taxonID	type	description	language	source
002087EAFFA5947864A8FCCFFEA9FD52.taxon	description	(Figs. 1, 7 – 10)	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA5947864A8FCCFFEA9FD52.taxon	materials_examined	Type material. Holotype ♁ (Fig. 1): ‘ PAPUA N. GUINEA | Onerunka VIII. 79 [handwritten in black] | nr Kainantu | W. G. Ullrich` <printed>, ‘ HOLOTYPE | Paraphloeostiba | conjuncta sp. n. | Shavrin A. V. des. 2023`<red, printed> (MHNG). Paratypes: 5 ♁♁ (three specimens dissected), 2 unsexed specimens (without apical segment of abdomen): same data as the holotype, with additional printed red label: ‘ PARATYPE | Paraphloeostiba | conjuncta sp. n. | Shavrin A. V. des. 2023 (2 ♁♁: cSh; 3 ♁♁, 2 unsexed specimens: MHNG).	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA5947864A8FCCFFEA9FD52.taxon	description	Description. Measurements (n = 8): HW: 0.37 – 0.41; HL: 0.21 – 0.24; OL: 0.10 – 0.12; TL: 0.02; AL (holotype): 0.59; PL: 0.29 – 0.32; PWmax: 0.46 – 0.55; PWmin: 0.41 – 0.44; ESL: 0.51 – 0.56; EW: 0.60 – 0.66; MTbL (holotype): 0.32; MTrL (holotype): 0.16 (MTrL 1 – 4: 0.06; MTrL 5: 0.10); AW: 0.61 – 0.66; AedL: 0.37 – 0.40; BL: 1.78 – 2.10 (holotype: 1.80). Habitus as in Fig. 1. Body reddish-brown, with slightly darker head and pronotum and dark-brown posterior portions of elytra (holotype and three paratypes) and abdomen (middle portions of each elytron in three paratypes yellow-brown); antennomeres 6 – 11 brown; mouthparts, antennomeres 1 – 5, laterobasal and basal margins of pronotum, and legs yellow. Head and neck without visible punctures; pronotum with very sparse and fine punctation more visible in latero-apical portions and finer in middle; scutellum without visible punctures; elytra with moderately dense, large and deep punctation, markedly finer and sparser in parascutellar portion and along suture; abdominal tergites with very fine and dense punctation, indistinct on abdominal tergites VII – VIII. Head with regular and strong microsculpture, transverse on vertex and isodiametric in middle portion; pronotum with dense and strong microreticulation, slightly transverse in medioapical and isodiametric in middle and lateral portions; scutellum with fine transverse meshes; microsculpture of elytra very dense, distinctly larger than that on pronotum (two paratypes with coarser sculpture along suture); abdomen with dense and moderately strong isodiametric sculpture, finer in abdominal tergites VII – VIII (three paratypes. Head indistinctly elevated in middle and posterior portions between ocelli and eyes, with wide and moderately deep postantennal depressions, 1.7 times as broad as long, with elongate and deep anteocellar foveae (grooves in front of ocelli) almost reaching level of middle of eyes; postocular carina missing. Apical maxillary palpomere very narrow, 1.7 – 1.9 times as long as preapical segment. Antenna reaching basal margins of elytra when reclined, with distinctly transverse antennomeres 6 – 10, progressively broadened apicad; basal antennomere about twice as long as broad, antennomere 2 distinctly narrower than basal antennomere, 3 slightly narrower and about as long as 2, 4 suboval, slightly longer than broad, 5 about as long as broad, 6 twice broader and about as long as 5, 7 broader and slightly longer than 6, 8 – 9 slightly broader than 7, 10 slightly longer than 9, apical antennomere about twice as long as preceding segment, from middle gradually narrowed toward rounded apex. Pronotum slightly convex, 1.5 – 1.7 times as broad as long, 1.2 – 1.3 times as broad as head, widest in middle, gradually and slightly narrowed anteriad toward widely rounded anterior angles not protruded apicad and strongly narrowed posteriad toward subtriangular, obtuse or subacute posterior angles; laterobasal margins strongly concaved; anterior margin straight or slightly concave in middle, slightly shorter than rounded posterior margin; middle portion with two wide paramedian impressions, from medioapical third gradually broadened posteriad; lateral margins distinctly impressed in some paratypes; laterobasal impressions very wide and deep. Elytra slightly broader than long, 1.7 times as long as pronotum, slightly broadened apicad; posterior margin straight or slightly rounded; surface of each elytron along suture narrowly impressed in three paratypes. Metatarsi twice shorter than metatibia. Abdomen about as broad as elytra, with two small and round tomentose spots in middle of abdominal tergite IV; apical margin of abdominal tergite VII with narrow palisade fringe. Male. Posterior margin of abdominal tergite VIII truncate (Fig. 9). Posterior margin of abdominal sternite VIII widely concave (Fig. 10). Aedeagus with wide basal bulb, gradually narrowed toward middle part of median lobe, and from about middle strongly narrowed toward small rounded apex; parameres moderately wide, with narrow apices, slightly exceeding apex of median lobe, with three long apical and one preapical setae and a row of 8 – 10 of moderately long setae on inner margins of each paramere; internal sac narrow and long, with several sclerotized elongate strunctures in basal portion (Fig. 7). Lateral aspect of the aedeagus as in Fig. 8. Female unknown. Comparative notes. Based on the shape of the lateral portions of the pronotum widely and deeply concave in front of posterior margin, P. conjuncta sp. n. is similar to P. flaveola sp. n., from which it can be distinguished by the darker coloration of the body, the presence of distinct wide and long paramedian impressions on the disc of pronotum, and narrower apical portions of the median lobe and the parameres.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA5947864A8FCCFFEA9FD52.taxon	distribution	Distribution. Paraphloeostiba conjuncta sp. n. is known only from the type locality in north-eastern part of Papua New Guinea.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA5947864A8FCCFFEA9FD52.taxon	etymology	Etymology. The specific epithet is the Latin adjective conjunctus, - a, - um (connected, related). It is related to P. flaveola sp. n.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA7947E64A8FCEBFF12FD6E.taxon	description	(Figs. 2, 11 – 14)	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA7947E64A8FCEBFF12FD6E.taxon	materials_examined	Material examined. PAPUA NEW GUINEA: 1 ♁ (Fig. 2; abdominal tergite VII strongly damaged; specimen was dissected prior to the present study, aedeagus and apical segments is placed in vial with glycerine pinned under the card with the specimen): ‘ Neth. Ind. - American New Guinea Exped. Sigi Camp 1500 m 26. ii. 1939 L. J. Toxopeus`, ‘ M. Cameron. Bequest. B. M. 1955 - 147.`, ‘ M. Cameron det., 19 Phloeonomus coriaceus COTYPE Cam.`, ‘ NHMUK 014663270`, ‘ Paraphloeostiba | coriacea (Cameron, 1952) | Shavrin A. V. det. 2023`(BMNH).	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA7947E64A8FCEBFF12FD6E.taxon	description	Redescription. Measurements: HW: 0.46; HL: 0.35; OL: 0.17; TL: 0.06; AL: 0.85; PL: 0.40; PWmax: 0.67; PWmin: 0.65; ESL: 0.74; EW: 0.90; MTbL: 0.34; MTrL: 0.16 (MTrL 1 – 4: 0.05; MTrL 5: 0.11); AW: 0.86; AedL: 0.50; BL: 2.56. Habitus as in Fig. 2. Body brown, with paler lateroapical portions of head, lateral and basal portions of pronotum and middle portion of elytra (abdomen slightly darker, with paler paratergites); antennomeres 6 – 11 yellow-brown; mouthparts, antennomeres 1 – 5 and legs yellowish. Head (including neck), pronotum and scutellum without punctures; elytra with moderately fine and dense punctation, finer along suture and indistinct in apical portions of each elytron; abdomen with very indistinct and moderately dense punctation, indistinct in middle portions of each tergite. Head with regular and very strong microsculpture; neck with moderately fine and dense isodiametric meshes; punctation of pronotum stronger and denser than that in middle portion of head; scutellum with dense and coarse sculpture, transverse in basal and somewhat isodiametric in apical portions; microsculpture of elytra isodiametric and very dense, but finer than that on pronotum; abdomen with very dense microsculpture, isodiametric in abdominal tergites IV – V and transverse in VI – VIII. Head flattened in middle, with slightly elevated infraorbital portions, 1.3 times as broad as long, with moderately wide and deep anteocellar foveae. Apical maxillary palpomere about twice as long as preapical segment. Antennomeres 8 – 10 slightly transverse; basal antennomere slightly more than twice as long as broad, antennomere 2 slightly narrower than basal antennomere, 4 distinctly longer than broad, 6 slightly transverse and distinctly broader than 5, 7 slightly broader and longer than 6, 8 – 9 slightly shorter than 7, 10 slightly longer than 9, apical antennomere about 1.2 times as long as 10. Pronotum distinctly convex, 1.6 times as broad as long, 1.4 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; posterior angles subacute; anterior margin distinctly concave in middle; middle portion with slightly impressed medioapical portion and with two indistinct and wide paramedian impressions; laterobasal portions widely and deeply impressed. Elytra convex, 1.8 times as long as pronotum, distinctly broadened apicad; posterior margin straight. Abdomen slightly narrower than elytra, with two very small and round tomentose spots in middle of abdominal tergite IV. Male. Posterior margin of abdominal tergite VIII truncate (Fig. 13). Posterior margin of abdominal sternite VIII slightly concave (Fig. 14). Aedeagus with wide basal bulb, strongly narrowed toward moderately small truncate apex; parameres distinctly exceeding apex of median lobe, gradually widened apically, with two long and two moderately short apical setae, and with with a row of 7 – 8 of moderately short setae on inner margins and one short seta on outer margins of each paramere; internal sac narrow and moderately short, with several sclerotized elongate stuctures in basal portion (Fig. 11). Lateral aspect of the aedeagus as in Fig. 12. Female unknown. Comparative notes. Paraphloeostiba coriacea can be distinguished from all species of the genus by the slightly transverse antennomere 6. Based on the general shape of the apical portion of the median lobe and the apical portions of parameres distinctly broadened apicad, and the presence of several short setae on inner margins of each paramere, it is somewhat similar to P. moluccensis Shavrin, 2022, known from Yamdena Island, Indonesia (Shavrin 2022), but differs from it by the darker coloration, stronger microsculpture of the head and the pronotum, significantly larger body, broader pronotum, longer elytra, slightly longer parameres and different morphology of the internal sac.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA7947E64A8FCEBFF12FD6E.taxon	distribution	Distribution. Paraphloeostiba coriacea is known from three localities in eastern part of Papua New Guinea.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA7947E64A8FCEBFF12FD6E.taxon	discussion	Remarks. Phloeonomus (s. str.) coriaceus was originally described from “ Rattan Camp, 1150 m ” (type) and “ Top Camp, 2100 m ... ” based on the material collected by L. J. Toxopeus. Steel (1960) studied “ one paratype ... from the Cameron collection ” and the male above (not type!), redescribed it and transferred to the genus Paraphloeostiba. I re-studied the latter specimen again, made figures of the aedeagus and genital segments, and redescribed the species.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA1947C64A8FCB7FE77FD9E.taxon	description	(Figs. 3, 15 – 21)	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA1947C64A8FCB7FE77FD9E.taxon	materials_examined	Type material. Holotype ♁ (Fig. 3; dissected): ‘ 12 VIII 79 [handwritten in black] | PNG / EHProv. | Umg. Kainantu | Onerunka <printed>, Papua | Nlle Guinée | W. G. Ullrich` <printed>, ‘ HOLOTYPE | Paraphloeostiba | flaveola sp. n. | Shavrin A. V. des. 2023`<red, printed> (MHNG). Paratypes: 3 ♀♀ (one specimen dissected): ‘ PAPUA N. GUINEA | Onerunka VIII. 79 [handwritten in black] | nr Kainantu | W. G. Ullrich` <printed>, ‘ PARATYPE | Paraphloeostiba | flaveola sp. n. | Shavrin A. V. des. 2023`<red, printed> (1 ♀: cSh, 2 ♀♀: MHNG).	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA1947C64A8FCB7FE77FD9E.taxon	description	Description. Measurements (n = 4): HW: 0.37 – 0.41; HL: 0.20 – 0.23; OL: 0.11 – 0.12; TL: 0.01 – 0.02; AL (holotype): 0.57; PL: 0.28 – 0.33; PWmax: 0.55 – 0.57; PWmin: 0.47 – 0.52; ESL: 0.50 – 0.56; EW: 0.63 – 0.68; MTbL (holotype): 0.30; MTrL (holotype): 0.16 (MTrL 1 – 4: 0.06; MTrL 5: 0.10); AW: 0.62 – 0.66; AedL: 0.45; BL: 2.00 – 2.35 (holotype: 2.18). Habitus as in Fig. 3. Body yellow-brown, with slightly darker apical portion of head, latero-apical and apical portions of elytra, and sometimes apical margins of abdominal tergites; antennomeres 6 – 11 brown; mouthparts, antennomeres 1 – 5 and legs yellow. Pronotum with indistinct and very sparse fine punctation, larger in lateral and indistinct or missing in middle portions; punctation of latero-apical and lateral portions of elytra slightly larger and deeper than that in pronotum; abdominal tergites without or with very fine and sparse punctation more distinct in lateral portion of each tergite. Head with coarse and dense microsculpture; neck with dense and fine transverse meshes; pronotum with very dense isodiametric sculpture, slightly finer than that in middle portion of head; microsculpture of elytra isodiametric, slightly coarser and larger than that on pronotum; abdomen with dense and transverse microreticulation. Head 1.7 – 1.8 times as broad as long, with short (about as long as diameter of ocellus) and deep anteocellar foveae; postocular carina present, indistinct and subacute, distance between posterior margin of eye and carina about as long as one nearest ommatidium. Apical maxillary palpomere narrow, slightly more than twice as long as preapical segment. Antennomere 7 slightly and antennomeres 8 – 10 strongly transverse; antennomere 2 slightly narrower than basal antennomere, 3 distinctly narrower than 2, 4 small, about twice shorter than 3, 5 slightly broader and shorter than 4, 6 about twice broader and slightly longer than 5, 7 – 8 slightly broader than 6, 9 – 10 slightly longer than 8, apical antennomere twice longer than 10. Pronotum distinctly transverse, 1.7 – 1.8 times as broad as long, 1.3 – 1.4 times as broad as head, widest in about middle; laterobasal strongly concaved in front of obtuse or subacute posterior angles; anterior margin widely concave in middle; middle portion without visible paramedian impressions except of two paratypes with very indistinct long and wide impressions. Elytra 1.2 times as broad as long, 1.6 – 1.7 times as long as pronotum; posterior margin straight. Abdomen slightly narrower than elytra,. Male. Posterior margin of abdominal tergite VIII truncate (Fig. 17). Posterior margin of abdominal sternite VIII widely and deeply concave (Fig. 18). Aedeagus with wide basal bulb, gradually narrowed toward moderately narrow and rounded apical portion; parameres significantly narrowed apically toward widely rounded apical lobes, distinctly exceeding apex of median lobe, with three moderately long apical setae, with a row of 4 – 6 of moderately short setae on inner margins and 2 – 3 small setae on outer margins of eache paramere; internal sac moderately long and wide, with several sclerotized and very long structures in basal portion (Fig. 15). Lateral aspect of the aedeagus as in Fig. 16. Female. Posterior margin of abdominal tergite VIII straight (Fig. 19). Posterior margin of abdominal sternite VIII widely rounded (Fig. 20). Female accessory sclerite as in Fig. 21. Spermatheca not recognized. Comparative notes. Based on the shape of the lateral portions of the pronotum widely and deeply concave in front of posterior margin, P. flaveola sp. n. is similar to P. conjuncta sp. n., from which it can be distinguished by the paler coloration of the body, the missing of paramedian impressions on the pronotum, and broader apical portions of the median lobe and the parameres.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA1947C64A8FCB7FE77FD9E.taxon	distribution	Distribution. Paraphloeostiba flaveola sp. n. is known only from the type locality in north-eastern part of Papua New Guinea.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA1947C64A8FCB7FE77FD9E.taxon	etymology	Etymology. The specific epithet is the Latin adjective flaveolus, - a, - um (yellowish). It refers to the general coloration of the body.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA3947C64A8FD27FD65FBED.taxon	description	(Fig. 4)	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA3947C64A8FD27FD65FBED.taxon	materials_examined	Material examined. PAPUA NEW GUINEA: 1 ♁, 1 ♀: Onerunka, near Kainantu. 08.05.1979. W. G. Ullrich leg. (MHNG); 1 ♀: same data. 01.06.1979 (MHNG); 2 ♁♁, 3 ♀♀: same data. 19.08.1979 (MHNG); 22 ♁♁, 52 ♀♀: same data. 08.1979 (MHNG, cSh); 2 ♁♁, 2 ♀♀: same data. 21.09.1979 (MHNG); 5 ♁♁, 13 ♀♀: same data. 09.1979 (MHNG); 8 ♁♁, 9 ♀♀: same data. 11.1979 (MHNG); 1 ♀: Rintebe, 30 km E Goroka. 1700 m a. s. l. 05.1979. W. G. Ullrich leg. (MHNG).	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA3947C64A8FD27FD65FBED.taxon	discussion	Remarks. Habitus as in Fig. 4. Jenkins Shaw et al. (2023) recorded it from Papua New Guinea without exact localities based on the material listed above.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA3947364A8FB36FA85FBB2.taxon	description	(Figs. 5, 22 – 25)	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA3947364A8FB36FA85FBB2.taxon	materials_examined	Type material. Holotype of Phloeonomus papuanus Cameron, 1931 ♀ (Fig. 5): ‘ Type` <round printed label with red margin>, ‘ Mt. Lamington | N. E. Papua | 1300 to 1500 feet | C. T. McNamara` <printed>, ‘ Phloeonomus | papuanus | TYPE [red] Cam.`<handwritten in black>, ‘ NHMUK 014663272 <printed label with barcode on the right side>, ‘ Paraphloeostiba | papuana (Cameron, 1952) | Shavrin A. V. det. 2023`<printed> (BMNH). Additional material examined. PAPUA NEW GUINEA: 1 ♀: ‘ Finschhafen, New Guinea 1.4 / [19] 44 Helfer` (BMNH).	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA3947364A8FB36FA85FBB2.taxon	description	Redescription. Measurements (n = 2): HW: 0.38 – 0.42; HL: 0.20 – 0.22; OL: 0.15; TL: 0.02; AL (holotype): 0.46; PL: 0.28 – 0.30; PWmax: 0.52 – 0.55; PWmin: 0.42 – 0.44; ESL: 0.47 – 0.52; EW: 0.60 – 0.62; MTbL (holotype): 0.29; MTrL (holotype): 0.15 (MTrL 1 – 4: 0.05; MTrL 5: 0.10); AW: 0.65 – 0.67; BL: 1.80 – 1.85 (holotype). Habitus as in Fig. 5. Body yellow-brown, with darker lateral portions of head and abdomen in holotype; mouthparts, antennae and legs yellow (antenomeres 7 – 11 of specimen from Finschhafen slightly darker). Head with indistinct and very fine punctation, denser in middle; neck without punctures; pronotum with very dense and fine punctation, without distinct punctures in middle; scutellum without punctures; punctation of elytra dense, slightly larger and deeper than that on pronotum, finer along suture and in apical portions of each elytron; abdomen without visible punctures on abdominal tergites IV – VI, with very indistinct, fine and sparse punctation on tergites VII – VIII. Head with coarse microscupture; neck with distinct transverse meshes; pronotum with very dense microreticulation, slightly coarser than that in middle portion of head; microsculpture of elytra dense, slightly more transverse than that on pronotum; abdomen with dense sculpture, slightly transverse on abdominal tergites IV – VI and transverse on tergites VII – VIII. Head with wide and moderately shallow postantennal depression, 1.9 times as broad as long, with very short (about as long as diameter of ocellus) and deep anteocellar foveae; postocular carina indistinct, acute, distance between posterior margin of eye and carina about as long as one nearest ommatidium. Apical maxillary palpomere sligthly more than twice as long as preapical segment. Antennomere 6 slightly and antennomeres 7 – 10 strongly transverse; antennomere 2 slightly narrower than basal antennomere, 3 slightly longer than 2, 4 slightly shorter than 4, 6 distinctly broader than 5, 7 – 9 significantly broader than 6, 10 slightly longer than 9, apical antennomere about twice as long as 10. Pronotum 1.8 times as broad as long, 1.3 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; posterior angles obtuse; anterior margin slightly concave in middle; middle portion without paramedian impressions; laterobasal impressions wide and shallow. Elytra slightly convex, 1.1 – 1.2 times as broad as long, 1.6 – 1.7 times as long as pronotum slightly more narrowed posteriad than annteriad. Abdomen slightly broader than elytra. Male unknown. Female. Posterior margin of abdominal tergite VIII truncate (Fig. 22). Posterior margin of abdominal sternite VIII rounded (Fig. 23). Female accessory sclerite as in Fig. 24. Spermatheca as in Fig. 25. Comparative notes. Based on the slightly convex elytra, the shape of the pronotum P. papuana is somewhat similar to P. obesa (see below), from which it can be distinguished by the smaller and slightly paler body, finer punctation of the pronotum, and different shape of the female accessory sclerite.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA3947364A8FB36FA85FBB2.taxon	distribution	Distribution. Paraphloeostiba papuana is known from two localities in eastern part of Papua New Guinea.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA3947364A8FB36FA85FBB2.taxon	discussion	Remarks. Phloeonomus papuanus was originally described based on the female from “ Mount Lamington ”, collected by C. T. McNamara. Steel (1960) redescribed it and transferred to the genus Paraphloeostiba. I re-studied the female which was previously studied by Steel. The spermatheca of P. papuana is figured for the first time.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAC947164A8FBCBFE76F990.taxon	description	(Figs. 6, 26 – 32)	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAC947164A8FBCBFE76F990.taxon	materials_examined	Type material. Holotype ♁: ‘ 12 VIII 79 [handwritten in black] | PNG / EHProv. | Umg. Kainantu | Onerunka` <printed>, ‘ Papua | Nlle Guinée | W. G. Ullrich <printed>, ‘ HOLOTYPE | Paraphloeostiba | ullrichi sp. n. | Shavrin A. V. des. 2023`<red, printed> (MHNG). Paratypes: 3 ♁♁ (two specimens dissected), 4 ♀♀ (one specimen dissected): same data as the holotype (2 ♁♁: cSh, 1 ♁, 4 ♀♀: MHNG); 1 ♁ (head damaged, aedeagus missing), 7 ♀♀ (one specimen without head): ‘ PAPUA N. GUINEA | Onerunka VIII. 79 [handwritten in black] | nr Kainantu | W. G. Ullrich` <printed> (1 ♁, 2 ♀♀: cSh; 5 ♀♀: MHNG). All paratypes with additional red printed label: ‘ PARATYPE | Paraphloeostiba | ullrichi sp. n. | Shavrin A. V. des. 2023`.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAC947164A8FBCBFE76F990.taxon	description	Description. Measurements (n = 16): HW: 0.34 – 0.39; HL: 0.18 – 0.24; OL: 0.12 – 0.13; TL: 0.02 – 0.00; AL (holotype): 0.55; PL: 0.28 – 0.34; PWmax: 0.52 – 0.57; PWmin: 0.41 – 0.46; ESL: 0.49 – 0.54; EW: 0.62 – 0.66; MTbL (holotype): 0.30; MTrL (holotype): 0.13 (MTrL 1 – 4: 0.05; MTrL 5: 0.08); AW: 0.59 – 0.64; AedL: 0.30 – 0.32; BL: 1.40 – 2.34 (holotype: 1.75). Habitus as in Fig. 6. Body yellow-brown (one paratype) to reddish-brown; mouthparts, antennae and legs yellow (antennomeres 7 – 11 sometimes slightly darker). Head with very sparse and fine punctation, middle portion of head usually without punctures; neck and scutellum without punctures; pronotum with sparse, moderately large and deep pucntation, sparser in middle portion; punctation of elytra sparse, distinctly larger and deeper than that on pronotum, sparser and finer in middle portions of each elytron; abdomen with fine and sparse punctation, sparser and finer in middle portion of each tergites. Head with very fine microsculpture, somewhat isodiametric in middle and sometimes indistinct in basal portions; neck with very fine transverse microreticulation; pronotum with fine transverse sculpture, indistinct in mediobasal portion in some paratypes; scutellum with very fine transverse meshes; elytra without microsculpture except basal portion with fine transverse microreticulation in some paratypes; abdomen with fine transverse meshes. Head slightly elevated in middle portion, 1.6 – 1.8 times as broad as long, with short and deep anteocellar foveae, slightly longer than diameter of ocellus, reaching basal third of level of eyes. Apical maxillary palpomere narrow and short, about 1.3 times as long as preapical segment. Antennomeres 6 – 10 strongly transverse; antennomere 2 about as broad as basal antennomere, 3 distinctly narrower and shorter than 2, 4 shorter than 3, 5 slightly broader than 4, 6 about twice broader than 5, 7 distinctly broader and longer than 6, 8 – 10 slightly shorter than 7, apical antennomere about three times as long as 10, from middle gradually narrowed toward subacute apex. Pronotum 1.6 – 1.8 times as broad as long, 1.4 – 1.5 times as broad as head, widest in middle, more narrowed posteriad than anteriad; posterior angles obtuse; anterior margin distinctly concave in middle; middle portion without paramedian impressions; laterobasal portions without or with with shallow and moderately wide impressions. Elytra 1.2 times as broad as long, 1.5 – 1.7 as long as pronotum, distinctly broadened apicad; posterior margin slightly rounded. Abdomen slightly narrower than elytra. Male. Posterior margin of abdominal tergite VIII slightly concave (Fig. 28). Posterior margin of abdominal sternite VIII widely and deeply concave (Fig. 29). Aedeagus with small basal bulb, gradually narrowed toward narrow and long median lobe with rounded apex; parameres short, not reaching apex of median lobe, with two moderately long apical and three preapical setae; internal sac moderately narrow and short, with sclerotized narrow struncture in basal portion (Fig. 26). Lateral aspect of the aedeagus as in Fig. 27. Female. Posterior margin of abdominal tergite VIII truncate (Fig. 30). Posterior margin of abdominal sternite VIII widely rounded (Fig. 31). Female genital segment as in Fig. 32. Female acessory sclerite and spermatheca not recognized. Comparative notes. Based on the glossy body with fine sculpture of the forebody and the general shape of the aedeagus, P. ullrichi sp. n. is similar to P. solomonensis Steel, 1960, known from “ Solomon Islands: Guadalcanal, Kukum ... ” (Steel 1960). It can be distinguished from it by the slightly broader apical portion of the median lobe and shorter parameres.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAC947164A8FBCBFE76F990.taxon	distribution	Distribution. Paraphloeostiba ullrichi sp. n. is known from two localities in eastern part of Papua New Guinea.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAC947164A8FBCBFE76F990.taxon	etymology	Etymology. Patronymic, the species is named to honour W. G. Ullrich (Lübeck), collector of the type specimens in Papua New Guinea.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAF947064A8FDBCFB3FFC12.taxon	description	(Figs. 33, 37 – 38)	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAF947064A8FDBCFB3FFC12.taxon	materials_examined	Material examined. INDIA: ARUNACHAL PRADESH: 1 ♀: Dirang vicinity, 27 ° 21 ’ N 92 ° 13 ’ E. 1800 ± 100 m a. s. l .. L. Dembický leg. (BMNH); PHILIPPINES: 1 ♁, 1 ♀: Mindanao, 30 km E of Malaybalay, Busdi. 1000 m a. s. l. 05 - 09.05.1996. Bolm leg. (SMNS); 1 ♀: same island, 30 km NW of Maramag, Bagongsilang. 13 - 17.05.1996. 1700 m a. s. l. Bolm leg. (SMNS).	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAF947064A8FDBCFB3FFC12.taxon	discussion	Remarks. Paraphloeostiba apicalis was known from Indonesia and Philippines (Herman 2001, Shavrin 2020). It is here recorded from India for the first time. Habitus as in Fig. 33. Aedeagus as in Figs. 37 – 38.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAF947064A8FBABFF58F9E1.taxon	description	(Fig. 34)	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAF947064A8FBABFF58F9E1.taxon	materials_examined	Material examined. MALAYSIA: 1 ♁: Borneo, Sabah, Kinabalu National Park: Poring. 500 m a. s. l. 29.11 - 02.12.1996. W. Schawaller leg. (SMNS).	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAF947064A8FBABFF58F9E1.taxon	discussion	Remarks. Homalium obesum was originallly described from “ Sumatra: région de Deli, dans des tabacs importés ”. Steel (1960) redescribed it, designated the lectotype, provided figures of the aedeagus and the female accessory sclerite, and recorded it from Java, Sarawak and Luzon. Phloeonomus densepunctatus Bernhauer, 1915 was originally described from “ Sarawak: Mt. Matang ”. Steel (1960) synonymized it with P. obesa. Paraphloeostiba obesa is known from Indonesia, Malaysia (including Sabah) and Philippines (Herman 2001). Habitus as in Fig. 34.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAF947664A8F93AFCEAFF76.taxon	description	(Figs. 35, 39 – 40)	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAF947664A8F93AFCEAFF76.taxon	materials_examined	Material examined. INDIA: 1 ♁, 2 ♀♀: UTTARAKHAND: Nainital District, Mukteshwar. 2100 – 2150 m a. s. l. 25.02.2004. S. Saluk leg. (cSh); TAMIL NADU: 6 ♁♁, 10 ♀♀: Kodaikanal, Madurai. 1400 m a. s. l. 28.08.1989. A. Riedel leg. (cSh, SMNS).	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFAF947664A8F93AFCEAFF76.taxon	discussion	Remarks. Phloeonomus subopacus was originally described from “ Chakrata District; Simla Hills ... ”. Steel (1960) redescribed it and transferred to the genus Paraphloeostiba. The species was known from the Himalayan regions of India (Himachal Pradesh, Uttarakhand). It is here recorded from southern India (Tamil Nadu) for the first time. Habitus as in Fig. 35. Aedeagus as in Figs. 39 – 40.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA9947664A8FE8FFCEAFD61.taxon	description	(Figs. 36, 41 – 42)	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA9947664A8FE8FFCEAFD61.taxon	materials_examined	Material examined. PHILIPPINES: 2 ♁♁: Leyte, Visca N Baybay. 22.02.1991. 200 – 500 m a. s. l., primary forest. W. Shawaller et al. leg. (SMNS).	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
002087EAFFA9947664A8FE8FFCEAFD61.taxon	discussion	Remarks. Phloeonomus tonkinensis was origianlly described from “ Tonkin: Hoa Binh ”. Steel (1960) redescribed it as Paraphloeostiba, and provided figures of the aedeagus, the female genital segment and the spermatheca. The species was known from Vietnam and Singapore (Herman 2001). It is here recorded from Philippines for the first time. Habitus as in Fig. 36. Aedeagus as in Figs. 41 – 42.	en	Shavrin, Alexey V. (2023): New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa 5301 (4): 486-500, DOI: 10.11646/zootaxa.5301.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5301.4.6
