identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
307B0A93379D5E00B002F58EB30B6027.text	307B0A93379D5E00B002F58EB30B6027.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leea compactiflora Kurz	<div><p>Leea compactiflora Kurz, J. Asiat. Soc. Bengal, Nat. Hist. 42 (2): 65. 1873</p><p>Figs 4, 5</p><p>Leea compactiflora Kurz, J. Asiat. Soc. Bengal, Nat. Hist. 42 (2): 65. 1873, Forest Fl. Burma 1: 279. 1877; C. B. Clarke in Trimen’s J. Bot. 19: 163. 1881; Ridsdale, Blumea 22: 88. 1974; J. Joseph, Fl. Nongpoh and vicinity 72. 1982; Harid. &amp; R. R. Rao, Forest Fl. Meghalaya 248. 1985; A. S. Chauhan, Contr. Fl. Namdapha 140. 1996; Hajra et al., Mat. Fl. Arunachal Pradesh 1: 318. 1996; B. D. Naithani in N. P. Singh et al. (Ed), Fl. India 2000; R. Shanpru in N. P. Singh et al. (Eds), Fl. Manipur 1: 240. 2000; N. P. Singh et al., Fl. Bihar, Analysis 114. 2001; T. K. Sarma &amp; A. K. Sarkar, Fl. Palamau Dist. 156. 2002; K. P. Singh in N. P. Singh et al. (Eds), Fl. Mizoram 1: 388. 2002; G. D. Pal, Fl. Lower Subansiri Dist. 1: 215. 2103, excl. syn. L. trifoliata; B. K. Shukla in K. P. Singh et al. (Eds), Fl. Uttar Pradesh 1. 351. 2016, excl. syn. L. trifoliata; S. S. Dash &amp; P. Singh, Fl. Kurung Kumey Dist. 384. 2017, excl. syn. L. trifoliata; Pusalkar &amp; S. K. Srivast., Fl. Uttarakhand 1: 989. 2018. Type: INDIA. Nagaland: Naga hills, 3 Nov 1836, Griffith Herb. Propr. N. 1297 (neotype, K K 004252754, here designated) .</p><p>Leea bracteata C. B. Clarke, Trimen’s J. Bot. 19: 164. 1881; Kanjilal &amp; Das, Fl. Assam 1 (2): 308. 1936; Deb, Fl. Tripura 1: 419. 1981; A. K. Mukh., Pachmarhi &amp; Bori Reserves 68. 1984. Type: INDIA. Sikkim, s. d., Hook. f. &amp; Thomson 169 (lectotype, designated by Ridsdale 1974, pg. 88: K [K 001615004]) .</p><p>Description.</p><p>Erect subshrub or shrub, 1–2 m high; roots non-tuberous. Stems erect or straggling, stout, jointed, with branches, young stem hispid or pubescent; internodes furrowed or round, green; nodes not swollen; pearl glands globose, more on inflorescences, white. Leaves alternate, pinnate to bipinnate, imparipinnate; petioles 7–21 mm long, base bulged, sheathing the stem, glabrous, green. Petiolar stipules narrowly winged, 4–8 cm long, extending to half of petioles; stipular scars triangular; rachises 5–31 cm long, channeled on upper surface, hispid, green. Leaflets opposite, ovate widely ovate or obovate, distal leaflets: lamina 14–32 × 6.5–13.5 cm, lateral leaflets: lamina 7.5–30 × 4–13 cm, glabrous, young leaves green or brown, mature leaves green on upper surface and pale green on lower, base cuneate to rounded, margins serrate, apex long acuminate or caudate; petiolules: central 3–8 cm long, green, lateral 0.4–2.5 cm long; secondary veins 7–16 pairs, 3 - nerved at base, vein raised on abaxial side. Inflorescences terminal, a leaf-opposed compound cyme, 8–22 cm long, many flowered, condensed, hispid; peduncles 1–4 cm long, furrowed, pubescent, green, hispid. Flowers 4–6.2 mm, actinomorphic, greenish with brown tinges. Pedicels slender, 1.5–2 mm long, hispid. Bracts 3.6–8.3 × 1.9–5 mm, broadly ovate, conspicuous, persistent, green, hispid. Calyx campanulate, 3.5–5.6 × 3–3.3 mm, 5 lobed, lobes 0.9–1.1 × 0.75–1.9 mm, ovate or widely ovate, apex acute, green, hispid; corolla lobes 5, ovate-oblong, 2.7–3.5 × 1.9–2.4 mm, apex acute, hooded, margin entire, glabrous, green; corolla tube with floral disc 3.1–3.3 × 3–3.2 mm. Floral disc 1.6–2.4 mm long, orange-red; upper free part 1.1–1.8 mm long, notched; sinuses shallow, 0.25–0.4 mm long; lower free part 0.8–1 mm long. Stamens 5, syngenecious; filaments 1.25–1.3 mm long, glabrous, white; anthers 1.2–1.3 mm long, dehiscing longitudinally, purple on backside, creamish on margin. Ovary globose, 0.95–1.1 mm long, superior, 4–6 - loculed, one ovule in each locule, placentation axile; style slender, 1.1–2.2 mm long, glabrous, creamy; stigma round. Fruits a berry, yellow when young, turning purple black at maturity, 6 - seeded. Seeds wedge-shaped, 4.72–4.91 × 3.72–3.85 mm, light brown when mature.</p><p>Flowering and fruiting.</p><p>May – November.</p><p>Habitat.</p><p>This species is usually found growing near roadsides and forest margins. The common associates are Elatostemma sp., Boehmeria glomerulifera Miq. (both Urticaceae), Chromolaena odorata (L.) R. M. King &amp; H. Rob. ( Asteraceae) and some ferns and grasses.</p><p>Distribution.</p><p>India (Arunachal Pradesh, Assam, Bihar, Manipur, Meghalaya, Mizoram, Sikkim, Uttarakhand, Uttar Pradesh and West Bengal), Bhutan, China, Laos, Myanmar, Nepal and Vietnam. Fig. 6.</p><p>Specimens examined.</p><p>China. Yunnan province: southern Yunnan between Keng Hung [Jinghong] and Muang Hing [Meng Hai], 02-25-1922 - 01-03-1922, J. F. Rock 2642 (US 1213169!) . India. • Arunachal Pradesh: Anjaw district, Hayuliang-Borpu, 25 Nov 1957, R. S. Rao 10819 (CAL) ; • Changlang district, Namdapha National Park, 05 Aug 2018, Amrutha 159829 (CALI) ; • East Siang district, Rengging, 24 Feb 1912, I. H. Burkill 36711 (CAL) ; • Rotung, 31 Dec 1911, I. H. Burkill 37599 (CAL) ; • Lohit district, Way to Tidding from Tezu, 10 Aug 2018, Amrutha 159836 (CALI) ; • Lower Subansiri district, Innerline beyond Kimin River up to 1 mile along the road to Ziro, 23 Nov 1957, G. Panigrahi 11469 (CAL) ; • Way to Raga, 16 Aug 2018, Amrutha 159850 (CALI) ; • Papum Pare district, 5 km from Chimpu, 03 Aug 1994, G. D. Pal 6709 (ARUN) ; • Itanagar-Hydel, 25 Sept 1978, G. D. Pal 76051 (ARUN) ; • Senki view, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.61061&amp;materialsCitation.latitude=27.103806" title="Search Plazi for locations around (long 93.61061/lat 27.103806)">Itanagar</a>, 27°06'13.7"N, 93°36'38.2"E, 300 m, 30 May 2022, Amrutha 164710 (CALI) ; • Tirap district, Lai Longsong, 19 June 1961, D. B. Deb 25781 (CAL) ; • Rusa-Bimalpur, 07 Sept 1958, G. Panigrahi 17025 (CAL) ; • Upper Siang district, Way to Hawa Camp, Yingkiong, 02 Aug 2018, Amrutha 159865 (CALI) ; • West Kameng district, Sessa Village, 01 June 2022, Amrutha 164722 (CALI) ; • West Siang district, Kane WLS, 13 Sept 2008, S. S. Dash 32228 (ARUN) ; • Assam: Darrang district, Darrang, 22 Apr 1914, Upendranath Kanjilal s. n. (ASSAM) ; • Meghalaya: East Khasi Hill district, Khasi hills, 30 Oct 1872, Clarke, C. B. 17726 F (K 004252751) ; • Ibid., 0-4000 ft, s. d., J. D. Hooker &amp; J. J. Thomson s. n. (K K 004252741, K 004252745, K 004252749); • Ri-Bhoi district, Balaiba Tilla, Nongpoh, 12 May 1965, J. Joseph 42328 (ASSAM) ; Nongpoh, 2000 ft, 02-15, May 1945, Thakur Rup Chand 1504 (MICH 1506767; L 0532034) ; • Ibid., 2000 ft, 16 Jun 1949, Thakur Rup Chand 22982 (MICH 1006084 A &amp; B; L 0532033); Ibid., 13 May 1914, Upendranath Kanjilal 3977 (ASSAM); • Sohra, 4000 ft, 24 May 1952, Walter N. Koelz 30039 (MICH 1506768) ; • West Garo Hills district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=90.20066&amp;materialsCitation.latitude=25.494917" title="Search Plazi for locations around (long 90.20066/lat 25.494917)">1 Km after entry gate, foothills of Tura Peak</a>, 25°29'41.7"N, 90°12'02.4"E, 03 June 2022, Amrutha 164735 (CALI) ; • Rongsingiri, Nokrek Biosphere Reserve, 17 Oct 2007, V. N. Singh &amp; Bikarma Singh 115870 (ASSAM) ; • Mizoram: Aizawl district, Seling, Aizawl Seling-Champhai Road, 800 m, 07 June 2022, Amrutha 164748 (CALI) ; • Nagaland: Naga hills, 17 Oct 1885, Clarke, C. B. 40845 A (K K 004252750) ; Jabocka, Naga Hills, Apr 1899, Dr. Prains Collector 907 (CAL) ; • Sikkim: s. d., Kurz s. n. (CAL); • Ibid., 11 Jan 1874, G. King 839 (CAL); • Ibid., 24 Aug 1875, King, G 2324 (L 0744659); • Ibid., Regio trop., 1-6000 feet, J. D. Hooker s. n. (K 004252736) ; • Ibid., 1-5000 feet, J. D. Hooker s. n. (K 004252729, K 004252730, K 004252731, K 004252732, K 004252735, K 004252758; P 02342891); • Sikkim Himalaya, 1881, G. King s. n. (US 01151425!) ; Gangtok district, Singtam, 18 May 1967, N. C. Mazumder &amp; R. M. Dutta 269 (CAL) ; • West Bengal: Darjeeling district, Rishop, 5 June 1870, Clarke, C. B. 11824 a, (K K 004252737) ; Jalpaiguri district, Dayamara, Jaldapara National Park, 09 May 2014, K. Karthigeyan 61470 (CAL barcode CAL 0000209976) ; • Rungbee, 8 Oct 1908, W. W. Smith 282 (CAL) . Laos. Sekong province, shrub in forest, 28 Apr 2013, fl, H. Sun et al. 14995 (KUN) .</p><p>Conservation status.</p><p>In India, this species is found in Northeast India, especially in Arunachal Pradesh. The Extent of Occurrence (EOO) is calculated to be 1417, 687 Km 2, and the Area of Occupancy (AOO) is 164.00 Km 2. They are mostly seen in forest margins and on the roadside. Because of its number of populations and localities, this species can be considered as Least Concern (LC) according to the IUCN Standards and Petitions Committee (2022).</p><p>Notes.</p><p>Leea compactiflora was first described by Kurz (1873) with a very short diagnosis, and he noted that the specimen was collected from Martaban, Myanmar. After an exhaustive search in various herbaria, we could not locate the sheet of Kurz from Martaban. Ridsdale (1974) also indicated that he did not trace the material. Therefore, a neotypification is required as per Art. ICN 9.8. Here, we select a specimen from the Naga hills, India, collected by Griffith, housed at the Kew herbarium (K 004252754) as the neotype following the Madrid Code (Turland et al. 2025), as it matches the protologue and was cited by Clarke (1881) in his revision of Indian species of Leea .</p></div>	https://treatment.plazi.org/id/307B0A93379D5E00B002F58EB30B6027	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Athalappil, Amrutha;Pradeep, Ayilliath Kuttiyeri;Wen, Jun;Krishna, Nikhil	Athalappil, Amrutha, Pradeep, Ayilliath Kuttiyeri, Wen, Jun, Krishna, Nikhil (2025): Resolving the taxonomy of Leea trifoliata (Leeaceae, Vitales): Reinstatement and typification. PhytoKeys 266: 281-295, DOI: 10.3897/phytokeys.266.173605
C3642DC9854E576AA5737DE313107501.text	C3642DC9854E576AA5737DE313107501.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leea trifoliata M. A. Lawson, Hook. f.	<div><p>Leea trifoliata M. A. Lawson, Hook. f. (Ed), Fl. Brit. India 1 (3): 666. 1875.</p><p>Figs 1, 2</p><p>Leea trifoliata M. A. Lawson, Hook. f. (Ed), Fl. Brit. India 1 (3): 666. 1875; C. B. Clarke, Trimen’s J. Bot. 19: 163. 1881; Kanjilal &amp; Das, Fl. Assam 1 (2): 305. 1936. Type: INDIA. Assam, s. d., Jenkins &amp; c. s. n. (lectotype, designated by Ridsdale 1974, pg. 88: K [K 000736385]; isolectotype: K [K 000323729]) .</p><p>Description.</p><p>Perennial weak herb, 0.8–1 (– 2.5) m high; roots non-tuberous. Stems erect or wiry, jointed, warted, glabrous; internodes furrowed, green; nodes swollen; pearl glands globose, white. Leaves alternate, trifoliate or 1 - pinnate with two rudimentary pairs of leaves at base; petioles 7–21 mm long, slightly bulged, sheathing the stem, hairy at base, green. Petiolar stipules narrowly winged, expanding the full length of the petiole, 2–2.5 cm long; stipular scars triangular; rachises 1–4 cm long, channeled and slightly winged on upper surface, hispid on margin, green. Leaflets opposite, elliptic oblong or obovate, distal leaflets lamina: 5.5–13 × 2.7–7.5 cm, lateral leaflets lamina: 4.5–12.5 × 2–6 cm, one row of stiff hairs in between nerves on upper surface, mature leaves green on upper surface and pale green on lower, base cuneate, margins serrate, apex cuspidate caudate, abruptly acuminate; petiolules: central 1–2.5 cm long, lateral 0.2–0.7 cm long, channeled and winged on upper surface, green; secondary veins 7–16 pairs, 3 - nerved at base, strigose on lower surface, vein raised on abaxial side. Inflorescences terminal or leaf-opposed, compound cyme, up to 4 cm long, axis hispid. Bracts 1.8–2.7 mm long, lanceolate-linear, pubescent. Calyx campanulate, c. 3.5 mm long, 5 lobed, each lobe c. 1–1.5 mm long, apex acute; corolla lobes 5, 2.7–2.8 × 1.1–1.3 mm, slightly hooded at apex, margin entire. Floral disc 1.1–1.5 mm long. Ovary 4–6 locular. Fruits noted as black in Lawson (1875).</p><p>Flowering and fruiting.</p><p>June – December.</p><p>Habitat.</p><p>This species mostly grows on roadsides and forest margins. They grow along with Adiantum sp. ( Pteridaceae), Impatiens minor (DC.) Bennet, Impatiens balsamina L. (both Balsaminaceae) and some grasses.</p><p>Distribution.</p><p>Bhutan, India (Assam &amp; Arunachal Pradesh). Fig. 3.</p><p>Specimens examined.</p><p>Bhutan. s. d., Booth s. n. (K 000323730) . India. • Arunachal Pradesh: Mishmi hills, s. d., Griffith 1344 (K 004252755, P 02342863) ; • East Kameng district, Pakke Wildlife Sanctuary, 25 Apr 2012, B. B. T. Tham 35028 (ARUN) ; • Kamle district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.07491&amp;materialsCitation.latitude=27.799833" title="Search Plazi for locations around (long 94.07491/lat 27.799833)">Raga</a>, 27°47'59.4"N, 94°04'29.7"E, 16 Aug 2018, Amrutha 159851 (CALI) ; • Assam: s. d., Masters s. n (K 004252752); • Cachar district, Bhuban Hills, Oct 1978, 74142 (ASSAM) ; • Cachar, June 1874, R. L. Keenen s. n. (K 004252761) ; • Darrang district, Aka Hills, s. d., N. L. Bohr 17378 (ASSAM) ; • Dibrugarh district, Dibru Reserve, eastern part, 20 Oct 1960, G. Panigrahi 21564 (ASSAM) ; • Golaghat district, s. loc, 300 ft, May 1891, Dr. King’s collectors s. n. (US 01151426, K 004252756, P 02342865, L 0744660); • Jorhat district, Gibbon Sanctuary, Camp- 2, 12 June 2010, Ranjit Daimary 122699 (ASSAM) ; • Ibid., Camp- 4, 26 Feb 2011, Ranjit Daimary 121691 (ASSAM); • Kamarup district, Moyang Residence, 09 Apr 1915, U. Kanjilal 5449 (ASSAM) ; Kamarup Metropolitan district, Guwahati, April 1902, A. C. Chatterjee (P 02342864, LY 0106862, MPU 703118) ; • Karbi Anglong district, • Garampani Wildlife Sanctuary, 24 Nov 1975, P. K. Hajra 65058 (ASSAM) ; Kaliani, 18 June 1963, D. B. Deb 35116 (ASSAM) ; • Nagaon district, Laopani, 28 Aug 1978, P. Chakraborty 78984 (ASSAM) ; • Sonitpur district, Batasipur, 13 May 1947, M. M. Srinivasan 22253 (ASSAM) ; • Kanchanjuri, Kazhiranga, 12 Aug 1973, P. K. Hajra 52836 (ASSAM) ; • Tinsukia district, Margherita Forest, 22 June 1938, G. K. Duka 17010 (ASSAM) .</p><p>Conservation status.</p><p>Leea trifoliata is restricted to two northeastern Indian states, Assam and Arunachal Pradesh, and one location in Bhutan. This species prefers to grow in open areas adjoining evergreen forests. The Extent of Occurrence (EOO) is calculated to be 114,777 km 2, and the Area of Occupancy (AOO) is 84 km 2. Considering its large Extent of Occurrence, this species does not fall into any threat category. Despite this, the Area of Occupancy is small, and each population has fewer than five mature individuals. Due to its narrow range of distributions and small AOO, this species could be categorized at present as Endangered (EN), B 2 b (ii, iii) according to the IUCN Standards and Petitions Committee (2022).</p><p>Notes.</p><p>In the protologue, Lawson (1875) described the anthers as distinct, and the fruits were black.</p><p>Ridsdale (1974) typified the name L. trifoliata by selecting the Jenkins &amp; c. material from Assam as the lectotype. We were able to trace two sheets of Jenkins &amp; c. from the Kew herbarium (K 000736385 &amp; K 000323729). However, Ridsdale has attached a slip to the sheet with barcode K 000323729 stating that it is the lectotype. Hence, the other sheet (K 000736385) becomes the isolectotype.</p></div>	https://treatment.plazi.org/id/C3642DC9854E576AA5737DE313107501	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Athalappil, Amrutha;Pradeep, Ayilliath Kuttiyeri;Wen, Jun;Krishna, Nikhil	Athalappil, Amrutha, Pradeep, Ayilliath Kuttiyeri, Wen, Jun, Krishna, Nikhil (2025): Resolving the taxonomy of Leea trifoliata (Leeaceae, Vitales): Reinstatement and typification. PhytoKeys 266: 281-295, DOI: 10.3897/phytokeys.266.173605
