identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B987A5BA543462A79638EF53D2F93C.text	03B987A5BA543462A79638EF53D2F93C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acianthera nikoleae A. Doucette & J. Portilla 2016	<div><p>Acianthera nikoleae A.Doucette &amp; J.Portilla, sp. nov. (Figs. 1, 11D)</p> <p>Type:— ECUADOR. Zamora-Chinchipe: Pangui, reported to grow on coffee trees around 1,200 m, flowered in cultivation at Ecuagenera Orchid Nursery, Gualaceo, Ecuador, 4 September 2015, Doucette 0319 (holotype: HA).</p> <p>Diagnosis: The new species is most similar to Acianthera melanochthoda (Luer &amp; Hirtz in Luer 1996: 169) Pridgeon &amp; M. W.Chase (2001: 244) but differs in the hispid sepals that are larger and expanded/fused into lamina.</p> <p>Epiphytic, repent herbs, roots 0.6 mm in diam. Stems 3.2–5.9 5 0.8–1.4 mm, enveloped by 2–3 papery sheaths, 5.0 5 3.0 mm. Leaves elliptical, verrucose, 26–48 5 14–24 mm. Inflorescence with a single flower; peduncle 1.4 5 0.4 mm; bracts 2.3 5 1.2 mm; pedicel 2.7 5 1.0 mm; ovary pubescent, 0.9 5 1.0 mm; dorsal sepal lanceolate, rose and orange, hispid, 11.0 5 2.9 mm; synsepalum oblong with a triangular recess, similar in colour to the dorsal sepal, 16.0 5 5.0 mm; petals spathulate with a toothed margin towards the acute apex, orange, 4.0 5 1.0 mm; lip lanceolate with a toothed margin, orange, 3.4 5 1.0 mm; column 3.2 5 0.6 mm.</p> <p>Etymology:— Named to honour Nikole Portilla, daughter of Jose Portilla, president of Ecuagenera.</p> <p>Notes:— This species belongs to a group of about eight (The Plant List 2015) attributable to Apoda-prorepentia (Luer 1986: 31) Luer (2004: 255), characterized by repent habit, single-flowered inflorescences and a papery bract. Acianthera nikoleae is most similar to A. melanochthoda, but differs in the larger flower size (e.g. synsepalum 11 mm vs. 7 mm long) and longer pubescence on the sepals and margin of the petals (toothed vs. entire). The sepalar morphology of A. nikoleae is unique in the group because the lateral sepals are expanded and fused into a thin layer extended beyond a triangular cavity formed in the centre of the synsepalum.</p> <p>The group of species similar in morphology to Acianthera nikoleae was originally attributed to Pleurothallis subgen. Apoda-prorepentia Luer (1986: 31). The species were transferred to Acianthera after molecular phylogenetic analyses (Pridgeon et al. 2001) indicated that Acianthera melanochthoda rendered Pleurothallis non-monophyletic (Pridgeon &amp; Chase 2001, Solano et al. 2011, Karremans &amp; Rincon-Gonzalez 2015). The International Plant Names Index (2016) indicated that A. nikoleae represents one of at least two species described since the key to the species (Luer 2007) was published. The second species recently described, Apoda-prorepentia hirtzii Luer in Luer &amp; Thorele (2013: 174), appears to be referable to Acianthera dodsonii, which is also reported to have purple spots on the leaves that are not raised (Luer 2007), and otherwise matches the latter in distribution and morphology of plant and flowers. We treat this a synonym of Acianthera dodsonii.</p> </div>	https://treatment.plazi.org/id/03B987A5BA543462A79638EF53D2F93C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA543461A7963D3755B3F818.text	03B987A5BA543461A7963D3755B3F818.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Masdevallia constricta fm. glabra A. Doucette & J. Portilla 2016	<div><p>Masdevallia constricta forma glabra A.Doucette &amp; J.Portilla, f. nov. (Figs. 2, 11A)</p> <p>Type:— ECUADOR. Without a specific locality: the typical form is reported from southern Ecuador and neighbouring Peru, flowered in cultivation at Ecuagenera Orchid Nursery, 4 September 2015, Doucette 0307 (holotype: HA).</p> <p>Diagnosis: this form differs from M. constricta f. constricta Poeppig &amp; Endlicher (1837: 6) in having glabrous sepals.</p> <p>Epiphytic, caespitose herbs, roots 1 mm in diam. Stems 30–35 5 2 mm, enveloped by 2–3 papery sheaths. Leaves spathulate, 100–135 5 15–22 mm. Inflorescences with a single flower; peduncle 80.0–100.0 5 1.5 mm; bracts 10–11 5 3 mm; pedicel 12–15 5 2 mm; ovary 7 5 3 mm; dorsal sepal spathulate, suffused with yellow, glabrous, 25 5 12–14 mm, contracted into a yellow tail, 32–59 mm long; lateral sepals sigmoid, broadest at the apex, similar in colour to the dorsal sepal, 25–30 5 10–15 mm, contracted into a tail similar to that of the dorsal sepal; petals oblong with a retrorse tooth, white, 5 5 3 mm; lip pandurate, white, suffused with pink, 5 5 3 mm; column 6.5 5 1.5 mm.</p> <p>Etymology:— From the Latin glaber, meaning hairless, referring to the trait that distinguishes this form from the typical.</p> <p>Notes:— Masdevallia constricta is reported (Luer 2002) to be frequent in the Andes from southern Ecuador into central Peru between 1500–1830 m. Luer (2002) stated that “ Masdevallia constricta is variable in size, and in the yellow or orange suffusion or veining of the sepaline tube.” The morphology of the new form described here is within the range of variation described by Luer (2002), except for the absence of pubescence on the sepals, which has never been reported for the species. No forms of Masdevallia constricta are reported by International Plant Names Index (2016) or Luer (2002). We feel the aberrant individual described here could be confused with an undescribed species, and therefore it is useful to indicate to other researchers its presence with this formal description.</p> </div>	https://treatment.plazi.org/id/03B987A5BA543461A7963D3755B3F818	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA563460A7963D2452CAF843.text	03B987A5BA563460A7963D2452CAF843.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Masdevallia pachyura fm. leptoura (Luer) A. Doucette	<div><p>Masdevallia pachyura forma leptoura (Luer) A.Doucette, stat. et comb. nov.</p> <p>Basionym: Masdevallia leptoura Luer (1983: 381)</p> <p>Homotypic synonym: Masdevallia pachyura subsp. leptoura (Luer) Luer (1988b: 44)</p></div> 	https://treatment.plazi.org/id/03B987A5BA563460A7963D2452CAF843	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA513466A7963A995462FD1A.text	03B987A5BA513466A7963A995462FD1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Masdevallia pachyura fm. pardinella A. Doucette & J. Portilla 2016	<div><p>Masdevallia pachyura forma pardinella A.Doucette &amp; J.Portilla, f. nov. (Figs. 3, 11B)</p> <p>Type:— ECUADOR: Azuay: Shanglly, 2600m, flowered in cultivation at Ecuagenera Orchid Nursery, 4 September 2015, Doucette 0312 (holotype: HA).</p> <p>Diagnosis: The form differs from M. pachyura f. pachyura Reichenbach (1874: 322) and M. pachyura f. leptoura in having entirely yellow sepals and a broader dorsal sepal.</p> <p>Epiphytic, caespitose herbs, roots 1.5 mm in diam. Stems 4–6 5 2 mm, and enveloped by 2–3 papery sheaths, 7–20 5 4 mm. Leaves spathulate, 80–195 5 12–20 mm. Inflorescence a raceme; peduncle 97.0–101.0 5 1.0– 1.5 mm; bracts 4–5 5 3 mm; pedicel 2.0–3.0 5 1.5 mm; ovary crested, 2.0 5 2.5 mm; dorsal sepal round, yellow, spotted with purple, 10–12 5 7–9 mm, contracted into a yellow tail, 3–5 5 1 mm; lateral sepals oblong, similar in color to the dorsal sepal, 11–12 5 3 mm, contracted into tails similar to that of the dorsal sepal; petals oblong, white suffused with yellow at the apex, tridentate, 6.0 5 2.2 mm; lip oblong, cleft at the base, constricted below the middle, the apex rounded, yellow suffused with orange at the apex, 5 5 2 mm; column 5.0 5 1.5 mm.</p> <p>Etymology:— From the Latin pardus meaning leopard, referring to the yellow and spotted flowers.</p> <p>Notes:— Masdevallia leptoura was first described to recognize a sympatric taxon resembling M. pachyura, but differing in the “larger [plant size] with more flowers with narrow, non-clavate tails”, and subsequently reduced to a subspecies of M. pachyura based on Luer’s (1988) opinion that “two of the most distinctive forms have been described as species. Masdevallia aureodactyla is little more than a color form of M. pachyura, and the other [M. leptoura] is reduced to a subspecies.” We agree with Luer (1988) and believe that this distinctive plant should be recognized at the rank of form.</p> <p>Given the overlap in morphology and geographic distribution, we hypothesize that M. leptoura and M. pachyura represent a single polymorphic species. Masdevallia pachyura f. leptoura is hypothesized to be a form with narrower tails, and M. pachyura f. pardinella is hypothesized to be a form of the species with yellow sepals and yellow, non-clavate tails. We feel that a form is the most appropriate rank for these two taxa because we hypothesize that they are embedded within a clade including populations matching the description of M. pachyura. Masdevallia aureodactyla is distinguished from M. pachyura f. pardinella by the orange and clavate rather than yellow and non-clavate tails. No forms of Masdevallia pachyura are reported by International Plant Names Index (2016) or Luer (2003).</p> </div>	https://treatment.plazi.org/id/03B987A5BA513466A7963A995462FD1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA503466A796391150CCF9F2.text	03B987A5BA503466A796391150CCF9F2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Masdevallia posadae fm. rugosepala A. Doucette & J. Portilla 2016	<div><p>Masdevallia posadae forma rugosepala A.Doucette &amp; J.Portilla, f. nov. (Figs. 4, 11C)</p> <p>Type:— ECUADOR. Without a specific locality, flowered in cultivation at Ecuagenera Orchid Nursery, 4 September 2015, Doucette 0308 (holotype: HA).</p> <p>Diagnosis: The form differs from M. posadae f. posadae (Luer 1982a: 74) in having rugose sepals and shorter thicker tails.</p> <p>Epiphytic, caespitose herbs, roots 1 mm in diam. Stems 14–17 5 3 mm, enveloped by 2–3 papery sheaths, 5–25 5 5–7 mm. Leaves spathulate, 100–170 5 17–20 mm. Inflorescence a successively flowered raceme; peduncle triquetrous, 15–280 5 3 mm; bracts 14–17 5 4–6 mm; pedicel 15–25 5 4–6 mm; ovary 4 5 2 mm; dorsal sepal ovate, yellow suffused with orange, 14 5 7 mm, contracted into a yellow tail, 31 5 2 mm wide; the lateral sepals ovate, rugose, 24 5 5.5 mm, contracted into a yellow tail, 20 5 2 mm; the petals white, oblong, 6 5 2 mm; column 5.5 5 1.5 mm.</p> <p>Etymology:— From the Latin ruga meaning “wrinkle” and sepalus meaning “sepal”. The name refers to the sepals, which distinguish this form from the typical form.</p> <p>Notes:— Masdevallia posadae f. rugosepala appears to be an unusual phenotype of M. posadae, a species that had not be previously reported from Ecuador. An individual of the form described here was self-pollinated at Ecuagenera and produced individuals with rugose and non-rugose sepals in addition to individuals with short thick tails and long thin tails. The form is striking and appears distinct from M. posadae, but the traits distinguishing the typical form and the new form described here appear to be phenotypic variation within the species. No forms of Masdevallia posadae are reported by International Plant Names Index (2016) or Luer (2000). We feel the aberrant individual described here could be confused with an undescribed species, and therefore it is useful to indicate to other researchers its presence with its formal description.</p> </div>	https://treatment.plazi.org/id/03B987A5BA503466A796391150CCF9F2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA50346BA7963C7951AEFD8A.text	03B987A5BA50346BA7963C7951AEFD8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleurothallis cardiostola fm. magnidraba A. Doucette & J. Portilla 2016	<div><p>Pleurothallis cardiostola forma magnidraba A.Doucette &amp; J.Portilla, f. nov. (Figs. 5, 11G)</p> <p>Type:— ECUADOR. Without a specific locality, flowered in cultivation at Ecuagenera Orchid Nursery, 4 September 2015, Doucette 0311 (holotype: HA).</p> <p>Diagnosis: The new form differs from Pleurothallis cardiostola f. cardiostola Reichenbach (1854: 128) in its larger flower and number of veins in the dorsal sepal.</p> <p>Epiphytic, caespitose herbs; roots 1 mm in diam. The stem 180–260 5 2.5 mm, enveloped by two to three tubular, papery sheaths 10–60 5 2.5 mm. Leaves sessile, cordate, 130–192 5 22–28 mm. Inflorescence fasciculate, protected by an erect, papery sheath, 18 mm long; peduncle 3 5 1 mm, with a tubular bract before the pedicel, 1–2 5 2 mm; pedicel 10 5 1 mm; ovary papillose, 7 5 2 mm, flowers non-resupinate, dorsal sepal broadly ovate, 11-veined, tan, minutely pubescent, 20 5 15 mm; synsepalum round, concave below the middle, similar in colour and texture to the dorsal sepal, 14 5 15 mm; petals lanceolate, similar in colour and texture to the sepals, 14 5 5 mm; lip white suffused with yellow, shiny, heater-shaped, 5 5 4 mm; column white suffused with yellow, with a shiny yellow bilobed stigma, 3 5 3 mm.</p> <p>Etymology:— From the Latin magnis meaning “large” and the English “drab” meaning a dull brown colour, referring to the large dull sepals.</p> <p>Notes:— The genus Pleurothallis is one of the largest plant genera including over 500 species (The Plant List 2015). Phylogenetic analyses confirmed the genus to be non-monophyletic (Pridgeon et al. 2001). We hypothesize the new form attributed to Pleurothallis cardiostola to belong to Pleurothallis s.s. based on morphology and the placement of Pleurothallis cardiantha Reichenbach (1876: 15) sister to the type P. ruscifolia (Jacquin 1760: 29) R.Br. in W.T. Aiton (1813: 211) with 100% bootstrap support based on a combined data set including two plastid genes (matK and trnL-F) and a nuclear ribosomal gene (nrITS; Pridgeon et al. 2001).</p> <p>Although there are morphological characters that easily distinguish the taxon described here from typical Pleurothallis cardiostola we hypothesize that it represents an aberrant individual embedded in P. cardiostola. The individual described here differs in traits that have been documented to vary within the species (Luer 2005) and is not reported from a locality disjunct from reported populations of P. cardiostola (i.e. Caribbean or Atlantic forests of Brazil).</p> <p>We feel that naming this new form of P. cardiostola brings it to the attention of scientists who can test the taxonomic hypothesis presented here using a molecular systematic approach on individuals representing different populations across the range of P. cardiostola. If the taxon described here is not embedded within P. cardiostola we feel that the number of veins of the dorsal sepal and larger flower size are sufficient to warrant recognition at the rank of species. No forms of Pleurothallis cardiostola are reported by International Plant Names Index (2016) or Luer (2005).</p> </div>	https://treatment.plazi.org/id/03B987A5BA50346BA7963C7951AEFD8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA5D346BA79638A1540AF9BA.text	03B987A5BA5D346BA79638A1540AF9BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleurothallis gigiportillae A. Doucette & J. Portilla 2016	<div><p>Pleurothallis gigiportillae A.Doucette &amp; J.Portilla, sp. nov. (Figs. 6, 11F)</p> <p>Type:— ECUADOR. Without specific locality, flowered in cultivation at Ecuagenera Orchid Nursery, 4 September 2015, Doucette 0303 (holotype: HA).</p> <p>Diagnosis: The species is most similar to Pleurothallis imperialis Luer (1975: 186) but differs in the free column foot and pubescent apex of the lip.</p> <p>Epiphytic, caespitose herbs, roots 1 mm in diam. Stems 140–420 5 2–5 mm, enveloped by 2–3 tubular, papery sheaths 15–75 5 205 mm. Leaves sessile, cordate, 140–160 5 50–75 mm. Inflorescence fasciculate, protected by a papery bract 20 mm long; peduncle 2 5 1 mm, with a tubular bract before the pedicel, 6.0 5 2.5 mm; the pedicel 11–16 5 2 mm; ovary 7 5 2 mm, dorsal sepal ovate with 15 veins, yellow suffused with purple towards the base, concave, 22–25 5 12–15 mm; synsepalum ovate with 16 veins, purple, flat, 18–20 5 13 mm; petals oblong, the apex acute, purple, spiculate, 11–12 5 4 mm; lip heater-shaped, purple, apex pubescent and bent downwards, 5 5 4 mm; column purple with a shiny, greenish, bilobed stigma, 2 5 3 mm.</p> <p>Etymology:— Named for Gigi Portilla, daughter of Jose Portilla, president of Ecuagenera.</p> <p>Notes:— Pleurothallis gigiportillae also appears to belong to Pleurothallis s.s. based on the similarity to P. cardiantha (solitary flower produced at the base of a cordate leaf and transversely bilobed stigma). The species is unusual in Pleurothallis subsection Macrophyllae -Fasciculatae Luer (1988a: 133) in having a free column foot, which may be unique in the section. It is difficult to hypothesize which species P. gigiportillae is most closely related to given the free column foot and pubescent lip. Superficially, the species resembles P. imperialis based on the size and habit of the plant the colour of the flowers. The new species keys to Pleurothallis megalorhina Luer &amp; R.Escobar (1996: 71) in Luer (2005), but differs in the free column foot and the pubescent, detrorse apex of the lip.</p> <p>Pleurothallis gigiportillae represents one of at least five species attributable to Pleurothallis subsection Macrophylla-Fasciculatae (Pleurothallis adventurae Karremans &amp; Bogarin (2011: 112); P. bogarinii Pupulin &amp; J.D.Zuniga (2007: 690); P. neobarbosae J.M.H.Shaw (2014: 77); P. oscarii Archila &amp; Chiron (2011: 201); P. silvae-pacis Karremans (2011: 156)) described since the key to the species (Luer 2005) was published. The species is not similar to any of the species listed by International Plant Names Index (2016) published between 2005 and 2015.</p> </div>	https://treatment.plazi.org/id/03B987A5BA5D346BA79638A1540AF9BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA5D346AA7963CB154DFF84D.text	03B987A5BA5D346AA7963CB154DFF84D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleurothallis papillingua A. Doucette & J. Portilla 2016	<div><p>Pleurothallis papillingua A.Doucette &amp; J.Portilla, sp. nov. (Figs. 7, 11E)</p> <p>Type:— ECUADOR. Without a specific locality, flowered in cultivation at Ecuagenera Orchid Nursery, 4 September 2015, Doucette 0314 (holotype: HA).</p> <p>Diagnosis: The species is most similar to Pleurothallis tiarata Luer &amp; Hirtz in Luer (2004: 238) differing in the fewer-flowered inflorescence, longer petals and papillose lip.</p> <p>Epiphytic, caespitose herb; roots 1 mm in diam. The stem 120–335 5 3 mm, and enveloped by two to three tubular, papery sheaths 13–55 5 3 mm. Leaves sessile, lanceolate, 90–150 mm 5 16–20 mm. Inflorescence with a single flower, rarely a two flowered raceme, protected by a papery sheath at the base, 18 mm long; the peduncle 24–27 5 1 mm, with a tubular bract before the pedicel, 7 5 2 mm; the pedicel 6 5 1 mm; ovary 5.0 5 1.5 mm, flowers non-resupinate, dorsal sepal ovate, beige, 7 5 6 mm; synsepalum broadly ovate, similar in color to the dorsal sepal, 7 5 8 mm; petal sigmoid, darker beige than the sepals, 10 5 2 mm; lip reniform, papillose, pink, 4 5 4 mm; column 3.0 5 1.5 mm.</p> <p>Etymology:— From the Latin papula, a small protuberance, and lingua, tongue, in reference to the texture of the lip, which distinguishes the new species from similar species.</p> <p>Notes:— The species belongs to Pleurothallis subsect. Macrophyllae -Racemosae Luer (1986: 61) and is excluded from the key (Luer 1999) at couplet 17 because it lacks both calli and broadly rounded lateral lobes. The species is unique in the subsection in having a densely papillose lip. No new species attributable to Pleurothallis subsect. Macrophyllae-Racemosae is reported by International Plant Names Index (2016) to have been published 2005–2015.</p> </div>	https://treatment.plazi.org/id/03B987A5BA5D346AA7963CB154DFF84D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA5F346FA7963CAC537AFE56.text	03B987A5BA5F346FA7963CAC537AFE56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stelis glaucus A. Doucette & J. Portilla 2016	<div><p>Stelis glaucus A.Doucette &amp; J.Portilla, sp. nov. (Figs. 8, 11I)</p> <p>Type:— ECUADOR. Without specific locality, flowered in cultivation at Ecuagenera Orchid Nursery, 4 September 2015, Doucette 0310 (holotype: HA).</p> <p>Diagnosis: The species is most similar to Stelis gemma Garay (1969: 77) but differs in its larger size, development of the inflorescence, and colour of the sepals.</p> <p>Epiphytic, caespitose herbs, roots 1mm in diam. Stems 17.0–20.0 5 1.5–2.0 mm, and enveloped by 2–3 papery sheaths, 10.0–20.0 5 1.5–2.0 mm. Leaves pendent, oval, petiolate, glaucous, 90–98 5 31–39 mm. Inflorescence a raceme; peduncle 30–35 5 1 mm; bracts papery, many, 2–4 5 3–5 mm; pedicel 2 5 1 mm; ovary 4 5 1 mm, dorsal sepal triangular, tan, minutely pubescent, 6 5 5 mm; lateral sepals similar to the dorsal sepal; petals rectangular, concave, dark tan, 0.5 5 1.5 mm; lip semicircular, dark tan, 1 5 1 mm; column bilobed, 1.2 5 1.0 mm.</p> <p>Etymology:— The name of the Greek sea-god, Glaucus, born mortal and turned immortal upon eating a magical herb, is used as a noun in apposition for the species epithet.</p> <p>Notes:— Stelis s.s. was discovered to be embedded in a grade formed by species of Condylago, Pleurothallis, and Salpistele. In order to maintain a monophyletic Pleurothallis the circumscription of Stelis was expanded to include these lineages (Pridgeon et al. 2001). The decision was controversial based on the absence of morphological characters that diagnose the clade. We hypothesize that the species described here, Stelis glaucus, belongs to the clade thought to include the type of the genus Stelis based on the shared traits of a racemose inflorescence, transversely bilobed stigma and reduced petals tightly flanking the column.</p> <p>Stelis glaucus is unusual in the genus given its habit and similar to only one other described species, S. gemma. Luer (2009) noted that Stelis gemma “is unique in the genus with the pendent, overlapping, glaucous green leaves”. The species described here represents a second species in the genus sharing this habit.</p> <p>Stelis glaucus represents one of over 120 new species attributable to Stelis s.s. described since the treatment of Stelis gemma by Luer (2009). Of the 120+ new species, S. glaucus is only similar to S. pendulata O. Duque (2010: 141) but differs in the leaf width (3.1–3.9 cm vs. 1.8 cm) and leaf (glaucus vs. green) and flower colour (beige vs. purple). Other than S. pendulata, S. glaucus not similar to any of the species listed by International Plant Names Index (2016) published between 2009 and 2015.</p> </div>	https://treatment.plazi.org/id/03B987A5BA5F346FA7963CAC537AFE56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA59346FA7963BDD542DFD1A.text	03B987A5BA59346FA7963BDD542DFD1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stelis pachyglossa (Lindley 1840) Pridgeon & M. W. Chase 2001	<div><p>Stelis pachyglossa (Lindley 1840: 68) Pridgeon &amp; M.W.Chase (2001: 265)</p> <p>Heterotypic synonym: Stelis guerrerensis Soto Arenas &amp; R.Solano in Solano (2014: 255)</p> <p>NOTE:— S. guerrerensis is distinguished by Solano (2014) based on the smaller habit and flowers and characters the author notes to be continuous with the new species (i.e. “ Stelis guerrerensis also differs by its more calyptrate and warty petals on the outer surface and broader callus of the lip.”). We hypothesize that S. guerrerensis is a superfluous renaming of S. pachyglossa given the shared distribution and morphological traits, including “calyptrate and warty petals”.</p> </div>	https://treatment.plazi.org/id/03B987A5BA59346FA7963BDD542DFD1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA59346FA796391153DAFB66.text	03B987A5BA59346FA796391153DAFB66.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stelis retusa (La Llave & Lexarza 1825) Pridgeon & M. W. Chase 2001	<div><p>Stelis retusa (La Llave &amp; Lexarza 1825: 40) Pridgeon &amp; M.W.Chase (2001: 266)</p> <p>Heterotypic synonym: Stelis sotoarenasii R.Solano (2010: 450)</p> <p>NOTE:— Stelis sotoarenasii was considered by the author to be a geographic variant of S. retusa and compared to S. aristocratica (L.O.Williams 1946: 237) Solano &amp; Soto Arenas in Hagsater &amp; Soto (2003: 11), which is considered to be a synonym of S. retusa (The Plant List 2015). Stelis sotoarenasii is distinguished from both by Solano (2010) in that it is intermediate in size between the two. We feel this is not a strong argument for the distinctness as a new species of what the author had previously considered a geographic variant of S. retusa and that it should be recognized as a synonym of S. retusa along with S. aristocratica.</p> </div>	https://treatment.plazi.org/id/03B987A5BA59346FA796391153DAFB66	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA59346FA7963E8D5273F9F2.text	03B987A5BA59346FA7963E8D5273F9F2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stelis transversalis Ames 1934	<div><p>Stelis transversalis Ames (1934: 91)</p> <p>Heterotypic synonyms: Stelis anagraciae Archila &amp; Szlach. in Archila (2013: 31)</p> <p>Stelis kaiae R.Solano &amp; C.Dietz. in Solano (2014: 258).</p> <p>NOTE:— Stelis anagraciae reported from Guatemala matches S. transversalis morphologically and appears to be a superfluous re-naming of the latter species, which has a distribution across Central America into Colombia and Ecuador. The species S. kaiae also matches S. transversalis morphologically and appears to be a superfluous re-naming of the same species based on an individual from Mexico.</p> </div>	https://treatment.plazi.org/id/03B987A5BA59346FA7963E8D5273F9F2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA59346EA7963C795561F85F.text	03B987A5BA59346EA7963C795561F85F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trisetella portillae A. Doucette 2016	<div><p>Trisetella portillae A.Doucette, sp. nov. (Figs. 9, 11J)</p> <p>Type:— ECUADOR. Morona-Santiago: Santiago, 800m, flowered in cultivation at Ecuagenera Orchid Nursery, 4 September 2015, Doucette 0313 (holotype: HA).</p> <p>Diagnosis: The species is most similar to Tristella gemmata (Reichenbach 1883: 294) Luer (1980: 57) but differs in the pandurate calyx and scabrose peduncle.</p> <p>Epiphytic, caespitose herbs, roots 0.5 mm in diam. Stems 4–5 5 1 mm, enveloped by tubular, papery sheaths 1.0–4.0 5 1.0– 1.5 mm. The leaf linear, 23–34 5 3 mm. Inflorescence a successively flowered raceme; peduncle scabrose, 50.0–56.0 5 0.2 mm; bracts 2 5 1 mm; pedicel 9.0–12.0 5 0.2 mm; ovary 1.0 5 0.5 mm; dorsal sepal semicircular, yellow suffused with red-orange, 5 5 4 mm, contracted into a yellow tail, 7.0 5 0.2 mm; synsepalum oblong, red-purple, 10.0 5 2.5 mm, contracted into yellow tails, 6.0 5 0.2 mm; petal ovate, beige with a red stripe, 2 5 1 mm; lip sagitate, purple, 2.8 5 1.0 mm; column green suffused with purple with a free column-foot, the apex yellow, a nodule before the stigma, 3 5 1 mm.</p> <p>Etymology:— Name to honour the president of Ecuagenera Orchid Nursery, Jose Portilla.</p> <p>Notes:— The genus Trisetella was originally attributed to Masdevallia but was later segregated by Luer (1980) based on sepal and lip morphology. The species were found to be sister to a clade composed of other genera formerly attributed to Masdevallia (Pridgeon et al. 2001). Tristella portillae keys to Trisetella triglochin (Reichenbach 1877: 648) Luer (1980: 58) but differs in the pandurate calyx and the nodule before the stigma. In most species there is a protrusion before the stigma, referred to by Luer (1989) as the “edge of the stigma”, but in the species described here the edge of the stigma is swollen into a nodule like that of Trisetella gemmata.</p> <p>Trisetella portillae represents one of three species (T. klingeri Luer (2007: 112); T. lasiochila Pupulin (2000: 30); T. dalstroemii Luer (1994: 136)) attributable to Trisetella described since the key to the species was published (Luer 1989). The species can be distinguished from the species listed by International Plant Names Index (2016) published between 2009 and 2015 by the pandurate calyx and the nodule before the stigma.</p> </div>	https://treatment.plazi.org/id/03B987A5BA59346EA7963C795561F85F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA5B3473A7963A9951A2FC16.text	03B987A5BA5B3473A7963A9951A2FC16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zootrophion machaqway A. Doucette & J. Portilla 2016	<div><p>Zootrophion machaqway A.Doucette &amp; J.Portilla, sp. nov. (Figs. 10, 11H)</p> <p>Type:— ECUADOR. Zamora-Chinchipe: Manchinaza, without a reported elevation, flowered in cultivation at Ecuagenera Orchid Nursery, 5 September 2015, Doucette 0315 (holotype: HA).</p> <p>Diagnosis: The species is most similar to Zootrophion atropurpureum (Lindley 1836: 1797) Luer (1982b: 80), but differs by the repent habit and broader flowers.</p> <p>Epiphytic, repent herbs, roots 1 mm in diam. Stems 3–4 5 1 mm, and enveloped by 3–5 sheaths, 1–12 5 2–3 mm. Leaves ovate, petiolate, 25–40 5 2–3 mm. Inflorescence with a single flower; peduncle 7.1–10.0 5 0.7 mm, with a tubular sheath 3.4–5.1 5 0.6–2.0 mm, before the pedicel; pedicel 4.0 5 1.5 mm; ovary crested, 4.5 5 2.5 mm, dorsal sepal ovate, purple, connate at the apex and below the middle to the synsepalum, 15.0–20.0 5 4.5–7.0 mm; synsepalum similar in colour and shape and connate to the dorsal sepal but with two low crests running from below the middle to the apex of the synsepalum, 15–20 5 5–7 mm wide; the petals lanceolate, purple, 4.0 5 1.5 mm; the lip sagittate, purple, 5 5 1 mm; column 3 5 1 mm.</p> <p>Etymology:— From the Quechua word for snake, in reference to the serpent-like growth habit and flowers, resembling the head of a snake, used as a noun in apposition.</p> <p>Notes:— Zootrophion Luer (1982b: 80) was recognized to accommodate species that were previously attributed to Cryptophoranthus Barbosa Rodriguez (1881: 79), based on the hypothesis that it was an artificial genus composed of unrelated species. That hypothesis was later supported by phylogenetic analysis of DNA sequence data (Pridgeon et al. 2001), and Zootrophion was found to be more closely related to Lepanthes Swartz (1799: 85) than the lectotype for the genus Cryptophoranthus, C. fenestratus Barbosa Rodriguez (1881: 79), now known to belong in Acianthera Scheidweiler (1842: 292).</p> <p>Zootrophion is presently circumscribed to include 21 species (The Plant List 2015) and at least one other (Z. ildephonsi P.Ortiz (2012: 37)) that can be divided into 19 species with a caespitose habit and two species (Zootrophion hirtzii Luer (1984: 1293), Z. serpentinum Luer (1984: 1297)) with a repent habit. At least 11 species were described after the publication of the key we used to the species (Luer 2004), and the time of writing this article Zootrophion machaqway is readily distinguished from the two species with a repent habit by the lateral sepal that is fused at the base to the lateral sepal instead of free at the base and in having a carinate dorsal sepal. Although the species is repent, we hypothesize that the new species is more closely related to the caespitose Zootrophion species based on the carinate sepals, fusion of the dorsal sepal to the base to the lateral sepals, petals about as long as the column and acute apex of the lip.</p> </div>	https://treatment.plazi.org/id/03B987A5BA5B3473A7963A9951A2FC16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
03B987A5BA453473A7963E1D5421FABE.text	03B987A5BA453473A7963E1D5421FABE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zootrophion serpentinum Luer (1984: 1297	<div><p>Zootrophion serpentinum Luer (1984: 1297)</p> <p>Heterotypic synonyms: Epibator ximenae Luer &amp; Hirtz in Luer (2004: 204)</p> <p>Zootrophion ximenae (Luer &amp; Hirtz) Pfahl (2014: epublished)</p> <p>NOTE:—The species matches Z. serpentinum in its distribution and vegetative and floral morphology (Luer 2004), but it was compared to Z. hirtzii in the original description from which it differs in the leaf shape and flower colour. We feel Z. ximenae was described as a new species in error and represents a superfluous re-naming of Z. serpentinum.</p> </div>	https://treatment.plazi.org/id/03B987A5BA453473A7963E1D5421FABE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Doucette, Alfonso;Portilla, Jose;Cameron, Kenneth M.	Doucette, Alfonso, Portilla, Jose, Cameron, Kenneth M. (2016): Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador. Phytotaxa 257 (3): 230-248, DOI: 10.11646/phytotaxa.257.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.257.3.2
