) longer than chb (24 – 28). Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 3) well visible. Four pairs of thin, smooth epimeral setae observed ventrally; all (1 a, 3 b, 4 a, 4 b) similar in length, short (8). Discidia triangular, pointed. Circumpedal carinae distinct. Anogenital region. Six pairs of genital (g 1, g 2, 12 – 16; g 3 – g 6, 8), one pair of aggenital (8), two pairs of anal (8) and three pairs of adanal (8) setae thin, smooth. Anterior edge of genital plates with two setae. Adanal lyrifissures iad located antero-laterally to adanal setae ad 3. Postanal porose area oval (24 × 12 – 16). Legs. Tridactylous; claw of each leg smooth. Formulae of leg setation and solenidia: I (1 – 4 – 3 – 4 – 20) [1 – 2 – 2], II (1 – 4 – 3 – 4 – 15) [1 – 1 – 2], III (1 – 2 – 1 – 3 – 15) [1 – 1 – 0], IV (1 – 2 – 2 – 3 – 12) [0 – 1 – 0]; homologies of setae and solenidia indicated in Table 12.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFFA01613282FE46FCF3F6B6.taxon	materials_examined	Type material. Collection data for holotype (female): locality 3 (soil); collection data for three paratypes (two females and one male): locality 3 (soil). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; one paratype in deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; two paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFFA01613282FE46FCF3F6B6.taxon	etymology	Etymology. The specific name “ longilineata ” refers to the long lamellar lines.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFFA01613282FE46FCF3F6B6.taxon	discussion	Remarks. The genus Neogalumna was proposed by Hammer (1973) with Neogalumna antenniger Hammer, 1973 as a type species. It is morphologically similar to the genus Galumna. The main morphological difference between these two genera is location of adanal lyrifissures: removed from anal plates, located laterally or anterolaterally to adanal setae ad 3 in Neogalumna versus located very close to anal plates, medially, antero-medially or postero-medially to adanal setae ad 3 in Galumna. However, this character state is inherent for some other taxa in the Galumnoidea (for example, Globogalumna Balogh & Balogh, 1990, some Heterogalumna Balogh, 1960). Thus, we consider that Neogalumna should be included as the subgenus in the genus Galumna: Galumna (Neogalumna) Hammer, 1973 stat. n. Hence, all known species of Neogalumna also should be combined: G. (N.) aethiopica (Mahunka & Mahunka-Papp, 2009) comb. n., G. (N.) antenniger (Hammer, 1973) comb. n., G. (N.) araujoi (Perez-Íñigo & Baggio, 1994) comb. n., G. (N.) curviporosa (Balakrishnan, 1986) comb. n., G. (N.) longiporosa (Liang, Yang & Tang, 2014) comb. n., G. (N.) seniczaki (Ermilov & Anichkin, 2010) comb. n. Similarly, the situation on taxonomic status of the genus Globogalumna is presented. It was proposed by J. & P. Balogh (1990) with Allogalumna globulifera Balogh & Mahunka, 1978 as a type species. It is morphologically similar to the genus Allogalumna, but adanal lyrifissures removed from anal plates in Globogalumna versus located very close to anal plates in Allogalumna. Thus, we consider that Globogalumna should be included as the subgenus in the genus Allogalumna: Allogalumna (Globogalumna) Balogh & & Balogh stat. n. Hence, all known species of Globogalumna also should be combined: A. (G.) biporosa (Ermilov & Anichkin, 2012) (d) comb. n., A. (G.) globulifera (Balogh & Mahunka, 1978) comb. n. The subgenus Galumna (Neogalumna) Hammer, 1973 comprises six species, which are distributed in the Oriental and Neotropical regions, north-eastern China, Polynesia and Kenya (data summarized by Subías 2004, updated 2014). In having the combination of long interlamellar setae, setiform and ciliate bothridial setae, presence of anterior margin of notogaster, presence of four pairs of circular porose areas and striate medial parts of genital plates, Galumna (Neogalumna) longilineata sp. nov. is similar to Galumna (Neogalumna) seniczaki (Ermilov & Anichkin, 2010) from Vietnam. However, it clearly differs from the latter by the presence of long lamellar lines, reaching the insertions of rostral setae (versus short, clearly not reaching the insertions of rostral setae in G. (N.) seniczaki), lamellar setae distinctly longer than rostral setae (versus both similar in G. (N.) seniczaki) and larger porose area Aa (diameter 26 – 36 versus 16 – 20 G. (N.) seniczaki).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFF1017C3282FB28FC22F66F.taxon	diagnosis	Diagnosis. Body size 381 – 415 × 265 – 298. Rostral and lamellar setae of medium size, interlamellar setae short. Bothridial setae setiform, ciliate. Lamellar lines straight, short. Four pairs of porose areas oval or circular. Median pore absent. Medial part of genital plates with longitudinal stria. Postanal porose area oval.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFF1017C3282FB28FC22F66F.taxon	description	Description. Measurements. Body length 398 (holotype: female), 381 – 415 (four paratypes: three females and one male); body width 282 (holotype), 265 – 298 (four paratypes). Integument. Body color yellowish-brown. Body surface smooth. Pteromorphs with weakly developed radiate wrinkles. Medial part of genital plates with longitudinal stria. Prodorsum. Rostrum widely rounded. Rostral and lamellar setae of medium size (both 20 – 24), setiform, slightly barbed. Interlamellar setae short (4 – 6), thin, smooth. Bothridial setae (118 – 131) longest on prodorsum, setiform, ciliate. Exobothridial setae absent. Lamellar and sublamellar lines distinct. Lamellar lines straight, short, clearly not reaching insertions of rostral setae. Sublamellar lines typical for Galumna: long, curving backwards. Porose areas Ad oval, transverse, elongate (20 × 4 – 8). Notogaster. Anterior notogastral margin developed. Dorsophragmata small. Notogastral setae represented by ten pairs (additionally c x present) of alveoli (eleventh pair located on pteromorphs). Four pairs of porose areas (Aa, 20 – 32 × 12 – 20, oval; A 1, 16 × 10 – 12, oval or 6 – 10, circular; A 2, 6 – 12, circular; A 3, 8 – 20 × 6 – 12, oval or 6 – 16, circular), with distinct borders. Alveoli of setae la located posteriorly to Aa. Lyrifissures im located between lm and A 1. Opisthonotal gland openings located laterally to A 1. Median pore absent in all specimens. Gnathosoma. Subcapitulum longer than wide (102 – 106 × 98). Subcapitular setae setiform; a (20 – 24) and m (16 – 20) slightly barbed, h (4 – 8) smooth. Two pairs of adoral setae (8 – 12) setiform, slightly barbed. Palps (82) with setation 0 – 2 – 1 – 3 – 9 (+ ω). Solenidion little longer than half of palptarsus length, straight, thickened, blunt-ended, attached to eupathidium. Chelicerae (110 – 123) with two barbed setae; cha (32 – 41) longer than chb (20 – 28). Trägårdh’s organ distinct. Epimeral and lateral podosomal regions. Apodemes (1, 2, sejugal, 3) well visible. Four pairs of thin, smooth epimeral setae observed ventrally; 3 b (12 – 16) little longer than 1 a, 4 a, 4 b (6 – 12). Discidia triangular, pointed. Circumpedal carinae distinct. Anogenital region. Six pairs of genital, one pair of aggenital, two pairs of anal and three pairs of adanal setae similar in length (6 – 12), thin, smooth. Anterior edge of genital plates with two setae. Adanal lyrifissures iad located antero-laterally to adanal setae ad 3. Postanal porose area oval (20 – 24 × 12 – 14). Legs. Tridactylous; claw of each leg smooth. Formulae of leg setation and solenidia: I (1 – 4 – 3 – 4 – 20) [1 – 2 – 2], II (1 – 4 – 3 – 4 – 15) [1 – 1 – 2], III (1 – 2 – 1 – 3 – 15) [1 – 1 – 0], IV (1 – 2 – 2 – 3 – 12) [0 – 1 – 0]; homologies of setae and solenidia indicated in Table 12.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFF1017C3282FB28FC22F66F.taxon	materials_examined	Type material. Collection data for holotype (female): locality 6 (soil); collection data for four paratypes (three females and male): locality 5 (soil). Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; three paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFF1017C3282FB28FC22F66F.taxon	etymology	Etymology. The specific name is dedicated to our colleague, acarologist, Dr. Andrei V. Tolstikov (Tyumen State University, Tyumen, Russia).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
03B82C73FFF1017C3282FB28FC22F66F.taxon	discussion	Remarks. In having the combination of the short and straight lamellar lines, setiform and ciliate bothridial setae, presence of anterior margin of notogaster, presence of four pairs of circular or oval porose areas and striate medial parts of genital plates, Galumna (Neogalumna) tolstikovi sp. nov. is similar to Galumna (Neogalumna) seniczaki (Ermilov & Anichkin, 2010) from Vietnam. However, it clearly differs from the latter by the presence of short (4 – 6) interlamellar setae (versus long, 69 – 77 in G. (N.) seniczaki).	en	Ermilov, Sergey G., Anichkin, Alexander E. (2014): Taxonomic study of oribatid mites (Acari, Oribatida) of Bi Dup — Nui Ba National Park (southern Vietnam). Zootaxa 3834 (1): 1-86, DOI: 10.11646/zootaxa.3834.1.1
