taxonID	type	description	language	source
077D87BC5F3F5213FF10FB75FF3914D6.taxon	materials_examined	Material examined: Tunisia: 1, Lake Ischkeul, 7. IV. 1986, leg. Schillhammer (NHMW).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3F5213FF10FB75FF3914D6.taxon	discussion	Comment: This species has been recorded only from Tunisia and Algeria (ASSING 2011).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3F5210FF10FA4DFC8A1046.taxon	materials_examined	Material examined: Kazakhstan: 1, 1, Alma-Ata region, Pristan-Dubinkaja, 43 ° 45 ' N, 80 ° 13 ' E, 550 m, 15. - 18. VI. 1993, leg. Lukthanov (NME); 1, Taldy-Kurgan region, Koktal, 44 ° 06 ' N, 70 ° 47 ' E, 500 m, 8. VI. 1993, leg. Lukthanov (cAss).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3F5210FF10FA4DFC8A1046.taxon	discussion	Comment: Tetartopeus rufonitidus is one of the most widespread species of the genus, its distribution ranging from West Europe to Middle Asia.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5210FF10FEBDFEE7135B.taxon	discussion	Comment: In a recent paper, BORDONI (2013) proposed several, partly rather bizarre synonymies, among them the (repeated) synonymy of T. ciceronii with T. rufonitidus REITTER (1909), without providing any new evidence whatsoever. As argued in detail earlier (ASSING 2008), T. ciceroni undoubtedly represents a distinct species differing from T. rufonitidus in numerous constant characters and is consequently revalidated.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5210FF10FC46FC601505.taxon	materials_examined	Material examined: Turkey: 2, Muğla, SE Muğla, 26 km W Köyceğiz, 17. V. 1991, leg. Schönmann & Schillhammer (NHMW, cAss).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5210FF10FC46FC601505.taxon	discussion	Comment: Previously, only the type material from Izmir was known.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5210FF10FDB6FD371392.taxon	materials_examined	Material examined: Spain: 2, Cádiz, Algeciras env., 36.08 ° N, 5.43 ° W, 50 m, 11. - 12. V. 2014, leg. Shavrin (cSha, cAss).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5210FF10FDB6FD371392.taxon	discussion	Comment: This species is apparently subject to remarkable variation in size. The two males listed above differ considerably in this respect.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5210FF10FC8EFEAE12EA.taxon	materials_examined	Material examined: Greece: 1, Ebros Delta, 17. V. 1991, leg. Schillhammer (NHMW).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5210FF10FC8EFEAE12EA.taxon	discussion	Comment: Tetartopeus scutellaris is one of the most widespread red-winged species of the genus.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5210FF10FBFCFBEA1458.taxon	materials_examined	Material examined: Turkey: 1 [identification based on a photo of the aedeagus], Hatay, 15 km WSW Antakya, Batıayaz, foot of Musa Daĝı, 500 m, pitfall, 6. - 23. IV. 2014, leg. Reuter (cFel).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5210FF10FBFCFBEA1458.taxon	discussion	Comment: The distribution of T. tezcani is confined to central southern Anatolia.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5211FF10FAB3FEC01045.taxon	materials_examined	Material examined: Turkey: 1, Artvin, Şavşat - Veliköy, pond, 5. VI. 1989, leg. Schönmann & Schillhammer (NHMW). Iran: 2 exs., Tehran province, N Tehran, Elburz Mts., Darake, Palanchal, 35 ° 51 ' N, 51 ° 23 ' E, 2250 m, 31. V. 2010, leg. Frisch (MNHUB, cAss); 1 ex., Tehran province, S-Elburz, Aghasht, 36 ° 00 ' N, 50 ° 53 ' E, 1600 - 1700 m, 27. V. 2008, leg. Weipert (cRou); 1 ex., Ilam province, 10 km S Ilam City, 33 ° 34 ' N, 46 ° 25 ' E, 1300 m, 19. X. 2011, leg. Frisch (MNHUB); 1 ex., Gilan province, S Astara, Lavandvil, 38 ° 18 ' N, 48 ° 50 ' E, 30 m, 10. X. 2011, leg. Frisch (cAss); 1 ex., Chahar province, Mahali-o-Bakhtiyari, E Haruni, 32 ° 24 ' N, 50 ° 34 ' E, 2300 m, 5. IV. 2012, leg. Weipert (cAss).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3C5211FF10FAB3FEC01045.taxon	discussion	Comment: The distribution of T. stylifer ranges from Turkey and Ukraine to Iran and Iraq.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3D5211FF10FEB3FC43109E.taxon	materials_examined	Material examined: Israel: 1, Berekhat Ya'ar, 32 ° 25 ' N, 34 ° 45 ' E, 10 m, shore of pond, pitfall, 17. III. 2010, leg. Drees (cAss).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3D5211FF10FEB3FC43109E.taxon	discussion	Comment: Previously, only the two type specimens from two localities in Israel (Ma'agar Bental and Ein Afek Reserve) were known (FELDMANN 2010).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3D5214FF10FD8BFE221000.taxon	description	(Figs 1 - 6, 8)	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3D5214FF10FD8BFE221000.taxon	materials_examined	Material examined: Russia: 1, Krasnodar, Temnolesskaia near Mezmai, 850 m, 19. VI. 1999, leg. Smetana [R 18] (cAss); 1, Krasnodar, S Labisnk, Mostovskoj env., 500 m, forest along Khodz river valley, edge of a water pool, 14. V. 1996, leg. Solodovnikov (ZMUC); 1, Krasnodar, foothills of Bambaki mountain range, ca. 20 km SW Psebai, Port Artur, ca. 1000 m, stream bank, 8. VIII. 1993, leg. Solodovnikov (ZMUC).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3D5214FF10FD8BFE221000.taxon	discussion	Comment: The identity of this species has been addressed in several previous articles (ROUBAL 1913; BORDONI 1980; COIFFAIT 1982; GUSAROV 1991). However, the descriptive details (especially regarding the coloration) and the illustrations of the male sexual characters, particularly those of the aedeagus, are either incorrect, misleading, or insufficient for a reliable identification. Therefore, a complete redescription and new illustrations are provided below.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3D5214FF10FD8BFE221000.taxon	description	Redescription: Body length 6.6 - 8.0 mm; length of forebody 3.7 - 4.3 mm. Habitus as in Fig. 1. Coloration distinctive: head black; pronotum red, posterior margin reddish or weakly infuscate; elytra bicoloured, with the anterior half more or less distinctly and more or less extensively infuscate, and with the posterior half reddish; abdomen black, segments VIII-X brown; legs dark-yellowish; antennae dark-brown, with the basal and the apical antennomeres reddish. Head (Fig. 2) weakly oblong, approximately 1.05 times as long as broad; punctation moderately coarse and dense, distinctly sparser in median dorsal portion; interstices without microsculpture. Eyes moderately large and weakly convex, approximately half as long as distance from posterior margin of eye to posterior constriction in dorsal view. Antenna slender, 2.2 - 2.8 mm long. Pronotum (Fig. 2) rather large, approximately 1.15 times as long as broad and 1.15 times as broad as head; punctation dense, slightly coarser than that of head; interstices on average slightly narrower than diameter of punctures, without microsculpture; midline narrowly impunctate. Elytra (Fig. 2) 0.8 - 0.9 times as long as pronotum; punctation rather fine, dense, and weakly defined. Hind wings present. Metatarsomere I as long as II. Abdomen slightly narrower than elytra; surface nearly completely matt due to extremely dense and fine punctation and distinct microsculpture; posterior margin of tergite VII with palisade fringe.: tergite VIII with broadly and rather weakly convex posterior margin; sternites IV-V in the middle with some coarse and irregularly spaced puncture-like impressions; sternite VI weakly impressed along the middle; sternite VII (Fig. 3) strongly transverse, with shallow median impression and with fine and dense pubescence, at posterior margin with transverse row of long marginal setae, posterior margin broadly and weakly concave; sternite VIII (Fig. 4) approximately 1.3 times as broad as long, with moderately deep, narrowly and asymmetrically V-shaped posterior excision in slightly asymmetric position, near this excision with denser pubescence; aedeagus (Figs 5 - 6) 1.5 mm long; ventral process conspicuously long and straight with weakly curved apex (lateral view); dorsal plate broad and apically acute in dorsal view (Fig. 8).: tergite VIII with strongly convex posterior margin.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3D5214FF10FD8BFE221000.taxon	discussion	Comparative notes: Tetartopeus lomnickii is characterized particularly by the coloration, the shape and chaetotaxy of the male sternite VIII, and by the morphology of the aedeagus. It is readily distinguished from all other red-winged Tetartopeus species with a dark abdominal apex (T. czwalinai (JAKOBSON, 1909) from West Anatolia; T. unguis ASSING 2010, T. vomer ASSING 2010, and T. inexcisus ASSING 2009 from North Anatolia; the widespread T. scutellaris (NORDMANN, 1837) and T. angustatus (LACORDAIRE, 1835 )) by the distinctly broader and more robust body and by the reddish pronotum alone. Aside from T. lomnickii, five Tetartopeus species have been recorded from the West Caucasus: T. terminatus (GRAVENHORST, 1802), T. quadratus (PAYKULL 1789), T. rufonitidus (REITTER, 1909), T. scutellaris, and T. stylifer (REITTER, 1909). In all these species the pronotum is usually black. The first four species have the elytra blackish, sometimes with the postero-lateral angles more or less distinctly yellowish. In T. stylifer, the abdominal apex is reddish and in T. scutellaris the head is distinctly more slender than in T. lomnickii. For illustrations of the aedeagi of these species see COIFFAIT (1982) and ASSING (2012 b).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F3D5214FF10FD8BFE221000.taxon	distribution	Distribution and natural history: The known distribution of T. lomnickii is confined to the West Caucasus. For previous records see ROUBAL (1913), GUSAROV (1991), and SOLODOVNIKOV (1998 a, b). The examined male from Temnolesskaia was sifted from wet leaf litter and other debris along a stream in a beech forest (SMETANA pers. comm.) at an altitude of 850 m, the other two specimens at the edge of a water pool and on a stream bank at altitudes of 500 and 1000 m (SOLODOVNIKOV pers. comm.).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F385214FF10FEF8FCFC13FB.taxon	description	(Figs 7, 9 - 10)	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F385214FF10FEF8FCFC13FB.taxon	materials_examined	Material examined: Russia: 1, 1, N-Mongolia, " Shangai ", leg. Leder (NHMW, cAss).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F385214FF10FEF8FCFC13FB.taxon	discussion	Comment: Tetartopeus lentus was originally described from the Transbaikal region; the aedeagus is illustrated by RYVKIN (1989). The previously unknown female secondary sexual characters are as follows:	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F385214FF10FEF8FCFC13FB.taxon	description	: tergite VIII (Fig. 9) transverse, with almost truncate and in the middle weakly pointed posterior margin; sternite VIII (Fig. 7) much longer than tergite VIII, posteriorly produced and broadly truncate; tergite IX (Fig. 10) deeply and narrow incised posteriorly.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F385214FF10FEF8FCFC13FB.taxon	discussion	The above specimens represent the first record from Mongolia.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F38521AFF10FD54FF331701.taxon	description	(Figs 11 - 16, 18 - 19)	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F38521AFF10FD54FF331701.taxon	materials_examined	Type material: Holotype: " Anatolia or., leg. H. Korge / Benliahmet sw. Kars, 31. VII. 1967 / Lathrobium (Tetartopeus) nov. spec. / Holotypus Tetartopeus korgei sp. n., det. V. Assing 2014 " (MNHUB). Paratypes: 1, 1 [both teneral]: same data as holotype.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F38521AFF10FD54FF331701.taxon	etymology	Etymology: This species is dedicated to the late Horst Korge (Berlin), who collected the type material and who took a special interest in the Staphylinidae fauna of Turkey.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F38521AFF10FD54FF331701.taxon	description	Description: Body length 6.2 - 6.8 mm; length of forebody 3.5 - 3.7 mm. Coloration: body (including apex of abdomen) black, except for the reddish posterior 3 / 5 of the elytra; legs yellowish; antennae dark-brown with antennomeres I-II dark-yellowish. Head (Fig. 11) approximately 1.1 times as long as broad; punctation rather sparse and relatively fine to moderately coarse; interstices on average distinctly broader than diameter of punctures, without microsculpture. Eyes moderately large and weakly convex, approximately half as long as distance from posterior margin of eye to posterior constriction in dorsal view. Antenna slender, approximately 2.2 mm long Pronotum (Fig. 11) approximately 1.2 times as long as broad and 1.15 times as broad as head; punctation moderately dense; interstices on average as broad as, or slightly narrower than, diameter of punctures, without microsculpture; impunctate midline moderately broad. Elytra (Fig. 11) short, 0.74 - 0.78 times as long as pronotum; punctation rather dense and coarse. Hind wings present, but possibly of reduced length. Metatarsomere I as long as II. Abdomen approximately as broad as elytra; surface nearly completely matt due to extremely dense and fine punctation and distinct microsculpture; posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII without appreciable sexual dimorphism, convex, indistinctly pointed in the middle (Fig. 12).: sternite VIII (Fig. 13) approximately 1.2 times as broad as long, in postero-median portion with longer and denser dark pubescence, posterior margin slightly convexly projecting in the middle, this projection with minute median concavity; aedeagus (Figs 14 - 16) 1.2 mm long; ventral process apically conspicuously bifurcate in lateral view; dorsal plate weakly sclerotized, apically convex in dorsal view.: tergite VIII (Fig. 18) weakly oblong, posterior margin nearly truncate in the middle; segments IX-X as in Fig. 19.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F38521AFF10FD54FF331701.taxon	discussion	Comparative notes: Regarding the shape of the ventral process of the aedeagus, the new species is most similar to T. tezcani ANLAŞ, 2009 from central southern Anatolia, from which it differs by the dark apex of the abdomen (T. tezcani: apex reddish), the anteriorly broadly blackish elytra (T. tezcani: very narrowly infuscate at most), and by the straight ventral process of the aedeagus with two apical projections of similar length (T. tezcani: ventral process of aedeagus distinctly curved in lateral view, dorso-apical projection much shorter than ventro-apical projection).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F38521AFF10FD54FF331701.taxon	description	Distribution and natural history: The type locality is situated to the southwest of Kars, Kars province, northeastern Anatolia. The paratypes are distinctly teneral.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F37521BFF10FF5DFBC8103E.taxon	materials_examined	Material examined: Russia: 4 exs., Transbaikal region, Werchne-Undinskoje, leg. Mandl (NHMW, cAss).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F37521BFF10FF5DFBC8103E.taxon	discussion	Comment: This East Palaearctic species is distributed in Siberia, the Russian Far East, and Mongolia. For illustrations of the external and sexual characters see ASSING (2011).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F37521BFF10FD7DFC8A12B8.taxon	materials_examined	Material examined: Japan: H o k k a i d o 4 exs., Tomakomai, Tokisatamappu marsh, 10 m, 25. VII. 1991, leg. Smetana [J 6] (CNC); 4 exs., Kushiro Marsh, between Horo and Oshima rivers, 5 m, 31. VII. 1991, leg. Smetana [J 21] (CNC, cAss). Honshu: 1, 1, Gumma Pref., near Mt. Shirane, 1500 m, 22. VII. 1980, leg. A. & Z. Smetana (CNC); 1, same data, but 2000 m (CNC); 1, Nikko National Park, Senjugahara, 1400 m, 15. VII. 1980, leg. A. & Z. Smetana (CNC); 1, Nikko National Park, Senjugahara, 1400 m, 15. VII. 1980, leg. A. & Z. Smetana (CNC).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F37521BFF10FD7DFC8A12B8.taxon	discussion	Comment: This species has been recorded only from Japan.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F37521BFF10FE15FCCD1385.taxon	materials_examined	Material examined: Russia: 1, Ussuri region, Novovarvarovka, 6. - 10. VII. 1989, leg. Nikodým (NHMB); 1, Russian Far East, Ussuryk District, " S. dk. Belodorodov ", Kamenushka, 10. - 18. VII. 1992 (cAss). Japan: H o k k a i d o: 10 exs., Tomakomai, Tokisatamappu marsh, 10 m, 25. VII. 1991, leg. Smetana [J 6] (CNC, cAss); 5 exs., Rubeshibe, Itonmuka river, marsh, 770 m, 29. VII. 1991, leg. Smetana [J 16] (CNC, cAss); 4 exs., Oshamanbe, Toyotsu, seashore, 27. VII. 1991, leg. Smetana [J 10] (CNC, cAss); 4 exs., Horokanai, Butokamabetsu river, 300 m, 3. VIII. 1991, leg. Smetana [J 31] (CNC, cAss); 27 exs., Kushiro Marsh, Iwahogi, 5 m, 31. VII. 1991, leg. Smetana [J 20] (CNC, cAss).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F37521BFF10FE15FCCD1385.taxon	discussion	Comment: In the Palaearctic region, the distribution of this Holarctic species is confined to the Russian Far East and Japan (ASSING 2011.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F375218FF10FB93FEA7170C.taxon	description	(Figs 17, 20)	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F375218FF10FB93FEA7170C.taxon	materials_examined	Type material examined L. gracilentum: see ASSING (2012 a). L. pallipes: see ASSING (2011). L. bimaculatum: see ASSING (2011) L. maculatum: Paratypes: 1: " S 264, Blackwater River NG, 6.74 Rff / Manchester Museum, Paratype / Paratype / F 3008.5152 / Lathrobium maculatum sp. n., H. R. Last. det., Paratype / Tetartopeus gracilentus (Kraatz), det. V. Assing 2014 " (MMUM); 1: same data, but " F 3008.5152 " (MMUM). Additional material examined: South Korea: 9 exs., Gangwon-do, 6 km NW Yeounpo-ri, Hajodae beach, 38 ° 02 ' N, 128 ° 42 ' E, 0 m, small lagoon lake 300 m W seashore, wet muddy litter under stand of reeds, flotation, 10. IX. 2010, leg. Makranczy et al. (HNHM, NIBR, cAss); 1, Jeollanam-do, Jiri-san, 2 km WNW Nogodan peak, waterfall near Kogae pass, 35 ° 18 ' N, 127 ° 31 ' E, 1270 m, from wet moss and litter at spraying water, flotation, 15. IX. 2010, leg. Makranczy et al. (HNHM). Japan: 3, 2, Oshamanbe, Toyotsu, seashore, 27. VII. 1991, leg. Smetana [J 10] (CNC, cAss); 1, Ryukyu Islands, Ishigaki Island, Maezato Dam, disturbed secondary forest, flight interception trap, 27. - 31. V. 2012, leg. Lackner (cAss).	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F375218FF10FB93FEA7170C.taxon	discussion	Comment: The original description of Lathrobium maculatum is based on a male holotype and six paratypes from " Blackwater River, Sepik " in New Guinea (LAST 1984). An examination of two paratypes yielded no evidence that they should be distinct from T. pallipes; the male sexual characters are practically identical (Figs 17, 20). The finding that T. pallipes is distributed southwards as far as New Guinea inspired a re-examination of material previously attributed to T. pallipes, and T. wui. Moreover, the characters indicated earlier (ASSING 2011, 2012) as distinguishing T. wui, T. pallipes, and T. gracilentus were reconsidered. These character conditions (head shape, coloration of the legs, punctation, size of aedeagus, shape of ventral process) are too variable (with overlap), not sufficiently pronounced, and linked by transitional conditions to further support previous hypotheses that T. gracilentus, T. wui, and T. pallipes represent different species. The observed differences are more plausibly attributed to intra- rather than interspecific variation of a widespread species with remarkable dispersal power, as evidenced also by its presence on numerous islands. Hence, Lathrobium pallipes, L. maculatum, Lobrathium wui, and Lobrathium bimaculatum are all placed in synonymy with the senior name Tetartopeus gracilentus.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
077D87BC5F375218FF10FB93FEA7170C.taxon	distribution	Distribution: The remarkably vast distribution of T. gracilentus ranges from North Japan and the Russian Far East across China to Sri Lanka in the southwest and New Guinea in the south. The previous doubtful record from South Korea (ASSING 2011) is now confirmed.	en	Assing, V. (2014): A revision of Tetartopeus IV. A new species from Turkey, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 46 (2): 1119-1131, DOI: 10.5281/zenodo.5307283
