identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
015C614EF40AFF995C94F9C5036EFCB1.text	015C614EF40AFF995C94F9C5036EFCB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Roncocreagris cavernicola (Vachon 1946) Vachon 1946	<div><p>Roncocreagris cavernicola (Vachon, 1946)</p><p>Microcreagris cavernicola Vachon 1946: 333 –334, figs 1–6. Roncocreagris cavernicola (Vachon): Mahnert 1976: 212.</p><p>Material. We have not examined the type specimens described by Vachon (1946) from Portugal, district of Leiria, Abiúl, Algar sul das Corujeiras (male) and Santiago da Guarda, Algar da Lapa (female), but Dr Mark Judson (MNHN), at our request, has kindly checked the most important diagnostic characters of the species, mentioned below. Recent searches for R. cavernicola at the type localities have been unsuccessful. In the case of the Algar da Lapa locality, the cave can no longer be found and the cave entrance was probably destroyed by building construction in the area (Grupo Protecção Sicó, pers. com.).</p><p>Diagnosis. No eyes or eye-spots. Strong troglomorphic adaptations. Carapace with 18 setae, of which 4 on posterior margin. Tergite I 4 setae. Galea short in both sexes, simple in male and with tiny apical rami in female. Pedipalp: femur ratio 5.9 in male and 6.9 in female, femur distinctly shorter than the chelal movable finger; chela+ ratio about 6.0 in male and 7.0 in female; ratio movable finger/hand+ 1.7 in both sexes; ratio trichobothria ib–ist / ist–it 1.1–1.3, ratio isb–ist / ib–isb 0.5–0.7.</p><p>Remarks. The following data are based on Vachon’s original description (1946) with additions or modifications from examination of the syntypes (3 MNHN Ps620-291, on 3 slides, Ƥ MNHN Ps620-292 on 2 slides).</p><p>Male, female in square brackets. Carapace with 4 posterior setae in both sexes. Chaetotaxy of tergites I–IV: 4:7:7:7 [4:6:6:7]. Manducatory process with 3 setae on each side in both sexes. Trichobothrial ratio ib–ist / ist–it 1.1 [1.3]; ratio isb–ist / ib–isb 0.5 [0.7]. Pedipalp measurements: femur 1.42/0.24 (5.9) [1.59/0.24 (6.9)]; patella 1.40/ 0.29 (4.9) [1.62/0.27 (6.0)], pedicel 0.36 [0.41], club 1.04/0.29 (3.6) [1.21/0.27 (4.5)], ratio club/pedicel 2.9 [3.0]; chela+ 2.45/0.42 (5.8) [2.76/0.40 (6.9)]; hand+ 0.94 (2.2) [1.05/0.4 (2.6)]; movable finger 1.59 [1.77]; ratio movable finger/hand+ 1.7 [1.7]; finger/femur 1.1 [1.1]; femur/patella 1.0 [1.0]; patella/hand+ 1.5 [1.5].</p><p>Vachon’s (1946) description stated that in R. cavernicola the manducatory process bears only 2 setae, which would be quite exceptional in the family Neobisidae (Harvey &amp; Volschenk 2007). Zaragoza (2000, 2002) also used this character for taxonomic comparisons. However, Vachon’s observation is incorrect because 3 setae are present on the manducatory process of both of the type specimens (M. Judson, in litt.).</p></div>	https://treatment.plazi.org/id/015C614EF40AFF995C94F9C5036EFCB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Reboleira, Ana Sofia P. S.;Zaragoza, Juan A.;Gonçalves, Fernando;Oromí, Pedro	Reboleira, Ana Sofia P. S., Zaragoza, Juan A., Gonçalves, Fernando, Oromí, Pedro (2013): On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species. Zootaxa 3670 (2): 283-299, DOI: 10.11646/zootaxa.3670.2.11
015C614EF409FF9D5C94F895061FFCBA.text	015C614EF409FF9D5C94F895061FFCBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Roncocreagris blothroides (Beier 1962) Beier 1962	<div><p>Roncocreagris blothroides (Beier, 1962)</p><p>(Figs 2–11)</p><p>Microcreagris blothroides Beier 1962: 25 –26, fig. 1.</p><p>Roncocreagris blothroides (Beier): Mahnert 1976: 212.</p><p>Material. Portugal, Sicó massif, district of Leiria, three caves: Santiago da Guarda, Gruta da Cerâmica (39º55ʹ36.57ʺN, 8º31ʹ0 3.63ʺW; 355 m. a.s.l.), 29.VIII.2009, 1 3 (DZUL), 28.XI.2009, 6 Ƥ (2 DEUA, 1 DZUL, 1 MNHN, 1 MNCN, 1 SR), 1 3 (DEUA), 21.III.2010, 3 Ƥ (1 DZUL, 1 MCNB, 1 SR), 27.XII.2010, 2 Ƥ (1 DEUA, 1 SR), 1 3 (SR), 10.VI.2011, 2 Ƥ (1 DEUA, 1 DZUL), 1 3 (DEUA); Redinha, Gruta de Santa Maria da Estrela (39º55ʹ41.15ʺN, 8º32ʹ59.4ʺW; 380 m. a.s.l.), 8.III.2009, 2 Ƥ (1 DEUA, 1 DZUL), 1 3 (DEUA); 11.VI.2009, 2 Ƥ (1 DEUA, 1 SR); 28.XII.2009, 1 3 (DEUA); Abiúl, Algar da Confraria I (39°55ʹ51.01"N, 8°31ʹ35.64"W; 420 m. a.s.l.), 22.VII.2007, 1 Ƥ (DEUA). All specimens lgt. A.S.P.S. Reboleira.</p><p>Diagnosis. No eyes or eye-spots. Extreme troglomorphic adaptations. Carapace usually with 18 setae, of which 4 on posterior margin. Tergite I with 4 setae. Galeal chelicera short in both sexes, with 3–5 tiny apical rami. Pedipalp: femur ratio about 8.0, femur roughly same length as the patella and almost as long as the movable chelal finger; chela+ ratio about 8.0 in males, 7.0– 7.8 in females; chelal hand maximum width distal of middle, ratio movable finger/hand+ about 1.7; trichobothrium ist about halfway between ib and it, ratio ib–ist / ist–it 0.9–1.2; ratio isb–ist / ib–isb 0.8–1.0.</p><p>Description. Males, followed by females in square brackets. Large species (Fig. 2). Opisthosoma and legs yellowish. Carapace, chelicerae and pedipalps pale brown.</p><p>Carapace markedly longer than broad (Fig. 3). Without eyes or eye-spots. Anterior margin moderately prominent medially, with blunt, low epistome with some tiny denticles (Fig. 4). Chaetotaxy: 18–19 setae, formula 4:4:6:4–5. Four microlyrifissures in ocular zone, two between median and posterior zones.</p><p>Coxal area. Manducatory process with 3 setae. Anterior process of coxa I long and apically pointed; medial process straight, with some low tubercles (Fig. 5). Pedipalpal coxa with 7 setae, pedal coxa I 5–7, II: 5–6, III: 3–4, IV: 6.</p><p>Tergal chaetotaxy I–X: 3–5:6:6–7:7–8:7:7:7:7:7:7 [4–6:6–7:6–7:7–8:7–8:7–10:7–8:7–8:6–8:7]. Male genital area with 10–11 setae on sternite II; sternite III with 11–16 setae, 6 of them along posterior margin of genital opening. Female genital area with 5–6 microsetae on sternite II, sternite III with 8–10 setae. Sternal chaetotaxy IV– X: 7–8:11:11–12:11–12:10–11:9–10:9 [6–7:10–11:10–11:11–13:10–11:10:9], each sternite VI and VII with 2 discal setae (included in sternal formula). Segment XI: 9–10 setae, including 6 TS. Chaetotaxy of stigmata of sternites III and IV 1–3 and 1–3 respectively. Anal cone with two dorsal and two ventral setae.</p><p>Chelicerae (Figs 6–7). Palm with 6 setae, subgaleal seta 0.63–0.68 from base of movable finger. Galea short (length 0.04 mm), apically with 3–5 short rami, about same length and shape in both sexes. Fixed finger with 10– 15 medium or small teeth, two medial teeth of them larger than the others; movable finger with 8–13 teeth, one large and blunt medial tooth, the others medium or small, dental row ending proximad to subgaleal seta. Rallum with 7–8 blades, all unilaterally pinnate on anterior face, basal blade about half length of others. Serrula exterior with 33–36 blades, serrula interior 27–31 blades.</p><p>Pedipalps (Figs 8–10). Trochanter, femur, distal third portion of the patella and chelal hand with low granulation, more pronounced on paraxial faces. Lyrifissures as in Figs 8–10. Femur with one tiny tubercle distad of middle on antiaxial face, 2–3 distal glandular pores present. Patella dorsally with one micropore at base of pedicel and 1–2 distal glandular pores. Chelal hand dorsally more or less parallel-sided, maximum width distad of middle; pedicel dorsal bears 2–3 dorsal micropores; antiaxial face with an irregular row of 3–5 glandular pores close to base of finger. Fixed finger with 139–148 teeth, most of them apically cusped, dental row reaching to level of trichobothrium esb; nodus ramosus short, reaching level of 3–5th distal teeth; trichobothrium it proximad to et, about half way between est and et; trichobothrium ist closer to the base of finger than to apex; distance between trichobothria ib and ist 0.9–1.2 times that between ist and it; distance between trichobothria isb and ist 0.8–1.0 times that between isb and ib. Chelal microsetae pattern (as defined by Zaragoza 2008): all groups present, Em: 4– 6 curved setae (same length as normal setae), Mm: 2–3, Im: 1. Movable finger with 132–141 teeth, most of them apically cusped, dental row shorter than on fixed finger, ending just distad of trichobothrium b; distance between trichobothria sb and st 1.3–1.8 times that between sb and b. One sensillum near tip of both fingers; one diploid sensillum pc close to dental margin and not raised, close to or slightly distad of trichobothrium sb.</p><p>Legs. Claws of legs I and IV with a tiny dorsal tooth proximad of middle (Fig. 11), subterminal setae with 3–4 rami (Fig. 12). Leg IV tibia with four long setae, TS ratio 0.33–0.49 for longest, basitarsus TS: 0.12–0.14, telotarsus TS: 0.21–0.38.</p><p>Measurements and ratios. Males, followed by females in square brackets. Body 3.08–3.1. Carapace 1.0–1.2/ 0.7–0.79 (1.4–1.5) [1.16–1.22/0.79–0.87 (1.4–1.5)]. Chelicera: palm 0.65–0.72/0.3–0.4 (2.1) [0.7–0.73/0.35–0.37 (2.0)], movable finger 0.41–0.45 [0.46–0.49]. Pedipalp: trochanter 0.93–1.02/0.23–0.24 (4.0–4.3) [1.02–1.04/ 0.24–0.26 (4.0–4.3)]; femur 1.84–2.12/0.23–0.26 (8.0–8.2) [2.0–2.09/0.29–0.3 (7.6–8.1)]; patella 1.75–2.1/0.26– 0.3 (6.8–7.1) [2.02–2.09/0.29–0.3 (6.8–7.2)], pedicel 0.44–0.5 [0.49–0.51], club 1.31–1.6/0.26–0.3 (5.1–5.4) [1.53–1.59/0.29–0.3 (5.1–5.5)], ratio club/pedicel: 3.0–3.2 [3.1–3.2]; chela+ 2.98–3.38/0.38–0.42 (7.9–8.1) [3.35– 3.44/0.44–0.48 (7.1–7.8)], chela - 2.82–3.16/0.38–0.42 (7.5–7.6) [3.15–3.23/0.44–0.48 (6.6–7.3)]; hand+ 1.15–1.26 (3.0–3.1) [1.28–1.31 (2.7–2.9)], hand - 0.99–1.04 (2.5–2.6) [1.07–1.1 (2.3–2.4)]; movable finger 1.85–2.15 [2.11– 2.19]; ratio finger/hand+ 1.6–1.7 [1.6–1.7)]; ratio chela+/carapace 2.8–3.0 [2.8–3.0]; femur/carapace 1.8 [1.6–1.8]; finger/femur 1.0 [1.0–1.1]; femur/patella 1.01–1.05 [1.0]; patella/hand+ 1.5–1.7 [1.5–1.6]. Leg I: femur 0.9–1.05/ 0.14–0.15 (6.4–6.9) [1.0–1.07/0.15–0.16 (6.6–6.7)]; patella 0.70–0.84/0.12–0.13 (6.0–6.5) [0.74–0.84/0.13–0.14 (5.7–6.2)]; tibia 0.86–1.02/.09–0.1 (9.4–10.7) [0.98–1.02/0.1 (10.2–10.4)]; basitarsus 0.47–0.56/0.08–0.09 (6.0– 6.6) [0.52–0.53/0.08 (6.5/6.7)]; telotarsus 0.60–0.71/0.07 (9.0–9.9) [0.66–0.7/0.07 (9.1–9.4)]; ratio femur/patella 1.3 [1.3–1.4]; telotarsus/basitarsus 1.3 [1.3]. Leg IV: femur+patella 1.58–1.78/0.21–0.22 (7.6–8.1) [1.7–1.8/0.21– 0.23 (7.5–8.8)]; tibia 1.55–1.79/0.13–0.14 (11.9–13.0) [1.68–1.74/0.14–0.15 (11.8–12.1)]; basitarsus 0.59–0.69/ 0.1–0.11 (6.1–6.5) [0.65–0.7/0.11 (6.0–6.4)]; telotarsus 0.71–0.84/0.08–0.09 (8.4–9.3) [0.82–0.87/0.09 (9.0–9.3)]; ratio femur+patella/tibia 1.0 [1.0]; telotarsus/basitarsus 1.2 [1.2–1.3].</p><p>Remarks. Beier (1962) described the tarsal claws of R. blothroides as lacking a dorsal tooth and used this as a characteristic to separate this species from R. cavernicola; as did Mahnert (1976), based on Beier’s description. Zaragoza (2002) suggested that, due to the tiny size of the teeth in some Roncocreagris species, it was not difficult to imagine that Beier might have overlooked them in R. blothroides . Given the agreement in all other studied characters, we have no doubt that the specimens described above belong to R. blothroides, hence the absence/ presence of teeth on the tarsal claws is not a valid diagnostic character for this species.</p><p>Beier’s original diagnosis (1962) of R. blothroides stated that it bears 4 setae on the posterior margin of the carapace and 4 setae on tergite I. Of the 24 specimens checked, 16 coincide with this (4+4), two bear 4+3, two 4+5, one 4+6, one 5+4, and two 5+5. No specimens bear 6 or more setae on posterior margin of carapace, and this number of setae is very rare on tergite I. This pattern is important for separating taxonomic groups within the genus Roncocreagris .</p><p>The discal setae of the sternites are glandular with a visible duct, as first described and illustrated by Judson (1992) and also mentioned and illustrated by Zaragoza 2002.</p><p>The sensillum of the movable finger, near dental margin and close to the trichobothrium st, is diploid as mentioned and illustrated for other species by Judson (1992) and Zaragoza (2003), but it is not raised in this species.</p><p>Roncocreagris blothroides and R. cavernicola share the feature of being the only species to usually have 4 setae on the posterior margin of the carapace and on tergite I. They differ as follows: the chelal measurements and ratios are distinctly larger in R. blothroides than in R. cavernicola; trichobothrium ist is medial with respect to ib / it, and it is medial with respect to est / et in R. blothroides, whereas in R. cavernicola ist is placed more basally and it is closer to est than to et; the male galea bears some rami apically in R. blothroides, but it is simple in R. cavernicola .</p><p>Distribution and habitat. All specimens of Roncocreagris blothroides were collected in the deepest parts of three caves, the two farthest separated by over 25 km. In addition to the type localities of Santa Maria da Estrela and Moura (= Arrifana) Caves, this species is newly recorded from Cerâmica and Confraria Caves, all of which are located in the Sicó karst massif (Fig. 1).</p><p>Roncocreagris blothroides is an abundant pseudoscorpion in the studied caves, with up to ten individuals being found during a single cave visit. It was found inhabiting a temperature range of 13.6–16.6ºC at soil level. This species shares its habitat with some frequent troglophile springtails, such as Neelus murinus Folsom, 1896, Willemia buddenbrocki Hüther, 1959, Onychiurus insubraruis Gisin, 1952, Onychiurus subgranulosus Gama, 1964, Folsomia candida (Willem, 1902), Proisotoma gisini Gama, 1964 and Tullbergia krausbaueri (Börner, 1901) (Gama 1962, 1965).</p></div>	https://treatment.plazi.org/id/015C614EF409FF9D5C94F895061FFCBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Reboleira, Ana Sofia P. S.;Zaragoza, Juan A.;Gonçalves, Fernando;Oromí, Pedro	Reboleira, Ana Sofia P. S., Zaragoza, Juan A., Gonçalves, Fernando, Oromí, Pedro (2013): On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species. Zootaxa 3670 (2): 283-299, DOI: 10.11646/zootaxa.3670.2.11
015C614EF40DFF935C94FCD403BFF945.text	015C614EF40DFF935C94FCD403BFF945.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Roncocreagris borgesi Zaragoza & Reboleira	<div><p>Roncocreagris borgesi Zaragoza &amp; Reboleira sp. nov.</p><p>(Figs 13–22)</p><p>Type material. Holotype: 3, Portugal, Sicó Massif, Santiago da Guarda, District of Leiria, Gruta da Cerâmica (39º55ʹ36.57ʺN, 8º31ʹ0 3.63ʺW; 355 m. a.s.l), 28.XI.2009, lgt. A.S.P.S. Reboleira (DEUA coll.).</p><p>Etymology. The species is dedicated to the Portuguese biologist Professor Paulo Borges, who has greatly contributed to our knowledge of the hypogean fauna of the Azores.</p><p>Diagnosis. No eyes or eye-spots. Moderately troglomorphic. Carapace with 20 setae, 6 on the posterior margin. Tergite I with 6 setae. Male cheliceral galea short and simple. Pedipalp: femur ratio about 6.0, femur as long as the movable finger; chela+ length/breadth ratio about 5.0; chelal hand widest proximad of middle, ratio movable finger/hand+ 1.4; trichobothium ist close to the middle of the finger, ratio ib–ist / ist–it 1.5, ratio isb–ist / ib– isb 1.5.</p><p>Description. Medium-sized pseudoscorpion. Opisthosoma elongate, moderate troglomorphic adaptations. Opisthosoma and legs yellowish. Carapace, chelicerae and pedipalps reddish brown.</p><p>Carapace longer than broad (Fig. 13). Without eyes or eye-spots. Anterior margin moderately prominent medially, epistome blunt and almost indistinguishable, with some tiny denticles not exceeding the anterior margin (Fig. 14). Chaetotaxy: 20 setae, formula 4:4:6:6. Five microlyrifissures on ocular zone, two between median and posterior zones.</p><p>Coxal area. Manducatory process with 3 setae. Anterior process of coxa I with simple tooth shape, long and apically pointed; medial process straight with a few denticles (Fig. 15). Pedipalpal coxa with 7 setae, pedal coxa I with 6, II: 3–4, III: 3–4, IV: 6.</p><p>Tergal chaetotaxy I–X: 6:7:9:9:9:11:11:10:10:9, most hairs lacking. Male genital area with 10 long setae on sternite II; sternite III with 10 setae, 4 of them along posterior margin of genital opening. Sternal chaetotaxy IV–X: 8:11:11:13:11:11:11; sternites VI and VII with 1 and 2 discal setae respectively (included in sternal formula); discal setae glandular, with visible duct. Segment XI 9 setae. Chaetotaxy of stigmata of sternites III and IV could not be checked due to partial contraction of opisthosoma. Anal cone with two dorsal and two ventral setae.</p><p>Chelicerae (Figs 16–17). Palm with 6 setae; subgaleal seta 0.65 from base of movable finger. Galea short (length 0.03 mm), pointed and simple. Fixed finger with 6 small, blunt, distal denticles, resembling protuberances, and 10 normal teeth, 4 basal ones larger than the others; movable finger with 17 teeth, one large and blunt subdistal tooth, the others medium or small, dental row ending just proximad of subgaleal seta. Rallum with 8 blades, all unilaterally pinnate on anterior face, the basal one about half length of others. Serrula exterior with 31 blades, serrula interior 25 blades.</p><p>Pedipalps (Figs 18–20). Trochanter, femur, distal third of the patella and the hand at base of the fingers with low granulation, more pronounced on paraxial faces. Lyrifissures as in Figs 18–20. Femur with one tiny tubercle distad of middle of antiaxial face, one distal glandular pore present. Patella with one micropore at base of pedicel. Chelal hand oval shaped in dorsal view, maximum width proximad of middle; pedicel bears one dorsal micropore; antiaxial face, close to finger base, bears an irregular row of 4 glandular pores. Fixed finger with 93 teeth, most of them apically cusped, dental row reaching up to level of trichobothrium esb; nodus ramosus short, at level of 4th distal tooth; trichobothrium it proximad of et, distinctly closer to est than to et; trichobothrium ist closer to base of finger than to apex; distance between trichobothria ib and ist 1.5 times longer than that between ist and it; distance between trichobothria isb and ist 1.5 times longer than that between isb and ib. Chelal microsetae pattern with all groups present, extending distad of trichobothrium isb, Em 3–4, Mm 3, Im 1. Movable finger with 85 teeth, most of them apically cusped, dental row shorter than on fixed finger, ending distad of trichobothrium b; distance between trichobothria sb and st 1.2 times longer than that between sb and b. One sensillum near tip of both fingers; diploid sensillum pc not raised, close to dental margin, level with trichobothrium sb.</p><p>Legs. Claws of legs I and IV with a tiny dorsal tooth proximad of middle (Fig. 21), subterminal setae with three rami (Fig. 22). Leg IV tibia TS 0.44, basitarsus TS 0.22, telotarsus TS 0.24.</p><p>Measurements and ratios. Body 2.1. Carapace 0.85/0.64 (1.3). Chelicera: palm 0.49/0.24 (2.0), movable finger 0.33. Pedipalp: trochanter 0.66/0.19 (3.5); femur 1.15/0.19 (6.1); patella 1.06/0.23 (4.6), pedicel 0.29, club 0.77/0.23 (3.4), ratio club/pedicel: 2.7; chela+ 1.98/0.39 (5.0), chela - 1.84/0.39 (4.7); hand+ 0.85 (2.2), hand - 0.71 (1.8); movable finger 1.16; ratio finger/hand+ 1.4; ratio chela+/carapace 2.3; femur/carapace 1.4; finger/femur 1.0; femur/patella 1.1; patella/ hand+ 1.2. Leg I: femur 0.57/0.11 (5.2); patella 0.41/0.12 (3.3); tibia 0.53/.009 (5.6); basitarsus 0.26/0.08 (3.5); telotarsus 0.40/0.06 (6.9); ratio femur/patella 1.4; telotarsus/basitarsus 1.5. Leg IV: both femoris contracted and not measurable, patella 0.51/0.22 (2.4); tibia 0.88/0.11 (8.2); basitarsus 0.33/0.09 (3.4); telotarsus 0.49/0.07 (5.1); ratio telotarsus/basitarsus 1.5.</p><p>Remarks. Among the species with 6 setae or more on posterior margin of carapace and tergite I, the new species shares with R. aurouxi Zaragoza, 2000 (from Cantabria, Spain) the simple shape of the cheliceral galea, although it should be noted that the types of the two species belong to different sexes (only the female is known for R. aurouxi). Also, both species have relatively low ratios for pedipalpal femur and chela for hypogean species of the genus. However, they differ in the chelal movable finger/hand ratio, which is 1.4 in R. borgesi vs 1.7 in R. auroxi; moreover the chelal finger length is almost equal than the femur in the new species, but distinctly longer in R. aurouxi .</p><p>Distribution and habitat. Only known from the type locality in Sicó massif, Roncocreagris borgesi sp. nov. was collected in the deepest galleries of Cerâmica Cave, where relative humidity is around 100% throughout the year and mean temperature at soil level is 15.3ºC. The presence of two hypogean pseudoscorpion species, R. blothroides and R. borgesi, in the same cave is infrequent but not unknown (e.g. Zaragoza 2007). Due to the moderate troglomorphism and scarce occurrence of R. borgesi in the cave, compared to R. blothroides, we suggest that this species might predominantly occur in the mesovoid shallow substratum, rather than in the cave itself.</p><p>Cerâmica Cave is the richest one of central Portugal in terms of known troglobiont fauna, harboring 10 caveadapted species: 3 pseudoscorpions, 1 spider, 1 millipede, 3 oniscidean woodlice, 1 dipluran and 1 beetle (Reboleira, 2012). Among the cave-adapted species, the pseudoscorpions R. blothroides, R. borgesi sp. nov. and Chthonius sp., the spider Lepthyphantes sp. and the rove-beetle Domene lusitanica Reboleira &amp; Oromí, 2011 are macro- and micropredators, while a new species of chordeumatid millipede, the woodlice Porcellio cavernicolus Vandel, 1964 and two trichoniscids, together with the dipluran Podocampa cf. fragiloides Silvestri, 1932, play a detritivorous role in this subterranean ecosystem (Reboleira et al. 2011a, 2011b). Other interesting troglophile species have stable populations in this biocoenosis, such as the gastropod Oxychilus draparnaldi (Beck, 1837), the centipede Lithobius pilicornis Newport, 1844 and the ground beetle Trechus fulvus Dejean, 1831 .</p></div>	https://treatment.plazi.org/id/015C614EF40DFF935C94FCD403BFF945	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Reboleira, Ana Sofia P. S.;Zaragoza, Juan A.;Gonçalves, Fernando;Oromí, Pedro	Reboleira, Ana Sofia P. S., Zaragoza, Juan A., Gonçalves, Fernando, Oromí, Pedro (2013): On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species. Zootaxa 3670 (2): 283-299, DOI: 10.11646/zootaxa.3670.2.11
015C614EF403FF905C94F88C036CFE7E.text	015C614EF403FF905C94F88C036CFE7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Roncocreagris gepesi Zaragoza & Reboleira	<div><p>Roncocreagris gepesi Zaragoza &amp; Reboleira sp. nov.</p><p>(Figs 23–32)</p><p>Type material. Holotype Ƥ, Portugal, Sicó Massif, district of Coimbra, Penela, Gruta do Algarinho/Dueça Cave system (39°59ʹ4.89"N, 8°23ʹ5.86"W; 208 m. a.s.l.), 25.IV.2009, lgt. A.S.P.S. Reboleira (DEUA).</p><p>Etymology. This species is dedicated to the speleological association Grupo Protecção Sicó (GPS), for its contribution to the knowledge and protection of caves and karst of Sicó region, and as recognition to its members for their effort and support during our field work.</p><p>Diagnosis. No eyes or eye-spots. Strong troglomorphic adaptations. Carapace with 21 setae, 7 on the posterior margin. Tergite I 6 setae. Female cheliceral galea long, with five short, apical rami. Pedipalp: femur ratio about 6.6, femur longer than movable finger; chela+ ratio about 5.9; chelal hand widest at middle, ratio movable finger/hand+ 1.4; trichobothium ist close to middle of finger, ratio ib–ist / ist–it 1.3, ratio isb–ist / ib–isb 1.8.</p><p>Description. Large pseudoscorpion. Opisthosoma elongate, advanced troglomorphic adaptations. Opisthosoma and legs yellowish. Carapace, chelicerae and pedipalps reddish brown.</p><p>Carapace distinctly longer than broad (Fig. 23). Without eyes or eye-spots. Anterior margin moderately prominent medially, epistome blunt and short (Fig. 24). Chaetotaxy: 21 setae, formula 4:4:6:7. Four microlyrifissures in ocular zone, two between median and posterior zones.</p><p>Coxal area. Manducatory process with 3 setae. Anterior process of coxa I with simple tooth shape, long and apically pointed; medial process straight with some small tubercles (Fig. 25). Pedipalpal coxa with 7 setae, pedal coxa I with 6, II: 5–6, III: 3, IV: 5.</p><p>Tergal chaetotaxy I–X: 6:6:7:10:9:10:10:10:9:9, most hairs lacking. Genital area with 5 thin setae on sternite II. Sternal chaetotaxy III–X: 9:7:12:12:12:9:9:8; sternites VI and VII each with 2 discal setae (included in sternal formula); discal setae glandular, with visible duct. Segment XI 9 setae. Chaetotaxy of stigmata of sternites III and IV with 3 microsetae on each side. Anal cone with two dorsal and two ventral setae.</p><p>Chelicerae (Figs 26–27). Palm with 6 setae, subgaleal seta 0.68 from base of movable finger. Galea 0.04 mm long, with five short, apical rami. Fixed finger with 5 small, blunt distal denticles and 14 other small, medium teeth; movable finger with 13 teeth, one large and blunt subdistal tooth, the others medium or small, dental row ending just proximad of subgaleal seta. Rallum with 8 blades, all unilaterally pinnate on anterior face, basal blade about half length of others. Serrula exterior with 32 blades, serrula interior 27 blades.</p><p>Pedipalps (Figs 28–30). Trochanter, femur, distal third of patella and hand at base of the fingers distinctly granulated, more pronounced on paraxial faces. Lyrifissures as in Figs 28–30. Antiaxial face of trochanter and femur with one tiny tubercle at middle or slightly more distad; femur bearing one distal glandular pore. Patella with one micropore at base of pedicel. Chelal hand elliptic in dorsal view, maximum width at middle; three dorsal micropores on pedicel; two glandular pores on antiaxial face close to finger base. Fixed finger with 98 teeth, most of them apically cusped, dental row almost reaching level of trichobothrium esb; nodus ramosus short, reaching level of 5th distal tooth; trichobothrium it proximad of et, closer to est than to et; trichobothrium ist slightly closer to base of finger than to apex; distance between trichobothria ib and ist 1.3 times longer than that between ist and it; distance between trichobothria isb and ist 1.7 times longer than that between isb and ib; distance between trichobothria et and it 1.2 times longer that between it and est. Chelal microsetae pattern: Em 3 curved setae, Mm 2, Im 1. Movable finger with 103 teeth, most of them apically cusped, dental row ending distad of trichobothrium b, shorter than that of fixed finger. Distance between trichobothria s b and st 1.2 times longer than that between sb and b. One sensillum near the tip of both fingers; diploid sensillum pc not raised, close to dental margin, slightly proximad of trichobothrium sb.</p><p>Legs. Claws of legs I and IV with a tiny dorsal tooth proximad to middle (Fig. 31), subterminal setae with three rami, trident shaped (Fig. 32). Leg IV tibia TS ratio 0.48, basitarsus TS 0.13, telotarsus TS 0.27.</p><p>Measurements and ratios. Body 2.19. Carapace 0.95/0.65 (1.5). Chelicera: palm 0.57/0.27 (2.1), movable finger 0.36. Pedipalp: trochanter 0.79/0.21 (3.9); femur 1.46/0.22 (6.6); patella 1.41/0.26 (5.5), pedicel 0.36, club 1.05/0.26 (4.1), ratio club/pedicel: 2.92; chela+ 2.39/0.41 (5.9), chela - 2.21/0.41 (5.5); hand+ 1.03 (2.5), hand - 0.85 (2.1); movable finger 1.39; ratio finger/hand+ 1.4; ratio chela+/carapace 2.5; femur/carapace 1.5; femur/finger 1.1; femur/patella 1.0; patella/ hand+ 1.4. Leg I: femur 0.73/0.14 (5.4); patella 0.51/0.12 (4.4); tibia 0.68/.08 (8.7); basitarsus 0.33/0.07 (5.0); telotarsus 0.45/0.06 (8.2); ratio femur/patella 1.4; telotarsus/basitarsus 1.4. Leg IV: femur+patella 1.12/0.25 (4.5); tibia 1.12/0.13 (8.4); basitarsus 0.41/0.1 (4.2); telotarsus 0.58/0.08 (7.4); ratio femur+patella/tibia 1.0; ratio telotarsus/basitarsus 1.4.</p><p>Remarks. The new species is easily distinguishable from R. borgesi sp. nov. by the shape of the chelal hand, which is elliptic and widest at middle in R. gepesi versus oval and widest proximad of middle in R. borgesi; also the pedipalps are larger and more slender in R. gepesi . The new species can be differentiated from R. cavernicola by the chelal movable finger/hand+ ratio (1.4, versus 1.7) and by the chaetotaxy of the posterior margin of the carapace (7 setae, versus 4) and of tergite I (6 setae, versus 4). Roncocreagris gepesi is more closely related to R. lucensis Zaragoza, 2002 (from Galicia, Spain), despite being 400 km away, but in the latter the pedipalps are shorter and less slender than in R. borgesi, and the distance between trichobothria et and it is 3.3 times longer than that between it and est in the Spanish species, versus 1.2 in the Portuguese species.</p><p>Distribution and habitat. Roncocreagris gepesi sp. nov. was collected in Algarinho, one of the caves of the Dueça system, opened in 1998 with the aid of a backhoe (Neves et al. 2005). This species was not found in Soprador do Carvalho, a cave located only 200 m from the entrance of Algarinho Cave. The mean cave temperature is 14.9ºC at soil level. The new species shares its habitat with other hypogean species, such as the woodlouse Porcellio cavernicolus and the dipluran Podocampa cf. fragiloides.</p></div>	https://treatment.plazi.org/id/015C614EF403FF905C94F88C036CFE7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Reboleira, Ana Sofia P. S.;Zaragoza, Juan A.;Gonçalves, Fernando;Oromí, Pedro	Reboleira, Ana Sofia P. S., Zaragoza, Juan A., Gonçalves, Fernando, Oromí, Pedro (2013): On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species. Zootaxa 3670 (2): 283-299, DOI: 10.11646/zootaxa.3670.2.11
015C614EF400FF955C94FD970463FB22.text	015C614EF400FF955C94FD970463FB22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Roncocreagris occidentalis Zaragoza & Reboleira	<div><p>Roncocreagris occidentalis Zaragoza &amp; Reboleira sp. nov.</p><p>(Figs 33–43)</p><p>Type material. Holotype Ƥ, Portugal, Serra de Montejunto, District of Lisboa, Pragança, Algar do Javali (39º11.965ʹN, 9º01.718ʹW; 380 m. a.s.l.), 19.IX.2009, lgt. A.S.P.S. Reboleira (DEUA). Paratypes: 1 deutonymph, same location as holotype, 19.IX.2009, lgt. A.S.P.S. Reboleira (DEUA); 1 Ƥ, same location as holotype, 5.IV.2009 lgt. A.S.P.S. Reboleira (DZUL); 1 Ƥ, Portugal, Cesaredas Plateau, District of Lisboa, Bolhos, Gruta dos Bolhos (39°18ʹ31.35"N, 9°16ʹ36.69"W; 145 m. a.s.l.), 26.XII.2010, lgt. A.S.P.S. Reboleira (DEUA).</p><p>Etymology. Named in reference to the westernmost location known for a hypogean species of the genus Roncocreagris .</p><p>Diagnosis. No eyes or eye-spots. Strongly troglomorphic. Carapace with 20 setae, 6 on posterior margin. Tergite I 6–7 setae. Female cheliceral spinneret reduced to a hyaline dome. Pedipalp femur ratio about 5.7, femur shorter than the movable finger; chela+ ratio about 5.5; chelal hand broadest proximad of middle, ratio movable finger/hand+ about 1.4; trichobothium ist close to finger base, ratio ib–ist / ist–it about 0.5–0.6.</p><p>Description. Female holotype, followed by female paratypes in brackets. Large pseudoscorpion with troglomorphic adaptations (Fig. 33). Opisthosoma elongate. Opisthosoma and legs yellowish. Carapace, chelicerae and pedipalps pale brown.</p><p>Carapace (Figs 34–35) longer than broad. Without eyes or eye-spots. Anterior margin moderately prominent medially, epistome blunt and almost indistinguishable. Chaetotaxy: 20 setae, formula 4:4:6:6. Three (four) microlyrifissures in ocular zone, two between median and posterior zones.</p><p>Coxal area. Manducatory process with 3 setae. Anterior process of coxa I simple, long and apically pointed; medial process straight with some small, blunt protuberances (Fig. 36). Pedipalpal coxa with 7 (6–8) setae, pedal coxa I with 7 (7–8), II: 6–7, III: 4 (5), IV: 6–7.</p><p>Tergal chaetotaxy I–X: 7:7:8:9:9:10:10:11:11:10 (6–7:7–8:8:9:9–10:9–10:11:10:10–11:9), most hairs lacking. Genital area with 5 (6) thin setae on sternite II. Sternal chaetotaxy III–X: 10:8:11:13:13:11:11:11 (10:7:11:12:12:11:11:9–10); sternites VI and VII each with 2 discal setae (included in sternal formula); discal setae glandular, with visible duct. Segment XI: 9 setae. Stigmata III and IV with 3 microsetae on each side. Anal cone with two dorsal and two ventral setae.</p><p>Chelicera (Figs 37–38). Palm with 6–7 setae, subgaleal seta 0.66 (0.62–0.68) from base of movable finger. Spinneret a low hyaline dome. Fingers with teeth worn (fixed finger with 5 distal denticles and 7–12 small and medium sized teeth; movable finger with 3 distal denticles and 14 teeth, one large medial tooth, the others medium or small, dental row ending proximad of subgaleal seta. Rallum with 8 blades, all unilaterally pinnate on anterior face, basal blade about half length of others. Serrula exterior with 32 (31–33) blades, serrula interior 24 (25–26) blades.</p><p>Pedipalp (Figs 39–41). Femur weakly granulated on paraxial face, more pronounced in basal half; chelal hand weakly granulated at base of finger; trochanter and patella smooth. Lyrifissures as in Figs 39–41. Femur with one tiny tubercle just distad of middle of antiaxial face, dorsally with one distal glandular pore. Patella with two dorsal micropores at base of pedicel. Chelal hand dorsal gently ovoid, widest proximad of middle; pedicel bears two dorsal micropores; antiaxial face with 4 glandular pores close to base of finger. Fixed finger with 100 teeth, most of them apically cusped, dental row reaching base of finger; nodus ramosus short, level with 4th distal tooth; trichobothrium it proximad of et, halfway between est and et; trichobothrium ist markedly closer to base of finger than to apex; distance between trichobothria ib and ist 0.5 (0.5–0.6) times shorter than that between ist and it. Chelal microsetae pattern: all groups present, extending distad of trichobothrium isb, Em 5 (4–5), Mm 2, Im 2. Movable finger with 97 fingers, most of them cusped, dental row shorter than on fixed finger, ending just distad of trichobothrium b; distance between trichobothria sb and st 1.1 (0.9–1.2) times that between sb and b. One sensillum near tip of both fingers; diploid sensillum pc not raised, close to dental margin, level with trichobothrium sb.</p><p>Legs. Claws of legs I and IV with a tiny dorsal tooth proximad of middle (Fig. 42), subterminal setae with two rami and some spinules (Fig. 43). Leg IV tibia TS ratio 0.49 (0.45), basitarsus TS 0.14 (0.13), telotarsus TS 0.33 (0.44).</p><p>Deutonymph. Body and appendages yellowish. Carapace longer than broad, no eyes or eye-spots, epistome absent; chaetotaxy 20 setae, formula 4:4:6:6. Manducatory process 3 setae, pedipalp coxa 5, pedal coxa I 4, II 4, III 3, IV 4; anterior process of coxa I long and tooth-shaped, medial process straight, with one denticle. Tergal chaetotaxy I–X: 6:6:7:7:7:7:7:7:7:8. Sternal chaetotaxy III–X: 4:4:9:9:8:9:8:9; sternites VI and VII each with 2 discal setae (included in sternal formula); discal setae distinctly closer to sternal row than in adults; segment XI 9 setae; stigmata of sternites III and IV with one microseta on each side. Chelicera with five setae on palm, subgaleal seta 0.6 from base, spinneret reduced to a hyaline dome; fixed finger with 12 teeth, movable finger with 10 teeth. Pedipalps smooth, except chelal hand at base of finger; femur with a tiny antiaxial tubercle in basal half; distal opening of patella oblique; chelal hand more oval than in adults, maximum width proximad of middle, antiaxial face with one glandular pore close to base of finger, one dorsal micropore on pedicel; chelal microsetae pattern: Em 2, Mm 1, Im 0; fixed finger with six trichobothria and 48 teeth, movable finger with two trichobothria and 45 teeth, dental row ending at level of diploid sensillum. Leg IV: telotarsus basally swollen; claws bearing a tiny basal tooth; subterminal setae with tiny distal denticles; tibia TS 0.35, basitarsus TS 0.20, telotarsus TS 0.27.</p><p>Measurements and ratios. Female holotype (followed by female paratype from Algar do Javali in square brackets). Body 3.74 [1.96, opisthosoma contracted]. Carapace 0.83/0.68 (1.2) [0.78/0.67 (1.2)]. Chelicera: palm 0.58/0.29 (2.0) [0.51/0.25 (2.0)], movable finger 0.36 [0.31]. Pedipalp: trochanter 0.68/0.22 (3.1) [0.61/0.19 (3.1)]; femur 1.28/0.22 (5.8) [1.13/0.20 (5.7)]; patella 1.21/0.26 (4.7) [1.06/0.23 (4.6)], pedicel 0.36 [0.34], club 0.85/0.26 (3.3) [0.72/0.23 (3.1)], ratio club/pedicel 2.4 [2.1]; chela+ 2.28/0.42 (5.5) [2.0/0.38 (5.3)], chela - 2.15/0.42 (5.1) [1.85/0.38 (4.9)]; hand+ 0.99 (2.4) [0.83 (2.2)], hand - 0.86 (2.1) [0.68 (1.8)]; movable finger 1.32 [1.18]; ratio chela+/carapace 2.8 [2.6]; femur/carapace 1.5 [1.5]; finger/femur 1.03 [1.04]; femur/patella 1.06 [1.07]; patella/ hand+ 1.22 [1.3]; finger/hand+ 1.3 [1.4]. Leg I: femur 0.60/0.15 (4.1) [0.55/0.15 (3.8)]; patella 0.48/0.12 (3.9) [0.41/ 0.13 (3.2)]; tibia 0.59/0.09 (6.9) [0.52/0.10 (5.2)]; basitarsus 0.29/0.07 (4.1) [0.26/0.07 (3.6)]; telotarsus 0.48/0.06 (8.0) [0.42/0.06 (6.7)]; ratio femur/patella 1.25 [1.3]; basitarsus/telotarsus 1.7 [1.7]. Leg IV: femur+patella 1.10/ 0.24 (4.7) [0.93/0.21 (4.4)]; tibia 0.99/0.13 (7.8) [0.88/0.12 (7.3)]; basitarsus 0.37/0.10 (3.7) [0.31/0.10 (3.0)]; telotarsus 0.58/0.08 (7.3) [0.53/0.09 (5.9)]; ratio femur+patella/tibia 1.1 [1.1]; telotarsus/basitarsus 1.6 [1.7].</p><p>Female paratype from Gruta dos Bolhos. Body 3.56. Carapace 0.78/0.66 (1.2). Chelicera: palm 0.52/0.26 (2.0); movable finger 0.34. Pedipalp: trochanter 0.60/0.21 (2.8); femur 1.12/0.22 (5.2); patella 1.06/0.27 (3.9), pedicel 0.32, club 0.74/0.27 (2.7), ratio club/pedicel 2.3; chela+ 2.04/0.40 (5.1), chela - 1.89/0.40 (4.7); hand+ 0.94 (2.3), hand - 0.79 (2.0); movable finger 1.12; ratio chela+/carapace 2.6; femur/carapace 1.4; finger/femur 1.0; femur/ patella 1.1; patella/hand+ 1.1; finger/hand+ 1.2. Leg I: femur 0.50/0.14 (3.7); patella 0.38/0.11 (3.4); tibia 0.48/0.08 (5.8); basitarsus 0.22/0.07 (3.4); telotarsus 0.40/0.06 (6.6); ratio femur/patella: 1.3; telotarsus/basitarsus 1.8. Leg IV: both legs IV absent.</p><p>Deutonymph. Body 1.52. Carapace 0.46/0.37 (1.2). Chelicera: palm 0.27/0.14 (2.0), movable finger 0.18. Pedipalp: trochanter 0.27/0.11 (2.4); femur 0.44/.012 (3.8); patella 0.37/0.14 (2.7), pedicel 0.12, club 0.25/0.14 (1.9), ratio club/pedicel 2.1; chela+ 0.86/0.21 (4.2), chela - 0.81/0.21 (3.9); hand+ 0.38 (1.8), hand - 0.33 (1.6); movable finger 0.48; ratio chela+/carapace 1.9; femur/carapace 1.0; finger/femur 1.1; femur/patella 1.2; patella/ hand+ 1.0; finger/hand+ 1.3. Leg IV: femur+patella 41/14 (3.0); tibia 0.32/0.7 (4.6); basitarsus 0.14/0.06 (2.4); telotarsus 0.22/0.06 (4.0); ratio femur+patella/tibia 1.3; telotarsus/basitarsus 1.6.</p><p>Remarks. The main diagnostic characters of the paratype from Gruta dos Bolhos generally coinciding with those of the holotype and paratype from Algar do Javali, but it has a less slender chela. We prefer to assign the specimen from Gruta dos Bolhos to R. occidentalis sp. nov. until new material is available for study.</p><p>Roncocreagris occidentalis represents the second hypogean Roncocreagris with the spinneret reduced to a hyaline dome, the other being R. iglesiasae Zaragoza, 2003 (Asturias, Spain). R. occidentalis differs from R. iglesiasae in the position of trichobothrium ist, which is distinctly nearer to the finger base than to it in the Portuguese species, as opposed to being closer to it than to the finger base in the Spanish species. Moreover, R. occidentalis has the pedipalps longer but less slender than in R. iglesiasae .</p><p>Distribution and habitat. Roncocreagris occidentalis sp. nov. was collected in two caves from two different karst units, Javali in the Montejunto massif and Bolhos in the Cesaredas plateau.</p><p>In Javali Cave, which has a year-round temperature of 15.1 + 0.5ºC at soil level, R. occidentalis is found with other interesting cave-adapted species, such as the ground beetle Trechus tatai Reboleira &amp; Ortuño, 2010 (Reboleira et al. 2010b), while in Bolhos Cave the biocoenosis is composed of a new trichoniscid woodlouse, the dipluran Podocampa cf. fragiloides and the thysanuran Coletinia sp. and the mean temperature is 16.3ºC. Also found in both caves was the pseudoscorpion Chthonius (Ephippiochthonius) cardosoi Zaragoza, 2012, previously known only from the Arrábida massif (Zaragoza 2012).</p></div>	https://treatment.plazi.org/id/015C614EF400FF955C94FD970463FB22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Reboleira, Ana Sofia P. S.;Zaragoza, Juan A.;Gonçalves, Fernando;Oromí, Pedro	Reboleira, Ana Sofia P. S., Zaragoza, Juan A., Gonçalves, Fernando, Oromí, Pedro (2013): On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species. Zootaxa 3670 (2): 283-299, DOI: 10.11646/zootaxa.3670.2.11
