taxonID	type	description	language	source
01017A38FFA1DB7CFF69FEDDFE15CE6D.taxon	discussion	Remarks. While examining species belonging to the subgenus Deltochilum, it became apparent that the status of the single species (D. orbiculare van Lansberge) included in the subgenus Telhyboma Kolbe should be reconsidered. After a careful examination, it appears that the only apomorphic character separating D. orbiculare from all other species included in the subgenus Deltochilum is the development of the 7 th interstrial carina. In D. orbiculare this carina is present on most of the length of the 7 th interstria and overlaps the elytral edge and the 9 th interstrial carina on most of the elytral length. The development of this carina is clearly linked to gender. Hyperthelic males always exhibit a much more developed structure than females, suggesting that the development of this carina in D. orbiculare is an additional secondary sexual character. In their study of the subgenera Calhyboma, Hybomidium and Telhyboma for Colombia, González et al. (2009) also noted the asymmetrical parameres and the additional internal sac sclerites, which conform to other species in the subgenus Deltochilum. The following apomorphies define the subgenus Deltochilum in its current sense: overall body shape in dorsal view with widest portion at mid-distance or slightly past mid-distance to elytral apex; elytral surface with a distinct network of irregular glossy bulges (not glossy in D. enceladus Kolbe) sharply contrasting from remaining surface; protibia lacking tarsus in both sexes; male protibia with a modification on internal edge basal third; male profemur with a teeth past mid-distance on internal edge; male mesofemur with a denticle on basal half; male metafemur with posterior surface completely matte, with punctures weakly impressed, more widely separated than remaining surface or absent; male sternite 3 with a strong bulging structure medially (note: this structure is absent in both Central American species, D. scabriusculum Bates and D. tumidum Howden. However, in some specimens of D. scabriusculum and D. tumidum the surface shows wrinkles that would correspond to the lateral edges of the bulge, suggesting that in these species there is a reversal to the primitive state); parameres with strong ventral projections, slightly asymmetrical.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA1DB7EFF69FACAFE84CCE1.taxon	materials_examined	Type locality. Brasilien, St. Catharina.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA1DB7EFF69FACAFE84CCE1.taxon	distribution	Distribution. Bolivia, Brazil, Colombia, Paraguay.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA1DB7EFF69FACAFE84CCE1.taxon	materials_examined	Material examined (1 male, 1 female). BRAZIL: PARANÁ, Col. Ing. Vitoria, x. 1945, coll. F. Justus - 1 male (CMNC); PARAGUAY: CAAGUAZÚ, Caaguazú, (25 ° 26 ' 48 '' S, 56 ° 1 ' 43 '' W), x. 1948, coll. F. H. Schade - 1 female (CMNC).	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA1DB7EFF69FACAFE84CCE1.taxon	discussion	Remarks. Only two specimens have been examined. Easily separated from all other species by the shape of parameres (Figs. 1 – 2) and the lustreless dorsal surface, which is strongly contrasting with the small glossy bulges on elytra.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA4DB79FF69FF25FC06CF5B.taxon	materials_examined	Type locality. Xingu (Central-Brasilien).	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA4DB79FF69FF25FC06CF5B.taxon	distribution	Distribution. Bolivia, Brazil.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA4DB79FF69FF25FC06CF5B.taxon	materials_examined	Material examined (14 males, 3 females). BRAZIL: MATO GROSSO, 13.4 km S Posto do Gil, elev. 240 m (14 ° 37 ' 55 '' S, 56 ° 16 ' 23 '' W), 11. i. 2001, coll. Génier & Vaz de Mello (2001 - 17) - 1 male (CMNC); same locality, 13. i. 2001, coll. Génier & Vaz de Mello (2001 - 29) - 5 males (CMNC); Fazenda São João, Municipio Diamantino, elev. 400 m (14 ° 14 ' 10 '' S, 56 ° 8 ' 11 '' W), 11. i. 2001, coll. Génier & Vaz de Mello (2001 - 04) - 1 female, 1 male (CMNC); same locality, 13. i. 2001, coll. Génier & Vaz de Mello (2001 - 46) - 1 female, 1 male (CMNC); same locality, 13. i. 2001, coll. Génier & Vaz de Mello (2001 - 39) - 2 males (CMNC); same locality, 13. i. 2001, coll. Génier & Vaz de Mello (2001 - 45) - 1 male (CMNC); MINAS GERAIS, Vespasiano, (19 ° 40 ' S, 43 ° 55 ' W), xii. 1951, coll. P. Pereira - 1 male (CMNC); PARÁ, IPEAN [= Instituto de Pesquisas e Experimentação Agropecuárias do Norte], Belem, (1 ° 26 ' 32 '' S, 48 ° 25 ' 59 '' W), xi. 1984, coll. L. Arnaud - 1 male (CMNC); Mujo, xii. 1921, coll. [anonymous] - 1 male (CMNC); SÃO PAULO, Fazenda Fugueira, Guararapes, (21 ° 15 ' S, 50 ° 38 ' W), 23. ii. 1941, coll. N. Sannomia - 1 female (CMNC).	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA4DB79FF69FF25FC06CF5B.taxon	discussion	Remarks. Enceladus is the proper epithet spelling. The name is derived from the Greek deity Enceladus, son of Gaia and brother of Eurymedon. The largest species of the subgenus, which can easily be separated by the shape of parameres (Figs. 3 – 4) and externally by the evenly dull dorsum.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA4DB78FF69FBD4FB8BCE89.taxon	materials_examined	Type locality. PANAMA: DARIÉN, Estación Ambiental Cana (7 ° 45.32 ' N, 77 ° 41.07 ' W), 600 – 750 m, 5 - 7. vi. 1996, coll. R. S. Anderson, 1996 - 110 - male holotype (CMNC)	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA4DB78FF69FBD4FB8BCE89.taxon	diagnosis	Diagnosis. D. eurymedon is unique in the genus Deltochilum in having the 7 th interstrial carina complete and reaching the elytral anterior edge combined with the sharply defined cephalic punctures throughout and from D. rosamariae in having the second interstria distinctly tuberculate apically. The shape of parameres (Figs. 15 – 16), especially the very long and curved dorsal projections, will also separate D. eurymedon from all other species in the genus.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA4DB78FF69FBD4FB8BCE89.taxon	description	Description. Holotype male (Fig. 13). Length 30.0 mm, width 21.0 mm. Colour. Black with feeble coppery sheen; antennal club and pubescence fulvous. Head. Clypeus with two sharp, upturned and widely separated teeth anteriorly, anterior margin slightly lobate medially and on each side of teeth. Head dorsal surface with sharply defined punctures throughout, punctures larger and more widely separated on disc; surface between punctures weakly microreticulated anteriorly, microsculpture becoming sharply defined posteriorly giving the surface a silky aspect. Eyes and ocular canthus unmodified. Pronotum. Moderately convex, anterior portion slightly gibbous anteriorly; surface completely punctate, punctures dense and evenly spaced, gradually becoming larger and umbilicate laterally; surface between punctures finely microreticulated on disc and anteriorly, microsculpture becoming strong laterally giving the surface a strong silky sheen on anterior and posterior angles; posterior edge distinctly bisinuate on each side, produced into a small denticle medially. Elytra. broadly arcuate in dorsal view, largest portion just behind mid distance; elytral striae 1 – 6 fine and distinct from base to elytral apex, with widely spaced oval punctures; interstriae with a network of strong irregular and glossy bulges, base surface with weakly-defined umbilicate punctures; interstria 7 with a strong carina from base to apex; interstria 8 with surface nearly perpendicular to dorsal surface, widest past middle; stria 9 visible and connecting to stria 2 apically; interstriae 8 - 9 strongly sericeous on most of surface, umbilicate punctures weakly-defined; apical portion with well-defined glossy calluses on interstriae 2 – 6, calluses gradually becoming longer externally. Thoracic sternites. Mesosternum with anterior margin raised and sinuous medially, remaining surface with coarse umbilicate punctures laterally and a double row of punctures along anterior margin medially. Metasternal median lobe with a more or less rectangular and glossy raised area medially, lateral portions with coarse umbilicate punctures throughout, disc with a more or less oval depression posteriorly, depression lacking tubercle anteriorly. Lateral lobe covered with closely set coarse umbilicate punctures. Legs. Profemur with a dent on anterior edge at about mid-distance to apex, dent almost triangular in shape, with pubescence covering ventral portion; protibia inwardly bent on apical half, with a wide lobate projection before mid-distance on internal edge; mesocoxa with a distinct triangular teeth on posterior edge, with a setal brush on internal side; mesofemur with a ventrally bent triangular process, posterior third surface strongly sericeous, sharply contrasting with glossy anterior surface; metatrochanter with a narrow setal brush ventrally to posterior edge, posterior edge produced into a ventrally bent lobate projection; metafemur strongly arcuate in ventral view, narrowest portion just before mid-distance, posterior surface completely matte and lacking punctures, dorsoposterior edge produced into a lobate projection separated from the trochantofemoral suture by its own width; metatibia sinuous and produced into a wide triangular tooth internally on basal fifth, apex spatulate, internal surface broadly concave. Abdomen. Sternite 3 produced into a strong approximately equilateral triangular process medially, widest portion of process in line with sternite 4. Pygidium approximately as wide as long; surface with transversally oval umbilicate punctures, punctures smaller and more widely separated basally; surface between punctures completely matte; margin very wide apically, internal edge broadly and evenly arcuate. Aedeagus. Parameres nearly symmetrical, with two straight and spiniform processes in lateral view, dorsal process longer, acute and more curved in dorsal view that the ventral one. Female (Fig. 14). Similar to male, lacking processes or modifications on legs; protibia wider, external teeth larger and obliquely directed. Sternite 3 unmodified medially. Variation. No variation except for body size and development of secondary sexual characters in males. Measurements (11 males, 5 females). Body length: male 22.0 – 30.0 (26.3 ± 2.5), female 26.0 – 28.5 (27.1 ± 1); body width: male 15.0 – 21.0 (17.8 ± 1.8), female 16.5 – 19.0 (17.8 ± 1.2) mm.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA4DB78FF69FBD4FB8BCE89.taxon	materials_examined	Material examined (11 males, 5 females). PANAMA: DARIÉN, Estación Ambiental Cana, elev. 460 m (7 ° 45.5 ' N, 77 ° 41 ' W), 3 - 9. vi. 1996, coll. Stockwell - 2 males (including 2 paratypes) (CMNC); same locality, 5 - 7. vi. 1996, coll. R. S. Anderson (1996 - 110) - 3 females, 2 males (including holotype, 3 paratypes) (CMNC); same locality, 7 - 10. vi. 1996, coll. R. S. Anderson (1996 - 117) - 2 females, 7 males (including 9 paratypes) (CMNC).	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA4DB78FF69FBD4FB8BCE89.taxon	etymology	Etymology. Eurymedon, a Giant of Greek mythology, son of Gaia and brother of Enceladus, relating to the large size and close relationship to D. enceladus. Natural history. All specimens with data have been collected in human feces baited pitfall traps set up during the afternoon. Current distributional data suggest that it is most likely an endemic of the Eastern Panamanian Mountain Forest. However, the elevation at which specimens were collected (460 – 700 m) is overlapping the Eastern Panamanian Mountain Forest which ranges from 500 – 1800 m and the Chocó-Darién Moist Forest which lies at elevations below 500 m. Further collecting will be necessary to settle the issue.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA5DB7AFF69F9A6FA19CA2E.taxon	materials_examined	Type locality. Bahia.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA5DB7AFF69F9A6FA19CA2E.taxon	distribution	Distribution. Bolivia, Brazil, Colombia, Ecuador, French Guiana, Peru.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA5DB7AFF69F9A6FA19CA2E.taxon	materials_examined	Material examined (38 males, 45 females). BOLIVIA: Alto Beni, elev. 1100 m, i. 1976, coll. Peña - 1 female, 1 male (CNC); Río Ichilo, elev. 350 m, [no date], coll. Steinbach - 1 female (CMNC); COCHABAMBA, 124 km E Cochabamba at Río Espíritu Santo, elev. 700 m (17 ° 3 ' 45 '' S, 65 ° 38 ' 38 '' W), 1 - 6. ii. 1999, coll. F. Génier (1999 - 31) - 1 female, 1 male (CMNC); 16.7 km W Villa Tunari, Avispas, elev. 500 m (17 ° 1 ' 13 '' S, 65 ° 32 ' 46 '' W), 10 - 12. ii. 1999, coll. F. Génier (1999 - 67) - 1 female, 1 male (CMNC); Estación Biológica El Valle del Sajta, Universidad Mayor de San Simón, elev. 300 m (17 ° 6.48 ' S, 64 ° 46.94 ' W), 16 - 27. xii. 2005, coll. S. & J. Peck (2005 - 47) - 1 female, 1 male (CMNC); Estación Biológica El Valle del Sajta, Universidad Mayor de San Simón, 67.5 km E Villa Tunari, elev. 300 m (17 ° 6 ' 19 '' S, 64 ° 46 ' 57 '' W), 7 - 9. ii. 1999, coll. F. Génier (1999 - 45) - 2 females (CMNC); same locality, 7 - 9. ii. 1999, coll. F. Génier (1999 - 48) - 2 females (CMNC); same locality, 7 - 9. ii. 1999, coll. F. Génier (1999 - 46) - 1 male (CMNC); same locality, 7 - 9. ii. 1999, coll. F. Génier (1999 - 47) - 1 male (CMNC); same locality, 7 - 9. ii. 1999, coll. R. Hanley (1999 - 57) - 1 male (CMNC); same locality, 9 - 13. ii. 1999, coll. F. Génier (1999 - 73) - 1 female (CMNC); same locality, 9 - 13. ii. 1999, coll. F. Génier (1999 - 74) - 1 female (CMNC); same locality, 9 - 13. ii. 1999, coll. R. Hanley (1999 - 78) - 1 female (CMNC); same locality, 9 - 13. ii. 1999, coll. F. Génier (1999 - 71) - 1 male (CMNC); same locality, 9 - 13. ii. 1999, coll. F. Génier (1999 - 75) - 1 male (CMNC); Villa Tunari, Hotel El Puente, elev. 357 m (16 ° 59.02 ' S, 65 ° 24.5 ' W), 15 - 27. xii. 2005, coll. S. & J. Peck (2005 - 45) - 1 male (CMNC); Provincia Chapare, Chimoré, elev. 250 m (16 ° 59 ' 56 '' S, 65 ° 9 ' 17 '' W), i. 1972, coll. A. Martínez - 2 males (CMNC); Regíon subandina, elev. 400 m, [no date], coll. Zischka - 1 male (CMNC); Villa Tunari, (16 ° 58 ' 48 '' S, 65 ° 25 ' 43 '' W), xii. 1985, coll. Martínez - 2 females, 2 males (CMNC); same locality, xi. 1992, coll. Arriagada - 1 female (CMNC); EL BENI, Guanay, (12 ° 31 ' 17 '' S, 66 ° 49 ' 39 '' W), xi. 1992, coll. L. Peña - 1 female (CMNC); LA PAZ, Chalalán, Río Tuichi, elev. 320 m (14 ° 25.6 ' S, 67 ° 55.2 ' W), 10 - 20. viii. 1995, coll. S. Spector - 1 male (CMNC); SANTA CRUZ, Hotel Flora y Fauna, 3.7 km SSE Buena Vista, (17 ° 29 ' S, 63 ° 33 ' W), 7 - 12. v. 2004, coll. A. R. Cline - 1 female (CMNC); Hotel Flora y Fauna, 4 - 6 km SSE Buena Vista, elev. 420 – 450 m (17 ° 29 ' S, 63 ° 33 ' W), 2 - 12. ii. 2000, coll. J. E. Wappes - 1 female, 1 male (CMNC); Hotel Flora y Fauna, 5 km SSE Buena Vista, elev. 440 m (17 ° 29.925 ' S, 63 ° 39.128 ' W), 6 - 15. xii. 2003, coll. S. & J. Peck (2003 - 130) - 1 female, 1 male (CMNC); same locality, 15 - 24. xii. 2003, coll. S. & J. Peck (2003 - 131) - 3 females, 1 male (CMNC); Provincia Ichilo, Buenavista, (17 ° 27 ' 32 '' S, 63 ° 39 ' 33 '' W), x. 1949, coll. Martínez - 1 female (CMNC); same locality, ii. 1950, coll. Martínez - 3 males (CMNC); same locality, iii. 1951, coll. Martínez - 1 male (CMNC); BRAZIL: AMAZONAS, Rio Parauary, (4 ° 36 ' S, 57 ° 47 ' W), 15. iii. 1937, coll. Zellibor-Hauff - 1 female (CMNC); MATO GROSSO, Fazenda São João, Municipio Diamantino, elev. 400 m (14 ° 14 ' 10 '' S, 56 ° 8 ' 11 '' W), 11. i. 2001, coll. Génier & Vaz de Mello (2001 - 04) - 3 females (CMNC); same locality, 13. i. 2001, coll. Génier & Vaz de Mello (2001 - 45) - 2 females (CMNC); same locality, 13. i. 2001, coll. Génier & Vaz de Mello (2001 - 46) - 1 female, 1 male (CMNC); PARÁ, Tucuruí, (3 ° 46 ' 26 '' S, 49 ° 41 ' 19 '' W), i. 1979, coll. Alvarenga - 1 female, 1 male (CMNC); COLOMBIA: META, Villavicencio, elev. 500 m (4 ° 9 ' 38 '' N, 73 ° 39 ' 43 '' W), 29. ii. - 5. iii. 1972, coll. S. & J. Peck - 1 female, 1 male (CMNC); ECUADOR: MORONA-SANTIAGO, Untsuants sitio 7, elev. 900 m, 23. i. 2002, coll. J. Celi & M. Ortega - 1 female, 1 male (CMNC); PERU: HUANUCO, Tingo María, elev. 700 m (7 ° 10 ' S, 76 ° 0 ' W), vii. 1974, coll. Bordón - 1 female, 1 male (CMNC); same locality, x. 1977, coll. P. Arnaud - 1 female, 1 male (CMNC); Universidad, Tingo María, (9 ° 10 ' S, 76 ° 0 ' W), xii. 1974, coll. Martínez - 8 females, 5 males (CMNC); same locality, vii. 1974, coll. Martínez - 1 male (CMNC); MADRE DE DIOS, Castañal Trail, Estación. Biológicas Pakitza, Parque Nacional del Manú, elev. 317 m (11 ° 56 ' 41 '' S, 71 ° 17 ' 0 '' W), 15 - 16. x. 2000, coll. R. Brooks (2000 - 13) - 1 female (CMNC); SAN MARTÍN, Alto Porongo, near Tocache, 18. x. 1976, coll. J. Schunke L. - 1 male (CMNC); Río Huallaga, near Tocache, (8 ° 11 ' 20 '' S, 76 ° 30 ' 51 '' W), 13. viii. 1975, coll. J. Schunke L. - 1 female (CMNC); UCAYALI, Lorento, Pucallpa, (8 ° 21 ' 57 '' S, 74 ° 33 ' 8 '' W), 11. ix. 1978, coll. J. Schunke L. - 1 male (CMNC); Pucallpa, (8 ° 23 ' 58 '' S, 74 ° 34 ' 39 '' W), 19. xi. 1974, coll. J. Schunke L. - 1 male (CMNC).	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA5DB7AFF69F9A6FA19CA2E.taxon	discussion	Remarks. Aside the characteristic shape of the parameres (Figs. 5 – 6), this species can be separated externally from all others by the shape of the 7 th interstrial carina, which overlaps the elytral lateral edge on most of the length.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA7DB7AFF69FE08FD7BCF4E.taxon	materials_examined	Type locality. Ecuador, provincia Los Ríos, Quevedo, Pichilingue, XII. 1978, A. Martínez.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA7DB7AFF69FE08FD7BCF4E.taxon	distribution	Distribution. Ecuador.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA7DB7AFF69FE08FD7BCF4E.taxon	materials_examined	Material examined (6 males, 15 females). ECUADOR: LOS RÍOS, Estación Experimental Tropical Pichilingue, Quevedo, (1 ° 0 ' 13 '' S, 79 ° 27 ' 50 '' W), v. 1976, coll. Martínez - 14 females, 5 males (including 19 paratypes) (CMNC); same locality, iv. 1976, coll. Martínez - 1 female (paratype) (CMNC); same locality, viii. 1976, coll. Martínez - 1 male (paratype) (CMNC).	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA7DB7AFF69FE08FD7BCF4E.taxon	discussion	Remarks. As for D. eurymedon, this species is known only from the original type specimens collected from a single locality. Interestingly, no additional specimens of this species were collected at the Estación Científica Río Palenque located only 50 km north of Estación Experimental Tropical Pichilingue with only a 200 m elevation difference despite extensive dung beetle trapping by H. F. Howden, S. B. Peck and others on several occasions from 1975 to 1979. Externally D. rosamariae is most closely related to D. eurymedon from which it can be separated by its smaller size and the shape of parameres (Figs. 7 – 8).	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA7DB74FF69FBE8FA0BCA20.taxon	materials_examined	Type locality. Jalapa, Mexico.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA7DB74FF69FBE8FA0BCA20.taxon	distribution	Distribution. Belize, Costa Rica, El Salvador, Guatemala, Mexico, Nicaragua, United States.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA7DB74FF69FBE8FA0BCA20.taxon	materials_examined	Material examined (69 males, 61 females, 1 specimen). BELIZE: 39 mi. W Hwy. at beaver dam, 6 - 12. viii. 1972, coll. S. & J. Peck - 1 female (CMNC); CAYO, Caves Branch, (17 ° 8 ' N, 88 ° 42 ' W), 23 - 29. viii. 1972, coll. S. & J. Peck - 1 male (CMNC); COSTA RICA: PUNTARENAS, R. Priv. Karen Morgensen, Send. El Viejo Níspero, 315 m, 23 Jun – 10 Jul 2003, D. Briceño, Tp. de Intersección, L _ N _ 205600 _ 420300 # 74609, 9.8670900 N 85.0600000 W - 1 specimen (INBio); GUATEMALA: BAJA VERAPAZ, 16.5 km N Salama, (15 ° 13 ' N, 90 ° 18 ' W), 3. vii. 1993, coll. F. Génier - 4 males (CMNC); PETÉN, Tikal, (17 ° 12 ' 46 '' N, 89 ° 37 ' 3 '' W), 23 - 26. viii. 1972, coll. S. & J. Peck - 1 female (CMNC); same locality, 28 - 30. vii. 1978, coll. Helava & Kukal - 2 females (CMNC); same locality, 12 - 28. v. 1991, coll. R. Limoges - 1 female (CMNC); SUCHITEPÉQUEZ, Cuyotenango, (14 ° 32 ' N, 91 ° 56 ' W), v. 1902, coll. Riedel - 1 male (CMNC); MEXICO: CAMPECHE, 87 mi. E Escárcega, elev. 244 m (18 ° 30 ' 55 '' N, 89 ° 29 ' 49 '' W), 14. viii. 1971, coll. A. Newton (337) - 1 female (CMNC); Chicaná, 10 km W Xpujil, (18 ° 30 ' 25 '' N, 89 ° 29 ' 11 '' W), 12 - 14. vii. 1983, coll. S. & J. Peck - 2 males (CMNC); CHIAPAS, 11 mi. E Trinitaria, elev. 1585 m (16 ° 8 ' 15 '' N, 91 ° 53 ' 43 '' W), 24. viii. 1971, coll. A. Newton (349) - 1 female, 1 male (CMNC); 4 mi. S Palenque, elev. 183 m (17 ° 27 ' 22 '' N, 91 ° 57 ' 59 '' W), 15. viii. 1971, coll. A. Newton (326) - 1 female (CMNC); 5 km E Ocozocoautla, (16 ° 44 ' 35 '' N, 93 ° 19 ' 42 '' W), 4. vi. 1990, coll. H. & A. Howden - 1 male (CMNC); Mirador La Coyota, Parque Nacional Cañón del Sumidero, elev. 1700 m (16 ° 49 ' 52 '' N, 93 ° 4 ' 27 '' W), 21. vi. 1989, coll. H. Howden - 1 male (CMNC); Parque Nacional Cañón del Sumidero, elev. 1000 m (16 ° 51 ' N, 93 ° 7 ' W), 25. v. - 1. vi. 1990, coll. B. Gill, H. & A. Howden - 2 females, 4 males (CMNC); same locality, 29. v. 1990, coll. H. & A. Howden - 1 female, 1 male (CMNC); same locality, 1. vi. 1990, coll. H. & A. Howden - 1 female (CMNC); same locality, 7. vi. 1990, coll. H. & A. Howden & B. Gill - 3 females, 1 male (CMNC); GUERRERO, 12.0 km SW Xochipala, elev. 1700 m (17 ° 41 ' N, 99 ° 38 ' W), 14. vii. 1992, coll. R. S. Anderson (1996 - 06) - 1 female (CMNC); 9 mi. NE Iguala, elev. 1341 m (18 ° 23 ' 39 '' N, 99 ° 28 ' 47 '' W), 29. viii. 1971, coll. A. Newton (378) - 1 female, 1 male (CMNC); JALISCO, 10 mi. SW Autlán, elev. 1280 m (19 ° 41 ' 48 '' N, 104 ° 23 ' 49 '' W), 19. ix. 1971, coll. A. Newton (422) - 1 male (CMNC); 16.5 mi. SW Autlán, elev. 701 m (19 ° 40 ' 6 '' N, 104 ° 25 ' 52 '' W), 12 - 20. ix. 1971, coll. A. Newton (420) - 1 male (CNC); 8 mi. W Atenquique, elev. 1798 m (19 ° 31 ' 45 '' N, 103 ° 31 ' 17 '' W), 10 - 18. ix. 1971, coll. A. Newton (410) - 4 females, 4 males (CMNC, CNC); MORELOS, Yautepec, (18 ° 53 ' N, 99 ° 4 ' W), 13. vii. 1963, coll. F. D. Parker & L. A. Stange - 1 female (CMNC); NUEVO LEÓN, 10 mi. W Linares, (24 ° 45 ' 51 '' N, 99 ° 41 ' 58 '' W), 31. v. 1971, coll. A. Newton (224) - 1 male (CMNC); S. Rosa Can., 29 km W Linares, (24 ° 44 ' 56 '' N, 99 ° 48 ' 33 '' W), 3 - 5. vi. 1983, coll. S. & J. Peck - 1 male (CMNC); OAXACA, 15 mi. S Sola de Vega, elev. 1829 m (16 ° 23 ' 18 '' N, 96 ° 58 ' 12 '' W), 30 - 31. v. 1971, coll. S. Peck - 2 females, 2 males (CMNC, CNC); 15 mi. S Sola de Vega on route 131, elev. 1829 m (16 ° 23 ' 18 '' N, 96 ° 58 ' 12 '' W), 30. v. 1971, coll. H. F. Howden - 1 female, 2 males (CMNC, CNC); 6 mi. S Valle Nacional, elev. 610 m (17 ° 43 ' 14 '' N, 96 ° 19 ' 2 '' W), 18 - 20. v. 1971, coll. H. Howden - 1 male (CMNC); QUIN- TANA ROO, 20 mi. S Felipe Carrillo Puerto, elev. 30 m (19 ° 17 ' 18 '' N, 88 ° 5 ' 40 '' W), 13. viii. 1971, coll. A. Newton (341) - 1 female, 1 male (CMNC); Felipe Carrillo Puerto, (19 ° 34 ' N, 88 ° 3 ' W), ix. 1984, coll. A. Martínez - 8 females, 9 males (CMNC); Kohunlich, 68 km W Chetumal, (18 ° 25 ' 11 '' N, 88 ° 47 ' 29 '' W), 15 - 17. vii. 1982, coll. S. & J. Peck - 1 female (CMNC); SAN LUIS POTOSÍ, 3 mi. W El Naranjo, (22 ° 31 ' 19 '' N, 99 ° 21 ' 49 '' W), 10 - 18. vi. 1971, coll. A. Newton (233) - 1 female, 2 males (CMNC, CNC); 50 km E Ciudad del Maíz, (22 ° 24 ' N, 99 ° 23 ' W), 12. vii. 1952, coll. F. Werner - 1 male (CMNC); 7 mi. W El Naranjo, elev. 732 m (22 ° 29 ' 54 '' N, 99 ° 22 ' 49 '' W), 10 - 18. vi. 1971, coll. A. Newton (235) - 1 female, 1 male (CMNC, CNC); 8 mi. E Xilitla, (21 ° 20 ' 19 '' N, 98 ° 51 ' 41 '' W), 28. vi. 1971, coll. A. Newton (252) - 1 female (CMNC); SINALOA, 15 mi. W El Palmito, elev. 1524 m (23 ° 34 ' N, 105 ° 57 ' W), 5. viii. 1964, coll. H. F. Howden - 1 female (paratype) (CNC); 31 mi. NE Concordia, elev. 1433 m (23 ° 28 ' 50 '' N, 105 ° 50 ' 19 '' W), 28. ix. 1971, coll. A. Newton (438) - 2 females, 2 males (CMNC); 8 mi. SW Concordia, elev. 91 m (23 ° 13 ' 22 '' N, 106 ° 8 ' 46 '' W), 28. ix. 1971, coll. A. Newton (430) - 2 females (CMNC); SONORA, Yécora, (28 ° 22 ' N, 108 ° 55 ' W), 20 - 22. v. 1961, coll. Gibson, Howden, Martin - 3 females, 2 males (including 4 paratypes) (CMNC, CNC); same locality, 12. vii. 1961, coll. W. W. Gibson - 2 females, 7 males (including holotype, 8 paratypes) (CMNC, CNC); TAMAULIPAS, El Abra S of Ciudad Mante, (23 ° 7 ' N, 98 ° 59 ' W), 7 - 9. vii. 1969, coll. S. & J. Peck - 2 females, 2 males (CMNC); El Nacimiento del Río Frío, Gómez Farías, (23 ° 1 ' 49 '' N, 99 ° 8 ' 52 '' W), 29. vi. 1969, coll. S. Peck - 1 female (CMNC); VERACRUZ, Lago de Catemaco, elev. 366 m (18 ° 21 ' 27 '' N, 95 ° 3 ' 36 '' W), 8 - 16. viii. 1960, coll. H. F. Howden - 2 females (CNC); same locality, 31. vii. - 4. viii. 1970, coll. A. Newton - 1 female (CMNC); Misantla, (19 ° 56 ' 10 '' N, 96 ° 49 ' 56 '' W), [nodate], coll. Hoge - 1 male (paralectotype) (CNC); Palma Sola, (21 ° 22 ' 24 '' N, 98 ° 17 ' 13 '' W), vi. 1972, coll. G. Halffter & P. Reyes - 1 female (CMNC); YUCA- TÁN, 3 mi. S Muná, (20 ° 26 ' 50 '' N, 89 ° 44 ' 6 '' W), viii. 1971, coll. A. Newton (346) - 1 male (CMNC).	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA7DB74FF69FBE8FA0BCA20.taxon	discussion	Remarks. After examining nearly 130 specimens from 42 unique localities, it must be concluded that although type specimens of D. scabriusculum montanum Howden vary slightly in the convexity of the elytra and in fine details of the parameres, the separation as a separate entity is hard to maintain. The variation, especially the length of the 7 th interstrial carina and the apical shape of the parameres (Figs. 9 – 10), is found in other populations. For example the length of the 7 th interstrial carina is extremely variable and recently examined specimens from a unique locality show a range covering the difference from D. s. scabriusculum and D. s. montanum. Similarly, the apical shape of the parameres is extremely variable, even within specimens from the same locality. All attempts to find a pattern with the variation have been unsuccessful. A much larger sample might be required. For the moment the following synonymy is proposed, the second epithet being valid: D. (Deltochilum) scabriusculum montanum Howden, 1966 = Deltochilum (Deltochilum) scabriusculum Bates, 1887 new synonymy. Externally, D. scabriusculum may be separated from all other species by the reduced 7 th interstrial carina, which covers the apical half to eight of the elytral length combined with the absence of the sternite 3 bulge in the male. The characteristic shape of the parameres (Figs. 9 – 10) and distinctly carinate 6 th interstria apically will separate it from D. tumidum Howden.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA9DB74FF69FE0FFE63CFC5.taxon	materials_examined	Type locality. Mexico, Sinalao, 5 miles north Mazatlán, 27. vii. 1964, at black light, Chemsak, Howden & Powell, (CNC No. 9146).	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA9DB74FF69FE0FFE63CFC5.taxon	distribution	Distribution. Mexico.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA9DB74FF69FE0FFE63CFC5.taxon	materials_examined	Material examined (7 males, 1 female). MEXICO: ESTADO DE MÉXICO, Tejupilco de Hidalgo [to] Temascaltepec de González, (18 ° 58 ' N, 100 ° 4 ' W), vi. 1933, coll. H. E. Hinton & R. L. Usinger - 2 males (including 2 paratypes) (CMNC, CNC); SINALOA, 26 mi. NE Villa Unión, elev. 305 m (23 ° 29 ' N, 106 ° 10 ' W), 17. viii. 1963, coll. E. Sleeper, W. Agnew, G. Noonan & P. Sullivan - 1 female (paratype) (CNC); 5 mi. N Mazatlán, (23 ° 21 ' N, 106 ° 25 ' W), 27. vii. 1964, coll. H. F. Howden, Chemsak & Powell - 1 male (holotype) (CNC); Rio Piaxtla, 29. vi. 1962, coll. [anonymous] - 1 male (paratype) (CNC); Villa Unión, elev. 700 m (23 ° 12 ' N, 106 ° 14 ' W), vii. 1968, coll. A. Martínez - 3 males (CMNC).	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
01017A38FFA9DB74FF69FE0FFE63CFC5.taxon	discussion	Remarks. Separated from other species by the shape of the parameres (Figs. 11 – 12) and the strongly reduced and blunt 7 th interstrial carina.	en	Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.213648
