taxonID	type	description	language	source
F63D5003A7C954F2AEF3A9082F7BD110.taxon	type_taxon	Type species. × Benzoscaphe stelleri Pupulin, Karremans & E. Salas	en	Pupulin, Franco, Karremans, Adam P., Salas Hidalgo, Elvira (2025): A new nothogenus and nothospecies in the Zygopetalinae (Orchidaceae), with a note on euglossine pollination in the subtribe. PhytoKeys 266: 33-52, DOI: 10.3897/phytokeys.266.159892
F63D5003A7C954F2AEF3A9082F7BD110.taxon	diagnosis	Diagnosis. Nothogenus novum caracteribus intermediis inter Benzingiam Dodson et Chondroscaphen (Dressler) Senghas et G. Gerlach. Folia viridi-grisacea sicut Benzingiarum species sed erecta prod pendulis sicut Chondroscapharum species; sepalis lateralibus omnino reflexis subuncinatis, basi petalorum ad columna adnata, pede columnae elongato Benzingiae similiter; inflorescentia suberecta prod pendula, petalis subtrapezoideis, labello callis duobus ornato, anthera elongata anguste ovata, polliniis elongatis valde curvatis supra stipitem rectangulo concavo-convexo viscidioque incrassato sicut Chondroscapharum species; floris magnitudo et forma labelli intermediis inter genera dignoscenda.	en	Pupulin, Franco, Karremans, Adam P., Salas Hidalgo, Elvira (2025): A new nothogenus and nothospecies in the Zygopetalinae (Orchidaceae), with a note on euglossine pollination in the subtribe. PhytoKeys 266: 33-52, DOI: 10.3897/phytokeys.266.159892
F63D5003A7C954F2AEF3A9082F7BD110.taxon	description	Description. Epiphytic, caespitose, erect herbs with a tuft of leaves fan-like arranged. Roots slender, flexuous, whitish. Leaves narrowly elliptic, acuminate, with a short conduplicate petiole, grayish green, articulated with basal, ovate, conduplicate sheaths provided with hyaline margins. Inflorescence a single-flowered raceme, suberect, much shorter than the leaves. Floral bracts two, the outer bract larger, broadly ovate, loose, the inner bractlet lanceolate, smaller. Pedicellate ovary terete, gently curved toward the apex. Flower spreading, downward facing, the dorsal sepal porrect-recurved, the lateral sepals reflexed, subuncinate, the petals inserted along the basal side of the column, porrect with the apex gently recurved, the lip obscurely three-lobed, the lateral lobes erect, the midlobe geniculate-bent, with ruffled margins; disc with a median, irregularly dentate callus and a second thickening toward the apex of the midlobe. Column hemiterete, dilated around the stigma, provided with a long foot, the stigma transversal, slit-like, the clinandrium swallow, the anther ovate; pollinia 4 in two superposed pairs of different sizes, on a rectangular, erected stipe and a narrowly lanceolate, thick viscidium. The new nothogenus presents intermediate features between the genera Benzingia (Fig. 1) and Chondroscaphe (Fig. 2). The leaves are grayish green with the epidermis somewhat prismatic, like in Benzingia, but they are held erect as in species of Chondroscaphe. The size of the flower is intermediate between the smaller flower of Benzingia (natural spread of flower ca. 3 cm) and larger ones typical of Chondroscaphe (spread flowers> 6 cm). The flower of the nothogenus is similar to that of Benzingia in the lateral sepals completely reflexed and uncinate at the apex, the base of the petals adnate to the margins of the column, and the elongate column foot. It resembles Chondroscaphe in the subtrapezoid petals, the lip provided with a basal callus and a distinct apical bulging, the narrowly ovate, elongate anther cap, and the shape of the pollinarium, with four elongate, distinctly curved pollinia lying over a rectangular, concave-convex stipe, attached to a thick viscidium.	en	Pupulin, Franco, Karremans, Adam P., Salas Hidalgo, Elvira (2025): A new nothogenus and nothospecies in the Zygopetalinae (Orchidaceae), with a note on euglossine pollination in the subtribe. PhytoKeys 266: 33-52, DOI: 10.3897/phytokeys.266.159892
502E55463A735869920DEAA98F583823.taxon	diagnosis	Diagnosis. Nothospecies nova caracteribus intermediis inter Benzingiam reichenbachianam (Schltr.) Dressler et valde probabiliter Chondroscaphen bicolorem (Rolfe) Dressler dignoscenda. Folia viridi-grisacea sicut Benzingiae reichenbachianae sed erecta prod pendulis sicut Chondroscaphe bicolor; sepalis lateralibus omnino reflexis subuncinatis, basi petalorum ad columna adnata, pede columnae elongato Benzingiae similiter; inflorescentia suberecta prod pendula, petalis subtrapezoideis, labello callis duobus ornato, anthera elongata anguste ovata, polliniis elongatis valde curvatis supra stipitem rectangulo concavo-convexo viscidioque incrassato sicut Chondroscaphe bicolor; floris magnitudo et forma labelli intermediis inter duabus speciebus.	en	Pupulin, Franco, Karremans, Adam P., Salas Hidalgo, Elvira (2025): A new nothogenus and nothospecies in the Zygopetalinae (Orchidaceae), with a note on euglossine pollination in the subtribe. PhytoKeys 266: 33-52, DOI: 10.3897/phytokeys.266.159892
502E55463A735869920DEAA98F583823.taxon	description	Description. Epiphytic, caespitose, erect herbs with a tuft of 4 – 5 leaves arranged like a fan. Roots flexuous, slender, ca. 1 mm in diameter, whitish, the apex green. Leaves narrowly elliptic, acuminate, 8.0 – 20.0 × 1.0 – 1.8 cm, with a short conduplicate petiole to 0.5 cm long, grayish green, paler on the underside, articulated with basal, ovate, strongly conduplicate sheaths 1.4 – 3.0 cm long, the margins scabrous-hyaline. Inflorescence a single-flowered raceme, suberect, to 5 cm long; peduncle terete, slender, ca. 3 cm long. Floral bracts two, the outer bract larger, broadly ovate, loose, 7 × 6 mm, the inner bractlet lanceolate, smaller, 7 × 2 mm. Pedicellate ovary terete-subclavate, gently curved toward the apex, 2 cm long. Flower spreading, downward facing, the sepals and petals creamy white, the petals marked with few, scattered rose-purple spots near the base, the lip white, flushed pale yellow toward the base, with lines of purple spots (nectar guides) running from the base to about two-thirds of the blade, the callus on the disc yellow, the column white, sparsely dotted with purple-red at the base, the anther cap cream to light brown. Dorsal sepal lanceolate, acute, 15 × 7 mm, porrect-recurved. Lateral sepals narrowly lanceolate, acute, abruptly apiculate, 25 × 5 mm, strongly reflexed, subuncinate at apex, the labellar margin enrolled from the base to the middle. Petals inserted along the basal side of the column, obliquely obovate to narrowly subrhombic, truncate, 22 × 8 mm, porrect, the apex gently recurved. Lip obscurely three-lobed, 28 × 21 mm, the lateral lobes elliptic, erect to flank the column, the midlobe broadly ovate-trapezoid, apically bilobed, geniculate-bent, apically shortly reflexed, with ruffled margins; disc with a median, irregularly dentate callus, slightly elevated callus in the middle of laminate, and a second thickening toward the apex of the midlobe. Column hemiterete, ca. 10 mm long, dilated around the stigma into elliptic-subtriangular wings, 5 mm wide at the broadest point, provided with a long foot ca 10 mm long, the stigma transversal, slit-like, the rostellum acicular, flanked by 2 pararostellar teeth, the clinandrium low-cuspidate, swallow. Anther cap ovate, apically refuse, 2 - celled. Pollinia 4, narrowly ovate, complanate, in two superposed pairs of different sizes, on a short, rectangular, erected stipe and a narrowly lanceolate, thick viscidium.	en	Pupulin, Franco, Karremans, Adam P., Salas Hidalgo, Elvira (2025): A new nothogenus and nothospecies in the Zygopetalinae (Orchidaceae), with a note on euglossine pollination in the subtribe. PhytoKeys 266: 33-52, DOI: 10.3897/phytokeys.266.159892
502E55463A735869920DEAA98F583823.taxon	distribution	Distribution. Only known from central Costa Rica, where it has been so far documented only from about 1500 meters of elevation along the southeastern slopes of the Central Volcanic Cordillera, facing the Central Valley (Fig. 6).	en	Pupulin, Franco, Karremans, Adam P., Salas Hidalgo, Elvira (2025): A new nothogenus and nothospecies in the Zygopetalinae (Orchidaceae), with a note on euglossine pollination in the subtribe. PhytoKeys 266: 33-52, DOI: 10.3897/phytokeys.266.159892
