taxonID	type	description	language	source
6AD4F4F54C665DC4BCE076319828A672.taxon	description	Figs 4, 5, Table 2, Suppl. material 2	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
6AD4F4F54C665DC4BCE076319828A672.taxon	description	Measurements (in mm) and body mass (in g) of the holotype. HB: 43.25, EL: 16.18, EW: 8.61, TRL: 8.67, TRW: 2.29, HFL: 8.64, FL: 31.94, TIB: 18.42, GTL: 15.68, CCL: 12.50, BCW: 7.57, BCH: 7.78, ZYW: 8.20, MAW: 7.58, IOW: 4.24, CM 3 L: 4.98, C 1 C 1 W: 3.65, M 3 M 3 W: 4.61, RCM: 0.79, CM 3 L: 5.92, ML: 11.52, MDL: 11.98, CPH: 4.38; BW: 6.20.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
6AD4F4F54C665DC4BCE076319828A672.taxon	etymology	Etymology. The specific epithet beibengensis refers to the type locality of the new species: Beibeng Township, Medog County, Nyingchi City, Xizang Autonomous Region, China. We propose the common English name “ Beibeng Tube-nosed Bat ” and the Chinese name “ Bèi Bēng Guǎn Bí Fú (背崩管鼻蝠) ”.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
6AD4F4F54C665DC4BCE076319828A672.taxon	diagnosis	Diagnosis. Murina beibengensis sp. nov. can be distinguished from all other congeners by the following combination of characters: (1) small-size Murina, FL 31.94 mm, GTL 15.68 mm; (2) dorsal fur orangish-yellow overall, grey at the base, gradually transitioning to orange-yellow tips halfway from the base; (3) ventral fur silvery-gray overall, dark grey at the base, with silvery-gray tips; (4) ears broadly rounded, with smoothly convex anterior margins, no notch on posterior margins; (5) forearm and wrists covered with sparse hairs; (6) wing attachment point located at 1 / 3 from base of claw to base of toe; (7) sagittal and lambdoid crests well developed; (8) I 2 is situated laterally anterior to I 3 and partially visible in the lateral view, and I 2 slightly taller than I 3; (9) mesostyles of M 1 and M 2 are reduced; (10) C 1 taller than P 4, P 2 smaller than P 4 in height, and crown area of P 2 larger than 2 / 3 that of P 4; (11) P 2 approximately equal to P 4 in height, with a basal area 2 / 3 that of P 4; (12) C 1 larger than P 4 in height and basal area; (13) mandibular foramina clearly visible, situated below P 2.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
6AD4F4F54C665DC4BCE076319828A672.taxon	description	Description. Morphometric data of Murina beibengensis sp. nov. are provided in Table 2 and Suppl. material 2. Small-size Murina, HB 43.25 mm, FL 31.94 mm, EL 16.18 mm, HFL 8.64 mm, and BW 6.20 g. Nostrils tubular, open sideways, and slightly short. Ears small, short, oval, blunt at tips, with smoothly convex anterior margins, not notched on posterior margin. Tragus short, narrow, and tapering toward a pointed tip, with slightly convex anterior margin, concave posterior margin, and basal notch, and slightly curved outward, about half as long as ear. Body covered with thick and fluffy hair. Dorsal hairs orangish-yellow overall (bicolored, grayish black at the base, gradually transitioning to orange-yellow tips halfway from base). Dorsal hairs extend onto bases of wings; uropatagium, thumbs, forearm, wrist, tibia, and feet, with well-developed fringe of hairs around margin of uropatagium. Densely furred anterior 1 / 3 of the dorsal uropatagium, posterior 2 / 3 covered with sparse hairs. Ventral hairs are silvery-gray overall, tricolored, gray-white at the base, central dark gray, with silvery-gray tips. Flesh purple around the eyes, muzzle, and lower forehead, and the face is hairy except for the long, protuberant nostrils that are naked. TL 32.69 mm, tip of the tail extending significantly past the rear edge of the uropatagium, tip slightly free. Plagiopatagium attachment point located at the middle of the claw of the first toe. Skull robust and domed, relatively small, GTL 15.68 mm. Rostrum long, deep, gradually ascending to forehead; prominent median depression present. Sagittal and lambdoid crests well developed. In dorsal view, braincase nearly rounded; zygomatic arches weak and slender, gradually widening posteriorly, widest at the base of the zygomatic arches; posterior margin of skull not protruding; middle from snout to frontal region distinctly concave downward. In lateral view, skull slightly elongated, with elongated oval braincase; height gradually rising from snout to parietal, with slightly increasing slope from snout to frontal and decreasing slope from frontal to parietal; slight depression between snout and frontal, without distinct prominence at frontal; zygomatic arches gradually rising from anterior to posterior, highest at the middle of zygomatic arches. In ventral view, palatine wide and nearly flat, ending at midpoint of C 1; basisphenoid pits slightly shallowly teardrop-shaped, extending posteriorly to anterior third of cochlea. Mandible length 11.52 mm, inverted L-shaped. Line between coronoid process and condyle nearly flat; distinct inward depression between condyle and angle; angle slightly long and wide; mandibular foramina clearly visible, situated below P 2. Dental morphology: Dental formula is I 2 / 3, C 1 / 1, P 2 / 2, M 3 / 3 = 34 (Fig. 5). In the maxilla, I 2 is situated laterally anterior to I 3 and partially visible in the lateral view; the crown area of P 2 larger than 2 / 3 that of P 4 and smaller than that of C 1. Based on these characters, the species belongs to the “ cyclotis - type ”. Maxillary dentition converges slightly anteriorly (RCM = 0.79). I 2 and I 3 bicuspid, smaller secondary cusp situated posterior to primary cusp; I 2 is slightly taller than I 3, with a crown area half that of I 3; distinct gap between posterior of I 3 and C 1, not in contact, about 1 / 3 of height of C 1. C 1 taller than P 4, slightly elongated and lacking secondary cusps, crown area approximately equal to P 4; P 2 is slightly smaller, delicate and pointed, about 4 / 5 as high as P 4 and 2 / 3 as high as C 1, crown area of P 2 is 2 / 3 that of P 4. Mesostyles of M 1 and M 2 are reduced, but retaining distinct cusps; paracone, protocone, metacone, and parastyle well developed. M 3 are reduced, with only parastyle, paracone, and protocone. In the mandible, the first, second, and third lower incisors (I 1, I 2, and I 3) smaller, tricuspid, nearly equal in height and width; slight overlap of outer cusps of I 1, I 2, and I 3 with gradual increase in height from I 1 to C 1. C 1 contains pointed cusp on anterior inner margin, in contact with I 3 outer cusp, exceeding P 2 and P 4 in height and basal area; C 1 larger than P 4 in height and basal area. P 2 nearly equal to P 4 in height, and the basal and crown area of P 2 approximately 2 / 3 that of P 4. In lateral view, trigonids of M 1, M 2, and M 3, metaconid, and paraconid approximately 2 / 3 as high as protoconid. Talonids of M 1 and M 2 are approximately half the size of trigonids; entoconid and hypoconid distinctly separated from trigonid, nearly equal to metaconid and paraconid in height. M 1 and M 2 are nyctalodont types, with well-developed entoconids. M 3 reduced, talonid approximately 1 / 3 as long as trigonid, paraconid, protoconid, and metaconid complete and well developed.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
362A7E5E02915BE1A4F4515A47F3A554.taxon	description	Figs 6, 7, 8, 9, Table 3, Suppl. material 2	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
362A7E5E02915BE1A4F4515A47F3A554.taxon	description	Measurements (in mm) and body weight (in g) of the holotype. HB: 38.46, EL: 11.89, EW: 6.79, TRL: 6.19, TRW: 1.94, HFL: 6.94, FL: 29.75, TIB: 15.47, GTL: 15.01, CCL: 14.09, BCW: 7.55, BCH: 6.71, ZYW: 8.74, MAW: 7.11, IOW: 4.39, CM 3 L: 4.82, C 1 C 1 W: 3.61, M 3 M 3 W: 5.07, RCM: 0.712, CM 3 L: 5.86, ML: 9.1, MDL: 10.21, CPH: 3.42; BW: 4.20.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
362A7E5E02915BE1A4F4515A47F3A554.taxon	etymology	Etymology. The specific epithet medogensis refers to the type locality of the new species: Medog Town, Medog County, Nyingchi City, Xizang Autonomous Region, China. We propose the common English name “ Medog Tube-nosed Bat ” and the Chinese name “ Mò Tuō Guǎn Bí Fú (墨脱管鼻蝠) ”.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
362A7E5E02915BE1A4F4515A47F3A554.taxon	diagnosis	Diagnosis. Murina medogensis sp. nov. can be distinguished from all of the other congeners by the following combination of characters: (1) small-size Murina, FL 29.75 – 31.24 mm, GTL 15.01 – 18.74 mm; (2) dorsal fur dark grayish overall, dark grey at the base, gradually transitioning to gray-brown / white tips 2 / 3 from the base; (3) ventral fur silvery-gray overall, dark grey at the base, with silvery-gray tips; (4) ears broadly rounded, with smoothly convex anterior margins, no notch on posterior margins; (5) forearm and wrists without covered sparse hairs; (6) wing attachment point located at 1 / 3 from base of claw to base of toe; (7) sagittal crest absent, lambdoid crest present and poorly developed; (8) I 2 is situated laterally anterior to I 3 and partially visible in the lateral view, I 2 equal to I 3 in height; (9) mesostyles of M 1 and M 2 are well developed; (10) C 1 slightly lower than P 4 in height and crown area of P 2 larger than 2 / 3 that of P 4; (11) P 2 is approximately equal to P 4 in height, with a basal area 2 / 3 that of P 4; (12) C 1 taller than P 4 in height, crown area equal to P 2 and 2 / 3 of P 4; (13) mandibular foramina clearly visible, situated below P 4; (14) without distinct prominence at frontal aspect of skull.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
362A7E5E02915BE1A4F4515A47F3A554.taxon	description	Description. Morphometric data of Murina medogensis sp. nov. are provided in Table 3 and Suppl. material 2. Small-size Murina, HB 37.68 – 38.46 mm, FL 29.75 – 31.24 mm, EL 11.89 – 14.06 mm, HFL 6.94 – 8.52 mm, and BW 4.20 g. Nostrils tubular, open sideways, and slightly short. Ears small, short, oval, blunt at tips, with smoothly convex anterior margins, not notched on posterior margin. Tragus short, narrow, and tapering toward pointed tips, with slightly convex anterior margin, concave posterior margin, and basal notch, and it curves outwards slightly, about half as long as ear. Body covered with thick and fluffy hair. Dorsal hairs dark grayish overall (bicolored, dark grey at the base, gradually transitioning to gray-brown / white tips 2 / 3 from base). Dorsal hairs extend onto bases of wings, uropatagium, thumbs, tibia, and feet, with slight-developed fringe of hairs around margin of uropatagium. Densely furred anterior 1 / 3 of the dorsal uropatagium, posterior 2 / 3 covered with sparse hairs. Ventral hairs are silvery-gray overall, bicolored, dark grey at the base, with silvery-gray tips. Flesh-purple around the eyes, muzzle, and lower forehead, and the face is hairy except for the long, protuberant nostrils that are naked. TL 34.54 – 36.37 mm, slightly shorter than head-body length, tip of the tail extending significantly past the rear edge of the uropatagium, tip slightly free. Plagiopatagium attachment point located at about 1 / 3 from base of claw to base of toe, near base of claw (Fig. 6 F, G). Skull robust and domed, relatively small, GTL 15.01 – 18.74 mm. Rostrum long, deep, gradually ascending to forehead; prominent median depression present. Sagittal crest absent, lambdoid crest present, poorly developed. In dorsal view, braincase nearly rounded; zygomatic arches weak and slender, gradually widening posteriorly, widest at the base of the zygomatic arches; posterior margin of skull slightly protruding; middle from snout to frontal region distinctly concave downward. In lateral view, skull slightly elongated, with elongated oval braincase; height gradually rising from snout to parietal, with slightly increasing slope from snout to frontal and decreasing slope from frontal to parietal; slight depression between snout and frontal, without distinct prominence at frontal; zygomatic arches gradually rising from anterior to posterior. In ventral view, palatine wide and nearly flat, ending at posterior margin of C 1; basisphenoid pits slightly shallowly teardrop-shaped, extending posteriorly to anterior third of cochlea. Mandible length 9.1 mm, inverted L-shaped. Line between coronoid process and condyle nearly flat; distinct inward depression between condyle and angle; angle slightly long and wide; mandibular foramina clearly visible, situated below P 4. Dental morphology: Dental formula is I 2 / 3, C 1 / 1, P 2 / 2, M 3 / 3 = 34 (Figs 7, 8). In the maxilla, I 2 is situated laterally anterior to I 3 and partially visible in the lateral view; crown area of P 2 larger than 2 / 3 that of P 4 and smaller than that of C 1. Based on these characters, the species belongs to the “ cyclotis - type ”. Maxillary dentition converges slightly anteriorly (RCM = 0.71). I 2 and I 3 bicuspid, smaller secondary cusp situated posterior to primary cusp; I 2 equal to I 3 in height, crown area of I 2 less than half that of I 3; distinct gap between posterior surface of I 3 and C 1, not in contact, about half of height of C 1. C 1 slightly lower than P 4 in height, slightly elongated and lacking secondary cusps, crown area 2 / 3 that of P 4; P 2 slightly smaller, delicate and pointed, about half as high as P 4 and 2 / 3 as high as C 1, crown area 2 / 3 that of P 4. Mesostyles of M 1 and M 2 are reduced, but retaining distinct cusps; paracone, protocone, metacone, and parastyle well developed. M 3 reduced, with only parastyle, paracone, and protocone. In the mandible, I 1, I 2, and I 3 smaller, tricuspid, almost equal in height and width; slight overlap of outer cusps of I 1, I 2, and I 3; with gradual increase in height from I 1 to C 1. C 1 contains pointed cusp on anterior inner margin, in contact with I 3 outer cusp, exceeding P 2 and P 4 in height; C 1 equal to P 2 in basal area but smaller than P 4. P 2 slightly smaller than P 4 in height, and the basal and crown area of P 2 approximately 2 / 3 that of P 4. In lateral view, trigonids of M 1, M 2, and M 3, metaconid, and paraconid approximately 2 / 3 as high as protoconid in height. Talonids of M 1 and M 2 are slightly half the size of trigonids; entoconids and hypoconids distinctly separated from trigonids, lower than metaconid and paraconid. M 1 and M 2 are nyctalodont types, with well-developed entoconids. M 3 reduced, talonid approximately 1 / 3 as long as trigonid, paraconid, protoconid, and metaconid complete and well developed.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
43CF00CF05115D4EA66CE313F7025B21.taxon	description	Figs 10, 11, 12, Table 4, Suppl. material 2	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
43CF00CF05115D4EA66CE313F7025B21.taxon	description	Measurements (in mm) and body weight (in g) of the holotype. HB: 34.68, EL: 12.74, EW: 7.89, TRL: 7.81, TRW: 5.35, HFL: 5.31, FL: 30.02, TIB: 12.75, GTL: 14.07, CCL: 12.01, BCW: 7.03, BCH: 6.49, ZYW: 8.07, MAW: 6.68, IOW: 4.40, CM 3 L: 4.44, C 1 C 1 W: 2.21, M 3 M 3 W: 4.97, RCM: 0.44, CM 3 L: 4.53, ML: 9.06, MDL: 9.46, CPH: 3.20; BW: 3.36.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
43CF00CF05115D4EA66CE313F7025B21.taxon	etymology	Etymology. The specific epithet milinensis refers to the type locality of the new species: Nanyi Lhoba Township, Milin County, Nyingchi City, Xizang Autonomous Region, China. We propose the common English name “ Milin Tube-nosed Bat ” and the Chinese name “ Mǐ Lín Guǎn Bí Fú (米林管鼻蝠) ”.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
43CF00CF05115D4EA66CE313F7025B21.taxon	diagnosis	Diagnosis. Murina milinensis sp. nov. can be distinguished from all of the other congeners by the following combination of characters: (1) small-size Murina, FL 28.84 – 34.01 mm, GTL 14.07 – 14.27 mm; (2) dorsal fur brown-gold overall, black at the base, gradually transitioning to brown-gold tips 2 / 3 from the base; (3) ventral fur pale overall, dark black at the base, with grayish white at the tips; (4) ears narrow and oval, without smoothly convex anterior margins, no notch on posterior margins; (5) forearm and wrists covered with sparse hairs; (6) wing attachment point located at 1 / 3 from base of claw to base of toe; (7) sagittal crest absent, lambdoid crest present and poorly developed; (8) I 2 is situated anterior to I 3 and clearly visible in the lateral view, I 2 equal to I 3 in height; (9) mesostyles of M 1 and M 2 are slightly reduced; (10) C 1 less than P 4 in height, crown is about 2 / 3 of P 4; (11) P 2 approximately half of P 4 in height, crown area of P 2 larger than 2 / 3 that of P 4; (12) C 1 equal to P 4 in height and crown area; (13) mandibular foramina clearly visible, situated below anterior margin of P 4.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
43CF00CF05115D4EA66CE313F7025B21.taxon	description	Description. Morphometric data of Murina milinensis sp. nov. are provided in Table 4 and Suppl. material 2. Small-size Murina, HB 29.23 – 39.29 mm, FL 28.84 – 34.01 mm, EL 12.48 – 14.58 mm, HFL 5.31 – 9.18 mm, and BW 3.15 – 7.50 g. Nostrils tubular, open sideways, and slightly longer. Ears small, short, and narrow, oval, blunt at tips, with smoothly convex anterior margins, not notched on posterior margin. Tragus short, narrow, and tapering toward pointed tip, with slightly convex anterior margin, concave posterior margin, and basal notch, and it curves outwards slightly, about half as long as ear. Body covered with thick and fluffy hair. Dorsal hairs brown-gold overall (bicolored, black at the base, gradually transitioning to brown-gold tips from 2 / 3 from base). Dorsal hairs extend onto bases of wings, uropatagium, thumbs, forearm, wrist, tibia, and feet, with slight-developed fringe of hairs around margin of uropatagium. Densely furred anterior 1 / 3 of the dorsal uropatagium, posterior 2 / 3 covered with sparse hairs. Ventral hairs are pale overall, bicolored, bicolored, dark black at the base and grayish white at the tips. Dark flesh-purple around the eyes, muzzle, and lower forehead, and the face is hairy except for the long, protuberant nostrils that are naked. TL 24.51 – 30.32 mm, slightly shorter than head-body length, tip of the tail extending significantly past the rear edge of the uropatagium, tip distinctly free. Plagiopatagium attachment point located at 1 / 3 from base of claw to base of toe, near base of claw (Fig. 10 F). Skull robust, nearly oval, relatively small, GTL 14.07 – 14.27 mm. Rostrum long, deep, gradually ascending to forehead; prominent median depression present. Sagittal crest absent, lambdoid crest present, poorly developed. In dorsal view, braincase nearly domed; zygomatic arches weak and slender, gradually widening posteriorly, widest at the base of the zygomatic arches; posterior margin of skull slightly protruding; middle from snout to frontal region distinctly concave downward. In lateral view, skull slightly elongated, with elongated oval braincase; height gradually rising from snout to parietal, with slightly increasing slope from snout to frontal and decreasing slope from frontal to parietal; slight depression between snout and frontal, with slight distinct prominence at frontal; zygomatic arches gradually rising from anterior to posterior. In ventral view, palatine wide and nearly flat, ending at posterior margin of C 1; basisphenoid pits slightly shallowly tadpole-shaped, extending posteriorly to anterior half of cochlea. Mandible length 8.77 – 9.42 mm, inverted L-shaped in lateral view. Line between coronoid process and condyle nearly flat; distinct inward depression between condyle and angle; angle short and wide; mandibular foramina clearly visible, situated below anterior margin of P 4. Dental morphology: Dental formula is I 2 / 3, C 1 / 1, P 2 / 2, M 3 / 3 = 34 (Figs 11, 12). In the maxilla, I 2 is situated anterior to I 3, and I 2 clearly visible laterally; crown area of P 2 approximately 1 / 3 that of P 4 and slightly smaller than C 1. Based on these characters, the species belongs to the “ suilla - type ”. Maxillary dentition converges slightly anteriorly (RCM = 0.44 – 0.70). I 2 and I 3 bicuspid, smaller secondary cusp situated posterior to primary cusp; I 2 almost equal to I 3 in height, crown area of I 2 half that of I 3; distinct gap between posterior surface of I 3 and C 1, not in contact, about half of height of C 1. C 1 less than P 4 in height, slightly elongated and lacking secondary cusps, crown area half that of P 4; P 2 slightly smaller, delicate and pointed, about half as high as P 4 and 2 / 3 as high as C 1, and crown area of P 2 is half that of P 4 and slightly smaller than C 1. Mesostyles of M 1 and M 2 are reduced, but retaining distinct cusps; paracone, protocone, metacone, and parastyle well developed. M 3 reduced, with only parastyle, paracone, and protocone. In the mandible, I 1, I 2, and I 3 smaller, tricuspid, almost equal in height and width; slight overlap of outer cusps of I 1, I 2, and I 3; with gradual increase in height from I 1 to C 1. C 1 contains pointed cusp on anterior inner margin, in contact with I 3 outer cusp. C 1 taller than P 2 and P 4 in height, with a basal area larger than P 2 but smaller than P 4. P 2 equal to P 4 in height, and the basal and crown areas are about half of P 4. In lateral view, trigonids of M 1, M 2, and M 3 clearly bicuspid, metaconid, and paraconid approximately 2 / 3 as high as protoconid in height. Talonid of M 1 and M 2 is slightly half the size of trigonid; entoconid and hypoconid distinctly separated from trigonid, lower than metaconid and paraconid, nearly equal to metaconid and paraconid in height. M 1 and M 2 are nyctalodont types, with well-developed entoconids. M 3 reduced, talonid approximately 1 / 3 as long as trigonid, paraconid, protoconid, and metaconid complete and well developed.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
9AA00AF9954B538FA2AA1317B0560C78.taxon	description	Figs 15, 16, 17, Table 5, Suppl. material 2	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
9AA00AF9954B538FA2AA1317B0560C78.taxon	description	Measurements (in mm) holotype. FL: 37.50, GTL: 18.2 (Thomas 1914).	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
9AA00AF9954B538FA2AA1317B0560C78.taxon	etymology	Etymology. The specific epithet “ rubella ” comes from the Latin word rubellus, meaning “ slightly reddish ” or “ reddish ”. The common English name “ Fujian Tube-nosed Bat ” and the Chinese name “ Fú Jiàn Guǎn Bí Fú (福建管鼻蝠) ”.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
9AA00AF9954B538FA2AA1317B0560C78.taxon	diagnosis	Diagnosis. Murina rubella can be distinguished from all of the other congeners by the following combination of characters: (1) larger body size, FL 34.24 – 38.45 mm, GTL 17.08 – 18.51 mm; (2) dorsal fur tan overall, pale brown at the base, gradually transitioning to dark tan tips from 2 / 3 from base; (3) ventral fur uniformly pale brown overall, black at the base, gray-white at tips; (4) ears narrow and oval, with smoothly convex anterior margins, no notch on posterior margins; (5) forearm and wrists covered with sparse hairs; (6) wing attachment point located at 1 / 3 from base of claw to base of toe; (7) without off-white circumferential band around the neck; (8) sagittal and lambdoid crests absent; (9) I 2 is situated anterior to I 3 and clearly visible in lateral view, I 2 slightly smaller than I 3 in height; (10) mesostyles of M 1 and M 2 are slightly reduced; (11) C 1 slightly smaller than P 4 in height, crown is about 2 / 3 of P 4; (12) P 2 approximately half of P 4 in height, crown area 2 / 3 that of P 4; (13) C 1 equal to P 4 in height and crown area.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
9AA00AF9954B538FA2AA1317B0560C78.taxon	distribution	Geographic distribution. Based on this study, phylogenetic evidence, and previous literature, M. rubella can be definitively recorded as distributed in China (Anhui, Fujian, Guangxi, Guangdong, Hubei, Hunan, Jiangxi, Zhejiang, Xizang) (Zhang 1997; Wang 2003; Francis et al. 2010; Zhou et al. 2011; Jiang 2015; Zhang et al. 2016; Huang et al. 2018; Moratelli et al. 2019; Yu et al. 2020; Jiang 2021; Liao et al. 2023; Qin et al. 2023), northern Laos (Francis et al. 2010; Francis and Eger 2012; Nguyen et al. 2015 b), and Vietnam (Francis and Eger 2012; Tu et al. 2015; Tu et al. 2016). Although M. huttoni has also been recorded in northern Thailand, northern Myanmar, and Peninsular Malaysia (Moratelli et al. 2019), the authenticity of these records (Francis and Eger 2012) requires further evidence due to the lack of available genetic data. The M. huttoni sensu stricto may be confined to the periphery of the Himalayas (Moratelli et al. 2019).	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
015E85F518C256EDB5C021312CAA81AE.taxon	description	Figs 13, 14, Table 4, Suppl. material 2	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
015E85F518C256EDB5C021312CAA81AE.taxon	description	Measurements (in mm) and body weight (in g) of the holotype. HB: 35.54, EL: 14.02, EW: 6.20, TRL: 7.48, TRW: 2.14, HFL: 6.07, FL: 30.29, TIB: 13.57, GTL: 13.73, CCL: 12.61, BCW: 6.36, BCH: 5.66, ZYW: 8.72, MAW: 7.04, IOW: 3.51, CM 3 L: 5.31, C 1 C 1 W: 2.45, M 3 M 3 W: 3.80, RCM: 0.64, CM 3 L: 5.74, ML: 9.22, MDL: 9.49, CPH: 3.61; BW: 5.68.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
015E85F518C256EDB5C021312CAA81AE.taxon	etymology	Etymology. The specific epithet yadongensis refers to the type locality of the new species: Xiayadong Township, Yadong County, Shigatse City, Xizang Autonomous Region, China. We propose the common English name “ Yadong Tube-nosed Bat ” and the Chinese name “ Yà Dōng Guǎn Bí Fú (亚东管鼻蝠) ”.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
015E85F518C256EDB5C021312CAA81AE.taxon	diagnosis	Diagnosis. Murina yadongensis sp. nov. can be distinguished from all of the other congeners by the following combination of characters: (1) small-size Murina, FL 28.81 – 31.84 mm, GTL 13.73 – 14.11 mm; (2) dorsal fur brown-gold, black at the base, gradually transitioning to brown-gold tips 2 / 3 from base; (3) ventral fur grayish brown, black at the base, gray-white at tips; (4) ears narrow and oval, without smoothly convex anterior margins, no notch on posterior margins; (5) forearm and wrists covered with sparse hairs; (6) wing attachment point located at 1 / 3 from base of claw to base of toe; (7) an off-white circumferential band around the neck; (8) sagittal crest absent, lambdoid crest present, poorly developed; (9) I 2 is situated anterior to I 3 and clearly visible in lateral view, I 2 equal to I 3 in height; (10) mesostyles of M 1 and M 2 are slightly reduced; (11) C 1 taller than P 4, crown is about 2 / 3 of P 4; (12) P 2 exceeded 2 / 3 of P 4 in height, and the crown area is about 2 / 3 of P 4; (13) C 1 equal to P 4 in height and crown area. and crown area of P 2 larger than 2 / 3 that of P 4.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
015E85F518C256EDB5C021312CAA81AE.taxon	description	Description. Morphometric data of Murina yadongensis sp. nov. are provided in Table 4 and Suppl. material 2. Small-size Murina, HB 28.68 – 35.54 mm, FL 28.81 – 31.84 mm, EL 11.40 – 14.20 mm, HFL 6.07 – 8.55 mm, and BW 5.68 – 7.00 g. Nostrils tubular, open sideways, and slightly longer. Ears small, short, and narrow, oval, blunt at tips, without smoothly convex anterior margins, not notched on posterior margin. Tragus short, narrow, and tapering toward pointed tip, with slightly convex anterior margin, concave posterior margin, and basal notch, and it curves outwards slightly, about half as long as ear. Body covered with thick and fluffy hair. Dorsal hairs brown-gold (bicolored, black at the base, gradually transitioning to brown-gold tips from 2 / 3 from base). Dorsal hairs extend onto bases of wings, uropatagium, thumbs, forearm, wrist, tibia, and feet, with slight-developed fringe of hairs around margin of uropatagium. Densely furred anterior 1 / 3 of the dorsal uropatagium, posterior 2 / 3 covered with sparse hairs. Ventral hairs grayish brown overall, bicolored, black at the base, gray-white at tips. An off-white circumferential band around the neck. Dark flesh-purple around the eyes, muzzle, and lower forehead, and the face is hairy except for the long, protuberant nostrils that are naked. TL 28.32 – 32.12 mm, slightly shorter than head-body length, tip of the tail extending significantly past the rear edge of the uropatagium, tip distinctly free. Plagiopatagium attachment point located at 1 / 3 from base of claw to base of toe, near base of claw (Fig. 13 E, F). Skull robust, nearly oval, relatively small, GTL 13.73 – 14.11 mm. Rostrum long, deep, gradually ascending to forehead; prominent median depression present. Sagittal and lambdoid crests absent. In dorsal view, braincase domed; zygomatic arches weak and slender, gradually widening posteriorly, widest at the base of the zygomatic arches; posterior margin of skull slightly protruding; middle from snout to frontal region distinctly concave downward. In lateral view, skull slightly elongated, with elongated oval braincase; height gradually rising from snout to parietal, with slightly increasing slope from snout to frontal and decreasing slope from frontal to parietal; slight depression between snout and frontal, with slight distinct prominence at frontal; zygomatic arches gradually rising from anterior to posterior. In ventral view, palatine wide and nearly flat, ending at posterior margin of C 1; basisphenoid pits slightly shallowly tadpole-shaped, extending posteriorly to anterior half of cochlea. Mandible length 8.85 – 9.22 mm, inverted L-shaped in lateral view. Line between coronoid process and condyle nearly flat; distinct inward depression between condyle and angle; angle short and wide; mandibular foramina clearly visible, situated below anterior margin of P 4. Dental morphology: Dental formula is I 2 / 3, C 1 / 1, P 2 / 2, M 3 / 3 = 34 (Fig. 14). In the maxilla, I 2 is situated anterior to I 3, and I 2 clearly visible laterally; crown area of P 2 approximately half that of P 4 and slightly smaller than C 1. Based on these characters, the species belongs to the “ suilla - type ”. Maxillary dentition converges slightly anteriorly (RCM = 0.64 – 0.66). I 2 and I 3 bicuspid, smaller secondary cusp situated posterior to primary cusp; I 2 almost equal to I 3 in height, with crown area approximately half that of I 3; distinct gap between posterior surface of I 3 and C 1, not in contact, about half of height of C 1. C 1 smaller than P 4 in height, slightly elongated and lacking secondary cusps, crown area 2 / 3 that of P 4; P 2 slightly smaller, delicate and pointed, about half as tall as P 4 and C 1, with crown area of P 2 is half that of P 4 and slightly smaller than C 1. Mesostyles of M 1 and M 2 are reduced, but retaining distinct cusps; paracone, protocone, metacone, and parastyle well developed. M 3 reduced, with only parastyle, paracone, and protocone. In the mandible, I 1, I 2, and I 3 smaller, tricuspid, almost equal in height and width; overlap of outer cusps of I 1, I 2, and I 3; with rapid increase in height from I 3 to C 1. C 1 without pointed cusp on anterior inner margin, not in contact with I 3 outer cusp. C 1 almost equal to P 4 in height and basal area, about twice as much as P 2. P 2 exceeded 4 / 5 of P 4 in height, and the basal and crown areas are about 2 / 3 of P 4. In lateral view, trigonids of M 1, M 2, and M 3, metaconidhe slightly equal to protoconid in height, paraconid about 1 / 3 of protoconid. Talonid of M 1 and M 2 almost the same size as trigonid; entoconid and hypoconid distinctly separated from trigonid, lower than metaconid and paraconid, nearly equal to metaconid and paraconid in height. M 1 and M 2 are nyctalodont types, with well-developed entoconids. M 3 reduced, talonid approximately 1 / 3 as long as trigonid, paraconid, protoconid, and metaconid complete and well developed.	en	Luo, Tao, Mao, Ming-Le, Lan, Chang-Ting, Zhao, Zi-Fa, Wang, Zhong-Lian, Yu, Jing, Wang, Jia-Jia, Yan, Chen-Rui, Xiao, Ning, Zhou, Jiang (2025): Four new tube-nosed bat species of the genus Murina (Chiroptera, Vespertilionidae) from Xizang Autonomous Region, China, based on morphological and molecular data. Zoosystematics and Evolution 101 (3): 1023-1055, DOI: 10.3897/zse.101.144375
