taxonID	type	description	language	source
DC833AECC9B25AB5B79FC1F231D614DF.taxon	description	Figs 1 A-D, 11, 17, 25 A-E, 34 A-C, 40	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
DC833AECC9B25AB5B79FC1F231D614DF.taxon	diagnosis	Diagnosis. Dorsum black. PW / PL = 1.18 - 1.21; PAW / PBW = 0.99 - 1.00 (Fig. 17); pronotum with anterior angles rounded, moderately projected forward; disc sparsely punctate and pubescent with deep transverse rugosities, but with a glabrous area in the middle. Elytral intervals flat, densely punctate and pubescent. Hind wings reduced. Metepisterna short, MW / ML = 1.1 - 1.3 (Fig. 11). Apex of apical lamella rounded (Fig. 25 A-E).	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
DC833AECC9B25AB5B79FC1F231D614DF.taxon	description	Description. BL = 20.3 - 22.0 mm, BW = 7.6 - 8.9 mm [BL = 21.0 mm, BW = 7.0 mm in holotype], PAW = 3.6 - 3.9 mm, PBW = 3.6 - 3.9 mm, PW = 5.0 - 5.5 mm, PL = 4.3 - 4.5 mm, MW = 1.9 - 2.0 mm, ML = 1.5 - 1.8 mm. Head, pronotum, elytra, legs, and venter black; antennae, labial and maxillary palpi, apex of mouthparts, and tarsomeres dark brown. Head. Vertex punctate and pubescent with a glabrous and rugose area in the middle; antennae long, reaching middle of elytra; antennomere 3 ~ 1.5 x longer than antennomere 4. Pronotum cordiform, PW / PL = 1.18 - 1.21 (Fig. 17), widest at apical third; anterior margin slightly concave, its width equal to its basal width, PAW / PBW = 0.99 - 1.00; lateral margins rounded before middle, then distinctly narrowed to base, faintly sinuate before posterior angles; anterior angles rounded, moderately projected forward; posterior angles obtuse, slightly sharp at tips; disc gently convex, sparsely punctate and pubescent, with deep transverse rugosities, with a small glabrous area in the middle; median line distinct, not reaching anterior margin and base; basal foveae deep and long, punctate, pubescent. Elytra elongate, EL / BW = 1.45 - 1.61; gently convex, widest near posterior third, rounded at apex in males, subtruncate in females; striae with fine punctures; parascutellar striae well developed; parascutellar pores present; intervals flat, densely punctate and pubescent; sutural angles sharp; hind wings reduced. Venter densely punctate, pubescent, metepisterna (Fig. 11) short, MW / ML = 1.25 - 1.33; abdominal sternites III-VI with one setiferous puncture each side, sternite VII with one pair of setiferous punctures in males, two pairs in females; all abdominal sternites with distinct impressions laterally. Legs long and slender; tarsi nearly smooth dorsally. Male genitalia. Median lobe (Fig. 25 B-E) long, strongly bent to ventral side; apical orifice opened dorsally, long and wide, not reaching basal bulb, slightly turned to left side; in dorsal view, apical lamella (Fig. 25 A) slightly bent to left side, length nearly equal to its basal width, apex rounded; in left lateral view, apical portion distinctly bent into a hook ventrally (Fig. 25 B, E), basal orifice ~ 90 ° relative to preapical shaft; left paramere large and round; right paramere helical carved; endophallus with coiled flagellum; basal part of flagellum strongly bent with an oval sclerite facing the right; apical part of flagellum distinctly bent with a triangular bursa. Female genitalia. Spermatheca (Fig. 34 A-C) with seminal canal ~ 15 x as long as receptaculum; receptaculum short linear; seminal canal inserted at base of common oviduct; spermathecal gland rounded and inserted near apex of seminal canal; villous canal long, tortuously contorted, adhered to common oviduct.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
DC833AECC9B25AB5B79FC1F231D614DF.taxon	distribution	Distribution. (Fig. 40) This species is known only from Guidong, Hunan. Its distribution seems isolated by a mountain barrier from C. hunanus.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
DC833AECC9B25AB5B79FC1F231D614DF.taxon	etymology	Etymology. The new species caperatus is named for its rugosity on the vertex, pronotal disc, and abdominal sternites.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
C1F49D6A1F585AFB9C71D7D2124DAB99.taxon	diagnosis	Diagnosis. Sphodromimus can be distinguished from other subgenera in Chlaenius by the following character combinations: body large, BL 19 - 26 mm; body black or metallic colored, luster matt or strongly shiny, antennae, mandibles usually dark brown, elytra black, ventral side black; head finely punctate; penultimate labial palpomere with 5 - 7 setae, apex truncate; antennae long, antennomere 3 longer than 4; mentum tooth stout, bifid; pronotum long, with sides usually sinuate before posterior angles, posterior lateral seta situated before posterior angles, anterior lateral seta absent; proepisterna sparsely punctate and pubescent; elytral intervals flat or convex, not ribbed, densely punctate and pubescent, basal margin reaching the scutellum; hind wings reduced in all species except Chlaenius flavofemoratus Laporte, 1834 and Chlaenius peterseni (Louwerens, 1967); prosternal process unbordered at apex; metepisterna wider than long in all species except C. flavofemoratus and C. peterseni, coarsely punctate, pubescent; legs sparsely pubescent, tarsi nearly smooth dorsally, claws simple, protibiae sulcate on dorsal side; abdominal sternites finely punctate laterally; apical lamella of aedeagus denticulate on dorsal side; apical gonocoxite without ensiform setae; receptaculum very short to absent.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
C1F49D6A1F585AFB9C71D7D2124DAB99.taxon	distribution	Species and distribution. Subgenus is composed of 16 species distributed in the Oriental Region (China, Indonesia, Laos, Myanmar, Nepal, Philippines, Thailand, Vietnam), including the two new species described below.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
B5DB751649F25FBFB9B9635FA0BEBE81.taxon	description	Figs 9 A, B, 15, 23, 31 A-E, 39 A-C, 40	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
B5DB751649F25FBFB9B9635FA0BEBE81.taxon	materials_examined	Material examined. Total 10 specimens. China - Guangdong: Holotype female (DWC, photo), China, Guangdong, 1500 m, Xinyi: Datianding (22.16 / 111.15) - VIII- 1997 - leg. Li / Holotypus, Chlaenius (Sphodromimus) wrasei sp. nov. det. Kirschenhofer, 2001 [red label] / COLL WRASE, BERLIN; 1 ♂ and 4 ♀♀ (IZAS), Guangdong, Xinyi, Yunkai Shan, 1508.21 m / 22.291317 ° N, 111.209888 ° E, 2017. V. 30, Y. Z. Liu, S. P. Yu leg., Inst. of Zoology; 2 ♀♀ (IZAS), Guangdong, Xinyi, Yunkai Shan, 1250.55 m / 22.292692 ° N, 111.203833 ° E, 2017. V. 31, Y. Z. Liu, S. P. Yu leg., Inst. of Zoology; Guangxi: 2 ♂♂ (IZAS), Guangxi Prov, Daming Shan, Tianping Protect Station, N 23.49811, E 108.43715 / 1230 m, 2011. V. 27 N, Xinlei Huang Coll., Inst. of Zoology.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
B5DB751649F25FBFB9B9635FA0BEBE81.taxon	diagnosis	Diagnosis. Pronotum fully metallic purple, or greenish purple. PW / PL = 1.09 - 1.11; PAW / PBW = 0.83 - 0.93 (Fig. 23); pronotum cordate with posterior angles nearly right angled, rounded at tips; disc sparsely punctate and pubescent, with dense, shallow, transverse rugosities, without glabrous area in the middle. Elytral intervals convex basally, flat apically, densely punctate and pubescent. Hind wings reduced. Metepisterna short, MW / ML = 1.11 - 1.15 (Fig. 15). Legs totally black.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
B5DB751649F25FBFB9B9635FA0BEBE81.taxon	description	Description. BL = 21.3 - 24.4 mm, BW = 8.0 - 8.7 mm. PL = 4.5 - 5.0 mm, PW = 5.0 - 5.5 mm, MW = 2.0 - 2.1 mm, ML = 1.7 - 1.9 mm. Head, elytra, venter, and legs dark and black; pronotum fully purple or greenish purple; antennae, labial and maxillary palpi, apex of mouthparts and tarsomeres dark brown. Head. Vertex finely punctate, pubescent, without a distinct glabrous area; antennae long, reaching middle of elytra; antennomere 3 ~ 1.5 x longer than antennomere 4. Pronotum cordiform, PW / PL = 1.09 - 1.11 (Fig. 23), widest at apical third; anterior margin slightly concave, PAW / PBW = 0.83 - 0.93; lateral margins distinctly narrowed from middle to base, distinctly sinuate before posterior angles; anterior angles rounded, slightly projected forward; posterior angles nearly right angled, rounded at tips; disc gently convex, sparsely punctate and pubescent, with dense shallow transverse rugosities, without glabrous area in the middle; median line distinct, fine, not reaching anterior margin and base; basal foveae deeply arcuate, punctate and pubescent. Elytra elongate, EL / BW = 1.61 - 1.75, gently convex near anterior third, widest near posterior third, rounded at apex in males, subtruncate in females; striae with deep punctures; parascutellar striae well developed; parascutellar pores present; intervals convex at base, flat from middle to apex, densely punctate and pubescent; sutural angles obtuse and right; hind wings reduced. Venter densely punctate, pubescent, metepisterna (Fig. 15) short, MW / ML = 1.11 - 1.15; abdominal sternites III-VI with one setiferous puncture each side, sternite VII with one pair of setiferous punctures in males, two pairs in females; all abdominal sternites with distinct impressions laterally. Legs long and slender; tarsi nearly smooth dorsally. Male genitalia. Median lobe (Fig. 31 B-E) long, strongly bent to ventral side; apical orifice opened dorsally, long and wide, not reaching basal bulb; in dorsal view, apical lamella triangular (Fig. 31 A), distinctly bent to right side, longer than basal width, each side of middle apical lamella with a denticulation or absent on right side; in left lateral view, apical portion slightly bent dorsally at apex, basal orifice ~ 90 ° relative to preapical shaft; left paramere larger than right paramere, both helically curved; endophallus with flagellum thick and straight; basal part of flagellum with irregular bursa; apical part of flagellum with drop-shaped sclerite. Female genitalia. Bursa copulatrix (Fig. 39 A-C) round, base with a distinct long digitiform protrusion; villous canal long, tortuously contorted, adhered to common oviduct; spermatheca and spermathecal gland absent.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
B5DB751649F25FBFB9B9635FA0BEBE81.taxon	distribution	Distribution. (Fig. 40) China (Guangdong; Guangxi).	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
33485B2928AD52489B49E1065845E20A.taxon	description	Figs 8, 10 A, B, 16, 24, 32 A-F, 33 A-E, 40	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
33485B2928AD52489B49E1065845E20A.taxon	materials_examined	Material examined. Total 5 specimens. Vietnam: Holotype, Male (NHMB), Indo Chine coll. Dussault / En-Le 1908 / Chl (Chlaenius Macrochlaeniles) Chlaenius Macrochlaeniles flavofemoratus ssp. enleensis Dr. K. MANDL det. 1978 / Holotype [red label]. Paratype (DWC, photo), 1 ♂ (IZAS), Vietnam, Tam Dao, 20 - 28. VI. 1990, Dr. Blazicek lgt. / Paratypus, Chlaenius (Haplochlaenius) tamdaoensis mihi det. Kirschenhofer 2001 [red label] / COLL WRASE, BERLIN; 1 ♂ (IZAS), Vietnam, Tam Dao, 60 km NW Hanoi, 900 m, 1997 May-June, S. Ryabov [internal sac fully everted]; 1 ♂ (IZAS), Vietnam, Cao Bang, Nguen Binh, 800 m, 2003. V. 13, S. Ryabov [internal sac partially everted]; 1 ♂ (IZAS), Tonkin, Hoa-Binh, leg. A. de Cooman [genitalia damaged by dermestes beetle].	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
33485B2928AD52489B49E1065845E20A.taxon	diagnosis	Diagnosis. Pronotum green to coppery. PW / PL = 1.06 - 1.12; PAW / PBW = 0.82 - 0.95 (Fig. 24); pronotum cordate with anterior angles rounded, not projected forward; disc sparsely punctate and pubescent, with shallow transverse rugosities. Elytral intervals gently convex throughout; densely punctate and pubescent. Hind wings reduced. Metepisterna short; MW / ML = 1.11 - 1.17 (Fig. 16). Apex of femora dark brown or yellow-brown, the rest of legs black.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
33485B2928AD52489B49E1065845E20A.taxon	description	Description. BL = 21.7 - 24.1 mm, BW = 8.2 - 8.7 mm. PL = 4.5 - 4.9 mm, PW = 5.0 - 5.3 mm, MW = 2.0 - 2.1 mm, ML = 1.7 - 1.8 mm. Head, elytra, venter dark and black; pronotum green to coppery; antennae, labial and maxillary palpi, apex of mouthparts and tarsomeres dark brown; apex of femora dark brown or yellow-brown, the rest of legs black. Head. Vertex finely punctate, pubescent, without a distinct glabrous area; antennae long, reaching middle of elytra; antennomere 3 ~ 1.5 x longer than antennomere 4. Pronotum cordiform, PW / PL = 1.06 - 1.12 (Fig. 24), widest at apical third; anterior margin slightly concave, PAW / PBW = 0.82 - 0.95; lateral margins distinctly narrowed from middle to base, slightly sinuate before posterior angles; anterior angles rounded, not projected forward; posterior angles nearly right angled, rounded at tips; disc gently convex, sparsely punctate and pubescent, with shallow, transverse rugosities, without glabrous area; median line distinct, fine, not reaching anterior margin and base; basal foveae deeply arcuate, punctate and pubescent. Elytra elongate, EL / BW = 1.58 - 2.22, gently convex near anterior third, widest near posterior third, rounded at apex in males; striae with deep punctures; parascutellar striae well developed; parascutellar pores present; intervals gently convex throughout, densely punctate and pubescent; sutural angles sharp at tips; hind wings reduced. Venter densely punctate, pubescent, metepisterna (Fig. 16) short, MW / ML = 1.11 - 1.17; abdominal sternites III-VI with one setiferous puncture each side, sternite VII with one pair of setiferous punctures in males; all abdominal sternites with distinct impressions laterally. Legs long and slender; tarsi nearly smooth dorsally. Male genitalia. Median lobe (Figs 32 B-E, 33 B-E) long, strongly bent to ventral side; apical orifice opened dorsally, long and wide, not reaching basal bulb; in dorsal view, apical lamella linear, longer than basal width (Figs 32 A, 33 A), distinctly bent to right side, with a denticulation laterally in the middle, left denticulation distinctly larger than the right one; in left lateral view, apical portion slightly bent dorsally at apex, basal orifice ~ 90 ° relative to preapical shaft; left paramere larger than right paramere, both helically curved; endophallus (Fig. 32 F) with flagellum helically thick and straight; basal part of flagellum with irregular bursa; apical part of flagellum with triangular sclerite. Female genitalia unknown.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
33485B2928AD52489B49E1065845E20A.taxon	distribution	Distribution. (Fig. 40) Vietnam (Indo Chine). We mark En-le, Yunnan on the map with a question mark '? '.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
FCC2DE3B0936587DA515BAE3CFA8A826.taxon	description	Figs 7 A-C, 14, 22, 30 A-E, 38 A-C, 40	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
FCC2DE3B0936587DA515BAE3CFA8A826.taxon	materials_examined	Material examined. Total 83 specimens. China - Fujian: 1 ♀ (IZAS), Fujian, Nanping, 1985. VI. 7; 1 ♀ (IZAS), Fujian, Nanjing, 1991. V. 15; Guangdong: 3 ♂♂ (IZAS), Guangdong, Haifeng, Jimingsi, 23.037605 ° N, 115.25343 ° E, 178.09 m / 2017. V. 22, Y. Z. Liu, S. P. Yu leg., Inst. of Zoology; Guangxi: 1 ♂ (IZAS), Guangxi, Longsheng, Liluo, 1985. IV Jun Li leg.; 1 ♂ (IZAS), Guangxi, Guilin, Longsheng, 2003. VII. 7 Jianxin Cui leg.; 1 ♀ (IZAS), Guangxi, Nanning, 1980. IV. 21, Rongquan Cai leg.; 1 ♂ (IZAS), Guangxi, Jingxi, 840 m, 1998. IV. 1 Chunsheng Wu leg.; Guizhou: 1 ♂ and 1 ♀ (IZAS), Guizhou, Luodian, 420 m, 1979. IV. 16, Qingqiang Li leg.; 1 ♀ (IZAS), Guizhou, Luodian, 1979. V. 1; Hainan: 1 ♂ (IZAS), Hainan, Baisha, Yinggeling, 2011. V. 1, 600 m, light trap, Wenxin Lin leg., Inst. of Zoology; Yunnan: 4 ♂♂ and 1 ♀ (IZAS), Yunnan, Jinghong, Virgin Forest Park, Peacock Villa, 22.0304 ° N, 100.8763 ° E, 682 m / 2021. VIII. 6 N, along road Pingzhou Zhu leg., Inst. of Zoology; 1 ♂ (IZAS), Yunnan, Xishuangbanna, Botanical Garden, Chao Wu leg.; 1 ♀ (IZAS), Yunnan, Hekou, Qiaotou, Bajiaotian village / 22.85348 ° N, 104.14211 ° E, 886 m, 2021. IV. 21 N, Y. Xu, Z. Q. Yan coll., Inst. of Zoology; 1 ♂ (CAS), China, Yunnan Province, Tengchong, Shangying, Longwenqiao, field, beach, 25 ° 01 ' 19.9 " N, 98 ° 40 ' 40.4 " E / 1290 m, 2003. X. 20 D., H. B. Liang, X. C. Shi Coll., Institute of Zool., CAS & California Acad. Sciences; 1 ♂ and 3 ♀♀ (IZAS), Yunnan, Xishuangbanna, Menglun, Botanical Garden, 549 m, 21 ° 56.035 ' N, 101 ° 15.154 ' E / 2005. V. 25, Guo Zheng leg, Inst. of Zoology, CAS; 4 ♂♂ and 6 ♀♀ (CAS), Yunnan, Xishuangbanna, Menglun, Botanical Garden, 558 m, 21 ° 55.035 ' N, 101 ° 16.500 ' E / 2007. V. 20, Guo Zheng leg, Inst. of Zoology, CAS; 3 ♂♂ and 8 ♀♀ (IZAS), Yunnan, Xishuangbanna, Menglun, Botanical Garden, 558 m, 21 ° 55.035 ' N, 101 ° 16.500 ' E / 2007. VII. 10, Guo Zheng leg, Inst. of Zoology, CAS; 1 ♂ and 4 ♀♀ (CAS), Yunnan, Xishuangbanna, Menglun, Botanical Garden, 572 m, 21 ° 54.646 ' N, 101 ° 16.257 ' E / 2007. I. 10, Guo Zheng leg, Inst. of Zoology, CAS; 2 ♀♀ (IZAS), Yunnan, Xishuangbanna, Menglun, Botanical Garden, 2009. XII. 1, Guo Tang leg, Inst. of Zoology; 2 ♀♀ (IZAS), China, Yunnan Prov., Nabanhe N. R. Guomenshan, alt. 1150 m, 2009. V. 6, Jiayao Hu, Ziwei Yin leg.; 1 ♂ (IZAS), Yunnan, Jinghong, Menghai, Nabanhe N. R. Guomenshan, Forest, 2009. VI. 26, 1114 m / 22.24644 ° N, 100.60610 ° E, pitfall, L. Z. Meng leg., Inst. of Zoology; 1 ♂ (IZAS), China, Yunnan, Hekou, Longpu, 240 m, 2011. IV. 13 D, 22.65404 ° N, 103.98193 ° E / Xinlei Huang leg., Inst. of Zoology, CAS; 1 ♂ (IZAS), Yunnan, Xishuangbanna, Menglun, Botanical Garden, 540 m, 21.92987 ° N, 101.24820 ° E, 2011. IV. 21 N / Hongbin Liang, Kaiqing Li leg., Institute of Zool., CAS; 1 ♂ (CAS), Yunnan, Xishuangbanna, Menglun, Botanical Garden, 540 m, 21.92987 ° N, 101.24820 ° E, 2011. IV. 22 / Yan Li leg., Institute of Zool., CAS; 1 ♂ (IZAS), Yunnan Prov. Menglun, Botanical Garden, vegetation, 21.91175 ° N, 101.28163 ° E / 650 m, 2009. XI. 15, Guo Tang Coll., Inst. of Zoology; 1 ♀ (IZAS), Yunnan, Xishuangbanna, Menglun, Botanical Garden, 2011. V. 04, 560 m / 21 ° 55 ' 39.66 " N, 101 ° 15 ' 18.09 " E, Jingxin Liu leg., Inst. of Zoology; 1 ♀ (IZAS), Yunnan, Xishuangbanna, 29 km NW, Jinghong, vic. Da Nuo You / 22 ° 12.41 ' N, 100 ° 38.29 ' E, 790 m, 2009. V. 16 leg. L. Meng, rice follow; 1 ♀ (IZAS), Yunnan, Xishuangbanna, 20 km NW, Jinghong, vic. Man Dian (NNNR) / 22 ° 07.80 ' N, 100 ° 40.05 ' E, 740 m, 2008. V. 13 leg. A. Weigei rubb. plant; 1 ♂ and 1 ♀ (CAS), Yunnan, Lushui, Pianma, Gangfang, Xuetang 26.12218 ° N, 98.57546 ° E / 1625 m, 2005. V. 16 N, D. Kavanaugh, D. Z. Dong leg., Inst. of Zoology, CAS; 1 ♂ (CAS), China, Yunnan, Mengla, Biodiversity Corridor, 660 m, 2011. IV. 25 D, 21.40482 ° N, 101.63035 ° E / Xinlei Huang leg., G 213 1999 km, Inst. of Zoology, CAS; 1 ♂ (IZAS), Yunnan, Yangbi, Pingpo, 1422 m, 25 ° 35 ' 33 " N, 100 ° 2 ' 56 " E / 2002. VI. 27, Min Wu leg.; 1 ♀ (CAS), China, Yunnan Province, Tengchong, Qushi Town, Xiaojiangqiao, riverside, 25 ° 14 ' 22.2 " N, 98 ° 37 ' 38.0 " E / 1445 m, 2003. X. 21, night, H. B. Liang, X. C. Shi Coll., Institute of Zool., CAS & California Acad. Science; 1 ♀ (CAS), Yunnan, Gaoligongshan, Nujiang Prefecture, 1500 m, 26 ° 07.3 ' N, 98 ° 34.5 ' E, 1998. X. 14, D. H. Kavanaugh leg.; 2 ♂♂ (IZAS), Yunnan, Xishuangbanna, Menglun Town NO. 55, 2009. V. 04, Hu Li leg.; 4 ♀♀ (IZAS), Yunnan, Xishuangbanna, Menglun, Botanical Garden, 2012. V. 16, L. Z. Meng leg., Inst. of Zoology; 1 ♂ (IZAS), Yunnan, Xishuangbanna, Menglun, 1982. IV. 26, Linyao Wang leg.; 1 ♂ (IZAS), Yunnan, 1980. VI. 5, Fen Liu leg.; 1 ♀ (IZAS), Yunnan, Gaoligong, Cikai Town, Pulahe joint of Nujiang, 27.74843 ° N, 98.66498 ° E / 1530 m 2004. X. 23, night, D. Kavanaugh, D. Z. Dong leg. Inst. of Zoology, CAS; 1 ♂ (IZAS), Yunnan, Lancang, Mafang By pitfall traps, 22.57925 ° N, 99.99849 ° E / 1723 m, 2004. VI. 16, W. B. Gu coll, Inst. of Zoology, CAS; 1 ♀ (CAS), Yunnan, Lushui, Liuku, Gaoligong Shan, 25 ° 51 ' 20 " N, 98 ° 50 ' 58 " E / 800 m, 2002. IX. 19, H. B. Liang, night, Sino-American Exped., Inst. of Zoology, CAS; 1 ♂ (CAS), China, Yunnan, Gaoligongshan, Nujiang Prefecture, Gangfang, Sancha Lukou / 26 07.3 ' N, 98 34.5 ' E, 1500 m, 1998. X. 12, D. H. Kavanaugh collector.; Myanmar: 1 ♂ (NHML), Carin Cheba, 900 - 1100 m, L. Fea / Fry Coll., 1905.100. / Chlaenius flavofemoratus Cast. = nigricoxis Mots., comp with type, H. E. Andrewes det.; Vietnam: 1 ♀ (IZAS), Tonkin, Hoa Binh, leg: A. de Cooman.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
FCC2DE3B0936587DA515BAE3CFA8A826.taxon	diagnosis	Diagnosis. Pronotum metallic green to metallic coppery. PW / PL = 1.14 - 1.26; PAW / PBW = 0.76 - 0.92 (Fig. 22); pronotum subquadrate with anterior angles rounded, not projected forward; disc gently convex, sparsely punctate. Elytral intervals convex basally, flat apically, densely punctate and pubescent. Hind wings developed. Metepisterna long, MW / ML = 0.75 - 0.92 (Fig. 14). Distal half of femora red-brown, the rest of legs black. This species is similar to Chlaenius (Haplochlaenius) costiger Chaudoir, 1856, but intervals interval convex basally and flat apically, not costulate.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
FCC2DE3B0936587DA515BAE3CFA8A826.taxon	description	Description. BL = 22.3 - 25.3 mm, BW = 8.5 - 10.4 mm, PAW = 3.5 - 4.0 mm, PBW = 4.3 - 4.6 mm, PW = 5.5 - 5.8 mm, PL = 4.5 - 4.9 mm, MW = 1.9 - 2.5 mm, ML = 2.4 - 3.0 mm. Head, elytra, and venter black; pronotum metallic green to metallic coppery; antennae, labial and maxillary palpi, apex of mouthparts and tarsomeres dark brown; distal half of femora red-brown, the rest of legs black. Head. Vertex finely punctate and pubescent, with a glabrous area in the middle; antennae long, reaching middle of elytra; antennomere 3 ~ 1.7 x longer than antennomere 4. Pronotum subquadrate, PW / PL = 1.14 - 1.26 (Fig. 22), widest at apical four-ninth; anterior margin slightly concave, PAW / PBW = 0.76 - 0.92; lateral margins rounded or straight before posterior angles; anterior angles rounded, not projected forward; posterior angles obtuse; disc gently convex, sparsely punctate, without glabrous area in the middle; median line distinct, fine, reaching anterior margin and base; basal foveae deep, punctate and pubescent. Elytra elongate, EL / BW = 1.47 - 1.83; slightly convex, widest near posterior third, rounded at apex in males, subtruncate in females; parascutellar striae well developed; parascutellar pores present; striae with deep punctures; interval convex basally, flat apically, densely punctate and pubescent; sutural angles obtuse; hind wings developed. Venter densely punctate, pubescent; metepisterna (Fig. 14) long, MW / ML = 0.75 - 0.92; abdominal sternites III-VI with one setiferous puncture each side, sternite VII with one pair of setiferous punctures in males, two pairs in females. Legs long and slender; tarsi nearly smooth dorsally. Male genitalia. Median lobe (Fig. 30 B-E) large, long, strongly bent to ventral side; apical orifice opened dorsally, long and wide, not reaching basal bulb; in dorsal view, apical lamella (Fig. 30 A) slightly bent to left side, width slightly longer than length, apex rounded; in left lateral view, apical portion slightly bent ventrally at apex, left side near apical lamella with a large denticulation; internal sac with flagellum slender, apex with a helical sclerite, not reaching to apical orifice; left paramere larger than right paramere, both rounded; endophallus with flagellum fine and slightly bent; basal part of flagellum without a sclerite; apical part of flagellum with a drop-shaped bursa. Female genitalia. Bursa copulatrix (Fig. 38 A-C) very long, base with trapeziform protrusion; villous canal long, tortuously contorted, adhered to common oviduct; spermatheca and spermathecal gland absent.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
FCC2DE3B0936587DA515BAE3CFA8A826.taxon	distribution	Distribution. (Fig. 40) China (Fujian; Guangdong; Guangxi; Guizhou; Hainan; Hong Kong; Yunnan), Indonesia, Laos, Myanmar, Vietnam.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
148A3BC98D235200AD3F0D3C105A4752.taxon	description	Figs 6 A, B, 13, 20, 28 A-E, 40	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
148A3BC98D235200AD3F0D3C105A4752.taxon	materials_examined	Material examined. China - Yunnan Prov.: 4 ♂♂ (IZAS), Pingbian, Dawei Shan, 2100 m, 2010. V. 23, X. D. Yang, X. Y. Zhu leg.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
148A3BC98D235200AD3F0D3C105A4752.taxon	diagnosis	Diagnosis. Dorsum black. PW / PL = 1.02 - 1.07; PAW / PBW = 0.98 - 1.00 (Fig. 20); pronotum subquadrate with anterior angles rounded, slightly projected forward; disc densely and completely rugose, punctate. Elytral intervals flat and densely punctate and pubescent. Hind wings reduced. Metepisterna short, MW / ML = 1.3 - 1.4 (Fig. 13). Apical lamella of median lobe truncated, right side with a large denticulation, left side with a small denticulation (Fig. 28 A-E), sometimes, such denticulation absent.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
148A3BC98D235200AD3F0D3C105A4752.taxon	description	Description. BL = 19.6 - 21.9 mm, BW = 7.3 - 7.6 mm, PAW = 3.33 - 3.55 mm, PBW = 3.40 - 3.55 mm, PW = 4.35 - 4.55 mm, PL = 4.05 - 4.45 mm, MW = 1.65 - 1.80 mm, ML = 1.20 - 1.35 mm. Head, pronotum, elytra, legs, and venter black; antennae, labial and maxillary palpi, apex of mouthparts and tarsomeres dark brown. Head. Vertex punctate and pubescent with a rugose area; antennae long, reaching middle of elytra; antennomere 3 ~ 1.5 x longer than antennomere 4. Pronotum subquadrate, PW / PL = 1.02 - 1.07 (Fig. 20), widest at apical third; anterior margin slightly concave, its width equal to width of basal margin, PAW / PBW = 0.98 - 1.00; lateral margins slightly bent before middle, then gently narrowed to base, straight before posterior angles; anterior angles rounded, slightly projected forward; posterior angles right angled, slightly rounded at tips; disc gently convex, sparsely punctate, pubescent, with a few transverse rugosities, without glabrous area in the middle; median line distinct, deep, reaching anterior margin and base; basal foveae deep, short, broad, punctate, and pubescent. Elytra elongate, EL / BW = 1.64 - 1.77; gently convex near anterior third, widest near posterior third, rounded at apex in males; striae with deep punctures; parascutellar striae well developed; parascutellar pores present; intervals flat, densely punctate and pubescent; sutural angles sharp; hind wings reduced. Venter densely punctate, pubescent, metepisterna (Fig. 13) short, MW / ML = 1.30 - 1.38; abdominal sternites III-VI with one setiferous puncture each side, sternite VII with one pair of setiferous punctures in males, two pairs in females; all abdominal sternites with distinct impressions laterally. Legs long and slender; tarsi nearly smooth dorsally. Male genitalia. Median lobe (Fig. 28 B-E) stout, strongly bent to ventral side; apical orifice opening dorsally, long and wide, not reaching basal bulb; in dorsal view, apical lamella (Fig. 28 A) wide and short, wider than long, apex truncated; in left lateral view, apical right side with a large denticulation towards the base and outside, basal left side only thickened and convex or convex into a small denticulation, basal orifice ~ 90 ° relative to preapical shaft; left paramere large and round; right paramere helically curved; endophallus with flagellum coiled; basal part of flagellum with a disciform sclerite facing the right; apical part of flagellum with strip of sclerite. Female genitalia unknown.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
148A3BC98D235200AD3F0D3C105A4752.taxon	distribution	Distribution. China (Yunnan), Vietnam.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
3A0D0D884CFB57FF84620CC29715BF18.taxon	description	Figs 4 A-D, 12, 21, 29 A-E, 37 A-C, 40	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
3A0D0D884CFB57FF84620CC29715BF18.taxon	diagnosis	Diagnosis. Pronotum metallic coppery to green. PW / PL = 1.12 - 1.21; PAW / PBW = 0.84 - 0.91 (Fig. 21); pronotum cordate with posterior angles right angled, rounded at tips; disc sparsely punctate and pubescent with shallow, transverse rugosities. Elytral intervals distinctly convex, with a row of setae laterally and sparse setae centrally. Hind wings reduced. Metepisterna short or width nearly equal to length; MW / ML = 1.03 - 1.17 (Fig. 12). Distal half of femora red-brown, the rest of legs black.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
3A0D0D884CFB57FF84620CC29715BF18.taxon	description	Description. BL = 20.9 - 21.6 mm, BW = 7.5 - 7.9 mm [BL = 21.3 mm, BW = 7.7 mm in holotype], PAW = 3.4 - 3.6 mm, PBW = 3.7 - 4.0 mm, PW = 4.8 - 5.3 mm, PL = 4.1 - 4.5 mm, MW = 1.7 - 1.8 mm, ML = 1.5 - 1.6 mm. Head, elytra, and venter black; pronotum metallic green to metallic coppery; antennae, labial and maxillary palpi, apex of mouthparts, and tarsomeres dark brown; distal half of femora red-brown, the rest of legs black. Head. Vertex sparsely, finely punctate and pubescent; antennae long, reaching middle of elytra; antennomere 3 ~ 1.5 x longer than antennomere 4. Pronotum cordiform, PW / PL = 1.12 - 1.21 (Fig. 21), widest at apical third; anterior margin slightly concave, PAW / PBW = 0.84 - 0.91; lateral margins rounded before middle, then distinctly narrowed to base, straight before posterior angles; anterior angles rounded, moderately projected forward; posterior angles almost right angled, slightly sharp at tips; disc sparsely punctate and pubescent, with shallow transverse rugosities, without glabrous area in the middle; median line distinct, not reaching anterior margin and base; basal foveae deep and long, punctate, pubescent. Elytra elongate, EL / BW = 1.67 - 1.73, gently convex, widest near posterior third, rounded at apex in males, subtruncate in females; basal margin sinuate, reaching the scutellum (but slightly obsolete on one side in a female); striae with deep punctures; parascutellar striae well developed; parascutellar pores present; intervals distinctly convex, with a row of setae laterally and sparse setae centrally; sutural angles obtuse; hind wings reduced. Venter sparsely pubescent, punctate; metepisterna (Fig. 12) short or width nearly equal to length, MW / ML = 1.03 - 1.17; abdominal sternites III-VI with one setiferous puncture each side, sternite VII with one pair of setiferous punctures in males, two pairs in females; all abdominal sternites with a few impressions laterally. Legs long and slender; tarsi nearly smooth dorsally. Male genitalia. Median lobe (Fig. 29 B-E) long, strongly bent to ventral side; apical orifice opened dorsally, long and wide, not reaching basal bulb; in dorsal view, apical lamella (Fig. 29 A) wide and short, wider than long, apex subtruncate, each side distinctly widened and thickened; in left lateral view, apex with a denticulation bent to back; in right lateral view, each side convex into a denticulation, right side smaller than left side, basal orifice ~ 90 ° relative to preapical shaft; left paramere large and oval; right paramere helically curved; endophallus with flagellum thick and straight; basal part of flagellum with a disciform sclerite facing the left. Female genitalia. Bursa copulatrix (Fig. 37 A-C) asymmetric, base with a bifid irregular protrusion; villous canal long, tortuously contorted, adhered to common oviduct; spermatheca and spermathecal gland absent.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
3A0D0D884CFB57FF84620CC29715BF18.taxon	distribution	Distribution. (Fig. 40) China (Hainan).	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
3A0D0D884CFB57FF84620CC29715BF18.taxon	etymology	Etymology. The new species yinggelingensis is named for the type locality Yinggeling, Hainan.	en	Qin, Yuyao, Germann, Christoph, Liang, Hongbin (2022): Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species. ZooKeys 1135: 61-91, DOI: http://dx.doi.org/10.3897/zookeys.1135.93843, URL: http://dx.doi.org/10.3897/zookeys.1135.93843
