taxonID	type	description	language	source
03B987E6FFB31070FF05FCCFEFF2FBC8.taxon	diagnosis	Diagnosis. The Tephritis maccus species group can be recognized among all other species groups of Tephritis by the following combination of characters: wing pattern radiate; basal cells hyaline; pterostigma brownish without hyaline spot; apical fork connected to main wing pattern (except some male specimens of T. urelliosomima); cell r 1 usually without or with only one large hyaline spot or indentation on anterior margin of wing (sometimes an additional small hyaline spot present); crossvein r-m in dark area; and aculeus moderately broad, sharply narrowing and pointed at extreme apex. The latter character occurs only in species of the T. maccus group and is believed to be a synapomorphy of this group of species.	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB01073FF05FEA8ECB7FC46.taxon	materials_examined	Meterial examined. Type material. Holotype (female): Iran: West Azerbaijan province, 10 km W Ziveh, 2700 m, 37 ° 08 ' N, 44 ° 52 ' E, 24. vii. 2012 (Mohamadzade & Najarpoor leg.) (JAZM); Paratypes: Azerbaijan: Talysh, vic. Lerik, Dzhoni vil. [= Çoni, 38 ° 36.8 ' N 48 ° 30.5 ' E, h = 1390 m], 5. vi. 1981, 1 ♀ (V. Ermolenko leg.) (SIZK); Iran: same collection data as in holotype, 3 ♀, 2 ♂; East Azerbaijan Province, Sahand ski resort, 30 km of Tabriz, 37 ° 45.850 ' N 46 ° 30.754 ' E, 2900 m, 30. viii. 2011, 1 ♀, 1 ♂ (Mohamadzade & Najarpoor leg.); Ardabil Province, Sabalan Mountain, 2900 m; 12. vii. 2012, swept from Senecio sp., 1 ♀, 1 ♂ (Mohamadzade & Najarpoor leg.) (JAZM, SIZK and SMNC); Non-type: Sabalan Mountain, 2900 m; 4. vii. 2014, 1 ♂ (Korneyev & Mohamadzade) (SIZK).	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB01073FF05FEA8ECB7FC46.taxon	distribution	Distribution. Azerbaijan and Iran (Mohamadzade Namin & Korneyev 2012).	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB01073FF05FEA8ECB7FC46.taxon	diagnosis	Diagnosis. Tephritis azari is similar to T. maccus Hering and T. gharalii sp. nov. in the wing pattern (hyaline basally, with brown radiate pattern on apical two-thirds, and usually with one large hyaline spot in cell r 1), flagellomere 1 pointed, oviscape short, aculeus relatively blunt, with short acute apex and similar spermathecae shape. It differs from both species by the size of the brown ray in cell dm basal to the level of r-m, which is absent or short, usually reaching only the mid-width of cell dm, at most reaching Cu 1 (long, reaching middle of cell cu 1 in T. maccus and T. gharalii).	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB01077FF05FBBFEDB8F8E2.taxon	materials_examined	Type material. Holotype (female): Iran: Charmahal & Bakhtiari Province, South part of Choghakhor Lake, 2600 m, 31 ° 53 N, 50 ° 58 E, 27. v. 2014 (Mohamadzade) (JAZM). Paratypes: Iran: Kirman, 08. v. 1960, 1 ♀ (Hammad) (NMPC); Alburz, Tochal, 3500 – 3950 m, 30. vi. 1973, 2 ♂ (Loc. No. 262, Exp. Nat. Mus. Praha) (HMIM); Tehran, Evin, 3. vii. 1973, 2 ♀; 06. viii, 1973 2 ♀; 22. viii. 1973, 2 ♀; 27. viii. 1973, 2 ♀; 06. ix. 1973, 1 ♂, 1 ♀ (HMIM); 5 km North East Abali, 35 ° 50.304 ' N, 51 ° 58.980 ' E, 2360 m, 29. viii. 2008, 1 ♀ (Mohamadzade) (SMNC); same collection data as holotype, 08. vi. 2013, 2 ♂, 2 ♀ (Mohamadzade); Kohkiloyeh & Boyerahmad Province, Pass near Sisakht, 3300 m, 30 ° 52 ’ 29 N, 51 ° 31 ’ 26 E, 26. v. 2014, 1 ♀ (S. & V. Korneyev); Charmahal & Bakhtiari Province, Choghakhor Lake, 2290 m, 31 ° 55 ’ 23 N, 50 ° 56 ’ 05 E, 27. v. 2014, 1 ♀ (V. & S. Korneyev) (SIZK).	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB01077FF05FBBFEDB8F8E2.taxon	description	Description. Head (Fig. 21): Yellow, whitish microtrichose, except ocellar triangle, distal 2 / 3 of arista and Vshaped mark on upper part of occiput blackish. Flagellomere 1 yellow. Frontal stripe less distinctly microtrichose. Frons above lunule with 5 – 10 setulae, setulae whitish yellow, brownish on anterior part of gena. Postocular setae and setulae (both longer and shorter) whitish yellow. Length: height: width ratio = 1: 1.1: 1.54. Frons subquadrate, twice as wide as eye, which is about 1.3 times as high as long. Gena 0.55 times as high as length of flagellomere 1. Flagellomere 1 1.6 times as long as wide, pointed on apicodorsal angle. Thorax: Black (only postpronotal lobe narrowly yellow), densely white microtrichose. Setae yellowish brown; posterior notopleural seta whitish yellow. Setulae white; scutellum with 15 – 21 white marginal setulae on each side. Calypteres white. Halter yellow. Legs: Yellow, fore femur with 2 rows of white posterodorsal and one row of brown posteroventral setae; mid and hind legs with brown setae and setulae (Fig. 18). FIGURES 1 – 9. Wing, 1. Tephritis azari, ♂. 2. T. azari, ♀. 3. T. urelliosomima, ♂. 4. T. urelliosomima, ♀. 5. T. gharalii, ♂. 6. T. gharalii, ♀. 7. T. gharalii, ♀. 8. T. maccus, ♀. 9. T. sophus (redrawn from Gentilini et al., 2006). Wing (Figs. 5 – 7): Base hyaline, including all of cells c, bm and bcu; apical portion with brown radiate mark, with few hyaline spots and indentations; pterostigma entirely brown, cell r 1 with only one hyaline spot beyond pterostigma at level of crossvein r-m. This spot in male large, reaching R 2 + 3 (but never extending into cell r 2 + 3), but in 2 females small and narrow, and in 3 females moderately large but not reaching R 2 + 3. Cell r 2 + 3 hyaline between radial fork and level of apex of vein Sc, distally brown, with only 2 apical hyaline spots, posterior spot extending into cell r 4 + 5; cell br hyaline basal to level of apex of Sc, in apical third completely dark, without hyaline spots. Cell r 4 + 5 with round hyaline spot touching vein M at or near level of dm-cu (only one female without this spot [Fig. 7]), and apical spot bordered by 2 narrow apical rays extending to apices of veins R 4 + 5 and M (in 2 females, the ray on vein R 4 + 5 is extremely narrow, the apical spot in cell r 4 + 5 is short, and the apical fork is broadly connected to the main pattern [Fig. 6]). Cell dm hyaline on proximal 1 / 4 brown on apical part with 3 brown rays (including 1 on dmcu) with 1 large hyaline spot at level of crossvein r-m and 1 small round (or pear-shaped) subapical hyaline spot (in one female without smaller spot); both spots extend to Cu 1 and sometimes are fused with spots in cell cu 1. Cell m with 3 large hyaline spots, the most distal extending into cell r 4 + 5; cell cu 1 mostly hyaline, with brown bars in apical half. Vein R 4 + 5 with 4 – 5 setulae ventrally. Abdomen: Black, tergites entirely microtrichose, with white setulae and yellow marginal setae. Oviscape shining black, with whitish setae on basal part; shorter than tergites 5 and 6 combined, its dorsum 1.3 times as long as tergite 6. Aculeus brown, 3 times as long as wide, strongly tapered subapically to short, pointed apex (Figs. 20, 24). Eversible membrane with two pairs of taeniae 0.3 times as long as membrane itself; on ventral side membrane with small scales (Fig. 19). Spermathecae teardrop shaped (Fig. 23) with narrowed neck. Epandrium oval (Fig. 26); glans (Fig. 27) moderately short, mostly membranous. Measurements. Holotype: BL. 3.5, WL. 3.25; Female. BL. 4 – 4.5 mm, WL. 3.5 – 3.7 mm, AL. 0.75 – 0.87 mm; Male. BL. 3.5 – 3.9 mm, WL. 3.2 – 3.6 mm.	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB01077FF05FBBFEDB8F8E2.taxon	diagnosis	Diagnosis. The new species differs from most Palaearctic species of Tephritis by having only one marginal hyaline spot in cell r 1. T. gharalii is closely related to T. azari Mohamadzade & Korneyev and T. maccus Hering in having similar body and wing size and wing pattern (hyaline basally, with brown radiate pattern on apical twothirds, and usually with one hyaline spot in cell r 1, as well as flagellomere 1 pointed, oviscape short, aculeus relatively blunt, with short acute apex, and similar spermathecae shape). It differs from T. azari in having a broad brown ray in cell dm basal to the level of crossvein r-m that reaches the mid-width of cell cu 1. In addition the new species has only one hyaline spot in cell r 1 that does not extend into cell r 2 + 3 (in T. azari, the short brown ray in cell dm basal to the level of r-m is absent or usually reaches only the mid-width of cell dm, at most reaching Cu 1, and cell r 1 in males has two hyaline marginal spots). T. gharalii also differs from T. maccus in having thick brown rays in cell dm that usually are connect to each other along Cu 1. Tephritis gharalii is also similar to T. urelliosomima, in which the aculeus shape is similar (blunt, but pointed at very apex); also in 2 female paratypes the base of the apical fork is broadly connected to the main part of the pattern as in T. urelliosomima. It differs from T. urelliosomima in the wing pattern with a hyaline spot in cell r 1 and absence of brown pattern in basal part of cell dm (in T. urelliosomima the large hyaline spot in r 1 usually is lacking; only in some males one small or rarely large spot is present; cell dm is completely brown with only one round hyaline spot at the level of crossvein r-m). The wing pattern of this new species resembles Euarestella iphionae (Efflatoun, 1924), but the latter genus has only one pair of scutellar setae. E. iphionae was recorded from Iran by Dirlbek (1980) but recent studies by Dr. Valery A. Korneyev and the second author at NMPC and the first author at HMIM proved that this record was based on misidentification of this new species. The record of T. maccus from Iran (Gilasian & Merz 2008) was also based on misidentification of T. gharalii. The misidentified specimens were added to the type series, and these two species are deleted from the list of Iranian fruit flies.	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB01077FF05FBBFEDB8F8E2.taxon	etymology	Etymology. The species is named in honour of Dr. Babak Gharali, an outstanding Iranian dipterist, in recognition of his invaluable contribution to the study of flies (Diptera), especially of the family Bombyliidae in Iran.	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB51076FF05FF6AECEDFC51.taxon	materials_examined	Material examined. Type material. Holotype (female): France: Pyrenees-Orientales: Vernet, in June (ZMHU); Non-type: Iran: 5 km North East Abali, 35 ° 50.304 ' N, 51 ° 58.980 ' E, 2360 m, 29. viii. 2008, 1 ♀ (Mohamadzade) (SMNC); Kyrgyzstan: Alai range, Kichik-Alai ridge, Isfairam-Sai basin, 45 km S of Kyzyl-Kiya, 1800 – 3000 m a. s. l., 11. vii. 1999, 1 ♀ (Korneyev & Kameneva) (SMNC); Talas Ridge, h = 1500 – 2000 m, Kara-Bura valley, 43 km from Kok-Sai, 42 ° 23.0 ' N 71 ° 37.6 ' E, 28. vi. 1998, 1 ♀ (Korneyev & Kameneva) (SIZK); Alai, Kichik-Alai ridge, Isfairam-Sai basin, 45 km S of Kyzyl-Kiya, 1800 – 3000 m a. s., 05. vii. 1999, 1 ♀; 08. vii. 1999, 1 ♂; 11. vii. 1999, 1 ♂, 1 ♀; 30. vii. 1999, 1 ♀; 31. vii. 1999, 1 ♂ (Korneyev & Kameneva) (SIZK).	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB51076FF05FF6AECEDFC51.taxon	distribution	Distribution. Spain, France, Italy, Kyrghyzstan and Afghanistan (Norrbom et al. 1999).	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB51076FF05FF6AECEDFC51.taxon	description	Measurements. Holotype: WL. 3.9 mm, AL. 0.72 mm.	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFB51076FF05FF6AECEDFC51.taxon	diagnosis	Diagnosis. Tephritis maccus is similar to T. gharalii and T. azari which have similar wing patterns, pointed flagellomere 1, short oviscape, relatively blunt aculeus, with short acute apex, and similar spermathecae shape. T. maccus differs in that the brown ray of cell dm basal to the level of r-m is narrow and reaches the middle of cell cu 1, whereas in T. azari this ray is absent or usually reaches only the mid-width of cell dm, at most reaching Cu 1 (Fig. 8), and in T. gharalii this ray is thick and penetrates into cu 1. In addition, in T. maccus and T. azari cell r 1 has two hyaline spots beyond pterostigma in males (the larger spot penetrates into r 2 + 3) and one hyaline spot in females, but in T. gharalii both sexes have only one hyaline spot in cell r 1 that at most reaches to R 2 + 3. Detailed studies of the specimens collected in Kyrgyzstan shown that they belong to T. maccus. The structure of the ovum is similar to other Tephritis species.	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFBA1079FF05FF6AEE72FDE5.taxon	distribution	Distribution. Italy (Gentilini & Korneyev 2006).	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFBA1079FF05FF6AEE72FDE5.taxon	discussion	Comments. T. sophus is a fossil species. It is similar to T. azari, T. maccus and T. gharalii in having radiate wing pattern, and similar aculeus shape, but like T. azari, the short brown ray in dm cell basal to the level of r-m reaches only the mid-width of cell dm or Cu 1 (whereas this ray reaches the middle of cell cu 1 in T. maccus and T. gharalii). T. sophus also differs from T. azari in having the brown area around the apex of R 2 + 3 with a broad Ushaped spot (including the small hyaline spot at R 2 + 3 apex) darker than the remaining pattern (Fig. 9). Both male and female with 2 hyaline spots in cell r 1 (Gentilini et al., 2006: Figs. 30 – 33, 36 – 38). The oviscape in this species is about as long as the preceding three abdominal tergites.	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFBA1079FF05FD51ECE7F84B.taxon	materials_examined	Material examined. Type material: Paratype, Turkmenistan, 1 ♀, 10 km S of Ashhabat, 23 – 27.04.1987 (Antropov) (SIZK); Allotype male: Iran: Yazd province, Manshad, 2150 m, 31 ° 32 ’ 07 N, 54 ° 13 ’ 70 E, 06. v. 2010 (Mohamadzade); Non-type, Iran: 1 ♀, Khorasan Province, 1900 m; 1 ♂, Tehran Province, Tehran, Evin, 16. v. 1970 (L. T.); 1 ♂, 1 ♀, Fars Province, Bamu, 1700 m, 24. ix. 1996 (Parch., Mogh. & Bar.) (HMIM); 1 ♂, Yazd province, Manshad, 2150 m, 31 ° 32 ’ 07 N, 54 ° 13 ’ 70 E, 06. v. 2010 (Mohamadzade).	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFBA1079FF05FD51ECE7F84B.taxon	distribution	Distribution. Iraq, Turkmenistan (Korneyev & Dirlbek 2001) and Iran (Gilasian & Merz 2008).	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFBA1079FF05FD51ECE7F84B.taxon	description	Male description. Head Coloration as in T. gharalii. Length: height: width ratio = 1: 1.14: 1.57. Frons subquadrate, 1.6 as wide as eye, which is about 1.4 times as high as long. Gena 0.55 times as high as length of flagellomere 1. Flagellomere 1, 1.5 times as long as wide, pointed at apicodorsal angle. Thorax: Coloration of thorax and legs as in T. gharalii. Wing (Fig. 3): Base hyaline, including all of cells c, bm and bcu; apical portion with brown radiate mark, with few hyaline spots and indentations; pterostigma dark brown, cell r 1 with one hyaline spot beyond pterostigma at level of crossvein r-m and reaching to vein R 2 + 3 (in one male with small spot that reaches only mid-width of cell r 1 and in another male without real spot and with lightening beyond pterostigma; in females this spot is usually absent, rarely with one small hyaline spot). Cell r 2 + 3 brown with small hyaline dot distal to cell R 2 + 3 apex and one apical spot, isolated from apical spot in cell r 4 + 5 by narrow ray; cell br hyaline proximal to radial fork, in apical half completely dark, without hyaline spots. Cell r 4 + 5 with round hyaline spot touching vein M between r-m and dm-cu, and apical spot wider than long, bordered by 2 narrow apical rays. Cell dm mostly dark brown, with narrow hyaline border at crossvein bm-cu and with one hyaline spot at level of r-m and small hyaline dot at apex. Cell m with 3 large hyaline spots, cell cu 1 brown on anterior half. In one specimen apical ray on R 2 + 3 completely reduced and ray on R 4 + 5 present only as isolated brown spot at end of vein. Vein R 4 + 5 with 4 – 5 setulae ventrally. Abdomen: Black, tergites entirely microtrichose, with white setulae and yellow marginal setae. Oviscape shining black, with whitish setae on basal part; shorter than tergites 5 and 6 combined, its dorsum 1.3 times as long as tergite 6. Aculeus brown, 3 times as long as wide, rapidly tapered subapically to short, pointed apex (Figs. 35 – 38). Eversible membrane with two pairs of taeniae 0.3 times as long as membrane itself; on ventral side membrane with small scales (Fig. 34). Spermathecae teardrop shaped (Fig. 37) with narrowed neck. Epandrium oval, (Fig. 39); glans short, mostly membranous (Fig. 36). Measurements. Female. BL. 4.3 – 4.7 mm, WL. 3.7 – 4 mm, AL. 0.85 mm; Male. BL. 3.7 – 4 mm, WL. 3.5 – 3.8 mm.	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
03B987E6FFBA1079FF05FD51ECE7F84B.taxon	diagnosis	Diagnosis. Tephritis urelliosomima is recognizable from all other species of the genus by absence of large hyaline spots in cell r 1 beyond pterostigma or only one small or large hyaline spot is present (Figs. 3 – 4). In addition the base of the apical fork is broadly connected to the main pattern; and cell dm is dark with one hyaline spot at the level of crossvein r-m.	en	Namin, Saeed Mohamadzade, Korneyev, Severyn V., Parchami-Araghi, Mehrdad, Gilasian, Ebrahim (2015): Revision of the Tephritis maccus species group (Diptera: Tephritidae) with description of a new species from Iran. Zootaxa 3956 (4): 589-599, DOI: 10.11646/zootaxa.3956.4.10
