identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B83B6EFFB0FFFDFCD2F9B240B3FCF2.text	03B83B6EFFB0FFFDFCD2F9B240B3FCF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomechusini Flemming	<div><p>Lomechusini Flemming</p> <p>Diagnosis. The aleocharine tribe Lomechusini is a morphologically diverse tribe that has not been</p> <p>satisfactorily defined, but the following characters are often applied to the group: 1) galea elongate; 2) mesocoxae broadly separated; 3) metaventral process much longer than mesoventral process, which extends slightly between mesocoxae; 4) athetine bridge of median lobe present; 5) proximal separation of median lobe that receives compressor plate deeply divided, appearing as if almost reaching parameral proximal costa in lateral view; 6) tarsal formula 4-5-5 (Seevers 1978; Newton et al. 2000; Maruyama 2006).</p> <p>KEY TO THE NORTH AMERICAN GENERA OF LOMECHUSINI</p> <p>1. Abdomen petiolate; head and prothorax with smooth intervals between large setigerous impressions; host not known................................. Ecitocala rugosa Frank and Thomas</p> <p>1′. Abdomen not petiolate; sculpture of head and prothorax different........................................2</p> <p>2. Pronotum with deep, medial, longitudinal sulcus; occurs with ant species Neivamyrmex nigrescens (Cresson)............................................................. Ecitonidia wheeleri Wasmann</p> <p>2′. Pronotum without sulcus..............................3</p> <p>3. Head with prominent Y-shaped carina with bifurcation extending posteriorly; pronotum carinate; occurs with ants in the genus Neivamyrmex Borgmeier................... Ecitoxenidia Wasmann</p> <p>3′. Head and pronotum not carinate..................4</p> <p>4. Pronotum cordiform with anterior margin bisinuate and 2 large circular impressions at base of anterior emargination; host unknown.... Bothronotoxenus tishechkini new genus,....................................................... new species</p> <p>4′. Pronotum not cordiform and anterior margin not bisinuate with 2 large circular impressions......................................................................5</p> <p>5. Paratergites of abdominal segments III-V with prominent tufts of trichomes; occurs with ants in the genera Camponotus Mayr and Formica L.................... Xenodusa Wasmann</p> <p>5′. Paratergites without tufts of trichomes.........6</p> <p>6. Occipital suture complete to hypostoma......7</p> <p>6′. Occipital suture incomplete..........................9</p> <p>7. Mentum length subequal to width at base; galea and lacinia extremely elongate, galea length greater than 7 times width at base and closer to 10 times width at base..................................................... Myrmedonota Cameron</p> <p>7′. Mentum length shorter than width at base; galea and lacinia in most instances long but not extremely so, galea length less than 7 times width at base................................................8</p> <p>8. Labial palpomeres subequal in length; occurs with ants in the genera Crematogaster Lund and Liometopum Mayr............ Pella Stephens............ (s. str.) and Platyusa sonomae Casey</p> <p>8′. Labial palpomere I as long as II and III combined; occurs with ants in the genus Crematogaster....................... Zyras Stephens</p> <p>9. Abdominal tergites IV and V with prominent medial tubercles and inner surfaces covered with trichomes; occurs with ant species Tapinoma sessile (Say)................................................ Myrmoecia Mulsant and Rey</p> <p>9′. Abdominal tergites IV and V without tubercles....................................................................10</p> <p>10. Glossa fused; galea and lacinia short, length of galea 3 times width at base...................11</p> <p>10′. Glossa not fused; galea and lacinia long, length of galea more than 3 times width at base.........................................................12</p> <p>11. Integument with microsculpture; body with macrosetae not extremely long and bristlelike; not occurring with ants............................................. Meronera venestula (Erichson)</p> <p>11′. Integument without microsculpture; body with macrosetae extremely long and bristle-like; not occurring with ants............................................................... Apalonia seticornis Casey</p> <p>12. Eyes large and coarsely faceted, occupying entire lateral margin of head; band of setulate cuticular processes at apex of abdominal tergite VII wide and conspicuous; host unknown........................ Tetradonia megalops (Casey)</p> <p>12′. Eyes not so large and coarsely faceted, not occupying entire lateral margin of head; band of setulate cuticular processes at apex of abdominal tergite VII narrow and less conspicuous......................................................................... 13</p> <p>13. All antennomeres at least quadrate but most longer than wide; elytra shorter than pronotum and with longitudinal impressions along lateral margin..................................................14</p> <p>13′. Some antennomeres transverse; elytra not shorter than pronotum and without longitudinal impressions along lateral margin......15</p> <p>14. Abdominal tergite VII with small, posteriorly projecting medial spine and segment III with broad, smooth eminence; known from California; ecology unknown................................................................... Trachyota Casey</p> <p>14′. Abdominal tergites III and VII without modifications; not known from California; not occurring with ants................ Drusilla Leach</p> <p>15. Abdominal tergites without microsculpture; tergites III-VII with macrosetae but virtually no microsetae; antennae not stout in appearance and conspicuously clavate with antennomeral pedicels clearly visible; not occurring with ants...................................... Xesturida laevis Casey</p> <p>15′. Abdominal tergites with some microsculpture, at least in form of conspicuous setigerous punctures; tergites III-VII with macrosetae and at least tergite III with microsetae; antennae stout in appearance and at most slightly clavate, antennomeral pedicels visible or not.............16</p> <p>16. Body flattened, meso- and metasterna not swollen ventrally in lateral view; antennomeral pedicels at least slightly visible; occurs with ants in the genus Neivamyrmex................................................................... Microdonia Casey</p> <p>16′. Body not flattened, meso- and metasterna swollen ventrally in lateral view; antennomeral pedicels not visible; occurs with ants in the genus Neivamyrmex...................... Dinocoryna Casey</p></div> 	https://treatment.plazi.org/id/03B83B6EFFB0FFFDFCD2F9B240B3FCF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Eldredge, K. Taro	Eldredge, K. Taro (2011): Bothronotoxenus Eldredge, A New Genus Of Aleocharinae (Coleoptera: Staphylinidae) From Arizona, U. S. A. The Coleopterists Bulletin 65 (4): 381-386, DOI: 10.1649/072.065.0413
03B83B6EFFB2FFFDFF30FCE5411EFD45.text	03B83B6EFFB2FFFDFF30FCE5411EFD45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bothronotoxenus Eldredge	<div><p>Bothronotoxenus Eldredge, new genus</p> <p>Diagnosis. Bothronotoxenus and its only species, B. tishechkini, can be separated from all other Lomechusini by its unique pronotum with a bisinuate anterior margin and a pair of deep, circular depressions on the disk just posterior of the sinuations.</p> <p>Description. Integument without microsculpture; tarsal formula 4-5-5; antennae 11-segmented. Head (Figs. 1–2): Transverse (HW = 0.57 mm, HL = 0.42 mm, HW/HL = 1.36) and widest behind eyes; temples swollen posteriorly and extending beyond occipital foramen; occipital and frontal suture absent; neck absent. Antennae compact with antennomeres IV-XI parallel-sided and pedicels scarcely visible. Labrum transverse. Mandibles with apex inwardly recurved, surpassing basointerior articulation. Maxilla (Fig. 5) with cardo ventrally overlapping base of stipes; galea long and consisting of distal lobe with abundant setae; lacinia with several teeth on apex and margin partially delimited by medial dilation, overall accompanied by several layers of setae; palpi 4-segmented, palpomere IV with filamentous sensillae. Labium (Fig. 3) with 3-segmented palpi; palpomeres I and III subequal in length, palpomere II shorter with membranous zone; glossa divided to near base with single terminal sensillate element on each lobe. Mentum (Fig. 6) transverse and trapeziform, with apical margin broadly and shallowly arcuate, posterior-most extension at rounded basolateral angles. Pronotum (Figs. 1–2): Transverse (PW = 0.72 mm, PL = 0.42, PW/PL = 1.71), shape cordiform with anterior margin bisinuate and posterior margin evenly arcuate without posterolateral angles, disk with circular impressions just behind anterior sinuations, marginal bead present and complete, hypomeron fully visible in lateral view. Mesocoxae set in cavities with complete marginal bead, moderately separated by long metaventral and mesoventral process that barely extends between coxae, isthmus short. Elytra (Fig. 1) transverse (EW = 0.78, EL = 0.57, EW/EL = 1.37), laterodistal angles weakly produced but not sinuate. Hind wings fully developed. Abdomen (Figs. 1–2): Widest at base and slightly tapering towards apex, macrosetae inconspicuous and secondary setae extremely long and fine. Tergite VII with transverse defensive gland opening, occupying little less than half width of segment at base.</p> <p>Type species. Bothronotoxenus tishechkini, new species.</p> <p>Etymology. The generic name is derived from Greek to mean guest with pits on its back. Its gender is masculine.</p></div> 	https://treatment.plazi.org/id/03B83B6EFFB2FFFDFF30FCE5411EFD45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Eldredge, K. Taro	Eldredge, K. Taro (2011): Bothronotoxenus Eldredge, A New Genus Of Aleocharinae (Coleoptera: Staphylinidae) From Arizona, U. S. A. The Coleopterists Bulletin 65 (4): 381-386, DOI: 10.1649/072.065.0413
03B83B6EFFB2FFFAFD1DFD354069FEFC.text	03B83B6EFFB2FFFAFD1DFD354069FEFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bothronotoxenus tishechkini Eldredge 2011	<div><p>Bothronotoxenus tishechkini Eldredge, new species</p> <p>Holotype. ♀, “ ARIZONA: Cochise Co. / Chiracahua Mts., 2 mi /NW Portal. 31°55.5′N / 109°09′W. UV light. 17/July 2001 / A. Tishechkin // LSAM/0043201 ” (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-109.15&amp;materialsCitation.latitude=31.925" title="Search Plazi for locations around (long -109.15/lat 31.925)">Field Museum of Natural History</a>; abdominal apex from segment VII, both maxillae and labial complex dissected and preserved as described in Material and Methods).</p> <p>Description. Body (Figs. 1–2) approximately 2.2 mm in length and overall color flavus. Head and pronotum glabrous, except for a few scattered inconspicuous setigerous punctures; elytra with setae extending towards apicolateral angles; abdominal terga III-VII with 1 row of extremely long and fine setae at apical margin, tergite VIII with secondary setae throughout but not as long as on preceding segments; all abdominal sterna with extremely long and fine secondary setae throughout, not restricted to apical margin. Head: Antennae (Figs. 1–2) approximately 2.5 times longer than head (AL = 1.08 mm, AL/HL = 2.57); antennomeres IV-X subquadrate, XI slightly longer than wide (antennomere XI W = 0.15 mm, L = 0.18, W/L = 0.83). Lacinia (Fig. 5) with distal comb comprised of 6 teeth and proximally followed by small medial dilation, margin proximal to distal comb covered in multiple layers of setae; galea long, length approximately 4 times width at base, slightly dilated towards apex and distal lobe with numerous setulae forming tuft; base of palpi ventrally covered by membrane that appears granulose when cleared, segment II subulate and III elongate. Labium (Fig. 3) with setulae b and y absent (in addition, holotype with setula g of left sclerite of second palpomere migrated to base of outer lateral portion, consequently chaetotaxy asymmetrical), prementum with 2 pairs of real pores and 1 pair of setal pores present, pseudopores absent, lateral apodemes relatively short and medial apodeme prominent. Abdomen: Tergite VIII (Fig. 7) with slight constriction at spiracle, distally sides straight and converging to apicolateral angles which run to apicomedial point on distal margin; 4 pairs of inconspicuous macrosetae present along lateral margin (in addition, holotype with pair of large pores that most likely represent macrosetal pores near apicomedial margin); fine secondary setae not nearly as long as those on preceding tergites, sparsely scattered throughout disk. Sternite VIII (Fig. 8) arcuate throughout with rounded apical margin; 4 pairs of inconspicuous macrosetae, 3 pairs along lateral margin and 1 pair just proximal of lateral margin; extremely long and thin secondary setae sparsely scattered throughout disk and row of shorter setae at apical margin. Spermatheca (Fig. 4): Distally somewhat squared and internally bearing cone-like umbilicus; proximally with short bend followed by series of constrictions, creating bunched appearance; elbowed bend followed proximally by medially, asymmetrically inflated stem with inflated portion occupied by less sclerotized section of spermatheca; proximal end terminates in loop that gives rise to spermathecal duct.</p> <p>Etymology. The species name is a patronym dedicated to Alexey Tishechkin, the sole collector of this species and other valuable guests of army ants.</p> <p>Remarks. The morphology of the antennae (compact and robust) and pronotum (cordiform with deep depressions) in B. tishechkini suggests it is myrmecophilous. Based on the shared derived structure of the spermatheca, it is hypothesized that B. tishechkini belongs in a clade along with Dinocoryna, Ecitocala, Ecitoxenidia, and Microdonia. Among these genera, all species with available life history information are known to be guests of Neivamyrmex (Formicidae: Ecitoninae) army ants (Seevers 1959, 1965; Frank and Thomas 1981; Kistner et al. 1996; Newton et al. 2000; Kistner and Berghoff 2008). Therefore, it is likely, based on phyletic affinities, that B. tishechkini is a guest of Neivamyrmex but further collecting efforts will be necessary to confirm this hypothesis.</p> </div>	https://treatment.plazi.org/id/03B83B6EFFB2FFFAFD1DFD354069FEFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Eldredge, K. Taro	Eldredge, K. Taro (2011): Bothronotoxenus Eldredge, A New Genus Of Aleocharinae (Coleoptera: Staphylinidae) From Arizona, U. S. A. The Coleopterists Bulletin 65 (4): 381-386, DOI: 10.1649/072.065.0413
