taxonID	type	description	language	source
073A87C8E20F6B7CFF0C9E54FC065BA7.taxon	diagnosis	Diagnosis. Small-sized palaemonid shrimp with dorsoventrally somewhat flattened body. Carapace smooth, glabrous, without pubescence or pilosity. Rostrum moderately developed, mid-dorsally carinate, otherwise unarmed, including ventrally; lateral carina proximally expanded. Supraorbital, hepatic and epigastric teeth absent. Orbit well developed, inferior orbital (= infraorbital) angle distinct. Antennal tooth well developed, acute. Fourth thoracic sternite unarmed. Pleon smooth, glabrous; brood chamber of female strongly expanded laterally; posteroventral angle of first to fifth pleura rounded, sixth pleuron quadrate, posteriorly unarmed. Telson with two pairs of submarginal cuspidate setae on dorsal surface and two pairs of short spiniform setae on posterior margin. Eyes with well-developed, globular cornea; accessory eye (nebenauge) absent. Antennule with first article ending in blunt distolateral lobe, not sharp tooth; stylocerite small, triangular; lateral flagellum with accessory ramus composed of single unit. Antennal scaphocerite with well-developed, broad blade; distolateral tooth vestigial. Epistome unarmed. Mandible without palp; incisor process with five distal teeth. Maxillule with broad laciniae; palp (endopod) distally bilobed, ventral lobe with small, acute, tooth-like, distoventral extension and long stiff seta. Maxilla with palp (endopod) moderately broad; endite with deep cleft; scaphocerite well developed. First maxilliped with endites broad; palp (endopod) absent; exopod with well-developed caridean lobe; epipod moderately developed, bilobed. Second maxilliped typical for family, with epipod moderately developed, subquadrate, without podobranch. Third maxilliped pediform, not operculate; coxal lateral plate poorly developed, rounded; basis and ischiomerus fused; arthrobranch absent. First pereiopods short, fingers of chela not spatulate, not dentate. Second pereiopods (chelipeds) moderately enlarged and robust, somewhat elongate, sexually dimorphic, slightly more robust in males, subequal in size, symmetrical in shape; cutting edges of chela fingers with simple robust teeth proximally, without fossa. Ambulatory pereiopods relatively slender; ischium, merus, carpus and propodus unarmed; dactylus elongate, compressed, biunguiculate; main unguis slender, acute; corpus with single acute distal tooth (= secondary unguis), latter stouter but shorter than main unguis. Male second pleopod with well-developed appendix masculina. Uropod with lateral lobe ending bluntly; exopod with reduced distolateral tooth; diaeresis feebly developed, unarmed. Gill / exopod formula: 5 pleurobranchs (P 1 – 5), 2 epipods (Mxp 1 – 2), 0 arthrobranchs, 0 podobranchs, 3 exopods (Mxp 1 – 3).	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20F6B7CFF0C9E54FC065BA7.taxon	type_taxon	Type species. Opaepupu huna sp. nov., by monotypy and original designation. Other species included. None.	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20F6B7CFF0C9E54FC065BA7.taxon	distribution	Distribution. Central Pacific: presently only known from the Hawaiian Islands.	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20F6B7CFF0C9E54FC065BA7.taxon	etymology	Etymology. The generic name (Opaepupu) is derived from the Hawaiian word`ôpae, a general term for shrimp or prawn, and pűpű, a general term used for marine shells, referring to its association with a bivalve mollusk. Gender neuter.	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20F6B7CFF0C9E54FC065BA7.taxon	discussion	Remarks. The phylogenetic position of Opaepupu gen. nov. is not immediately obvious and it cannot be excluded that the new genus is more closely related to a genus or a group of palaemonid genera with species not associated with bivalves (see below). However, for obvious reasons, the new genus is herein morphologically contrasted with all other genera containing bivalve-associated species. Among the above-mentioned palaemonid genera known to contain shrimps associated with bivalves, Opaepupu gen. nov. appears to be morphologically most similar to the Indo-West Pacific genera Platypontonia, Anchiopontonia, Pinnotherotonia, Bruceonia, Cainonia and the Atlantic-East Pacific genus Pontonia. The remaining Indo-West Pacific genera, namely Conchodytes, Anchistus, Paranchistus and Neoanchistus appear to be much more distant to the new genus (see below). Opaepupu gen. nov. is easily distinguishable from Platypontonia, currently with two species, P. brevirostris (Miers, 1884) and P. hyotis Hipeau-Jacquotte, 1971 (Bruce 1968; Hipeau-Jacquotte 1971, 1974; Suzuki 1971). For instance, these two genera differ in the form of the rostrum, being distally undivided and having a distinct mid-dorsal carina in Opaepupu gen. nov. vs. distally bifurcate and dorsally non-carinate in Platypontonia; the shape of the dactylus of the ambulatory pereiopods, which is elongate and biunguiculate in Opaepupu gen. nov. vs. stouter and not biunguiculate in Platypontonia; the ventral lobe of the maxillular palp (endopod) with a long stiff seta dorsal to a small hook-like process in Opaepupu gen. nov. vs. with a hook-like process but without a seta, in Platypontonia; and the first maxilliped with two distinct endites and lacking palp (endopod) in Opaepupu gen. nov. (but see below) vs. with the endites completely fused and with a small palp in Platypontonia (cf. Bruce 1968; Hipeau-Jacquotte 1971). Additional, minor, differentiating characters between these genera are in the size of the cuspidate setae of the telson, which are relatively small (especially in the female) in the new genus vs. large and stout in the two species of Platypontonia; and in the first article of the antennular peduncle distolaterally unarmed (ending in a blunt lobe) vs. armed with a small distolateral tooth in Platypontonia) (cf. Bruce 1968; Hipeau-Jacquotte 1971). In addition, P. brevirostris associates with the cockscomb oyster, Lopha cristagalli (Linnaeus, 1758), whilst P. hyotis associates with large, heavily calcified oysters of the genus Hyotissa Stenzel, 1971 (as Pterostrea in Suzuki 1971); both hosts are morphologically and ecologically very different from the trapezid clam host of the type species of Opaepupu gen. nov. (see below). Opaepupu gen. nov. can be easily separated from the monotypic Anchiopontonia by the shape of the dactylus of the ambulatory pereiopods, which is elongate and strongly biunguiculate in the new genus vs. very stout and with a unique distal armature in Anchiopontonia (cf. Bruce 1992). The two genera also differ in the configuration of the rostrum, which is proximally broad and dorsally carinate in Opaepupu gen. nov. vs. rather narrow and not carinate in Anchiopontonia; the shape of the anterolateral margin of the carapace, with a strong antennal tooth and without an infraorbital lobe in Opaepupu gen. nov. vs. with a small antennal tooth and with a rounded, protruding infraorbital lobe in Anchiopontonia; the development of the distolateral tooth of the scaphocerite, which is greatly reduced in the new genus vs. well developed in Anchiopontonia; and the first maxilliped lacking a palp in Opaepupu gen. nov. vs. with a small palp in Anchiopontonia (cf. Bruce 1992). In addition, A. hurii (Holthuis, 1981) lives exclusively in association with spiny oysters of the genus Spondylus Linnaeus, 1758, which are morphologically and ecologically very different from the trapezid host of the type species of Opaepupu gen. nov. (see below). Opaepupu gen. nov. also differs in many respects from the monotypic Pinnotherotonia, containing P. rumphiusi Marin & Paulay, 2010, a rare and peculiar shrimp associated with the burrowing clam Periglypta crispata (Deshayes, 1854). The new genus can be separated from Pinnotherotonia by the body and appendages being glabrous vs. densely pubescent in Pinnotherotonia; the rostrum dorsally carinate and distally pointed vs. not carinate and distally broadly truncate in Pinnotherotonia; the posterior margin of the telson armed with two pairs of small spiniform setae vs. fringed with simple setae in Pinnotherotonia; the ventral lobe of the maxillular palp with a long stiff seta dorsal to a small hook-like process vs. with a small spinule and without a stiff seta in Pinnotherotonia; the first maxilliped with two distinct endites and lacking a palp vs. with the endites completely fused and with a small palp in Pinnotherotonia; the third maxilliped relatively slender, pediform vs. broad, suboperculiform in Pinnotherotonia; the second pereiopods more slender, with proportionally longer fingers; the dactylus of the ambulatory pereiopods elongate and not covered by setae vs. much stouter and densely covered by numerous setae on both mesial and lateral surfaces in Pinnotherotonia (cf. Marin & Paulay 2010). Some of the characters separating Opaepupu gen. nov. from Pinnotherotonia also apply to Bruceonia, presently containing only the type species, B. ardeae (Bruce, 1981), associated with the Pacific jewel-box, Chama pacifica Broderip, 1835 (Fransen 2002). These are the absence of pilosity on the body and appendages in the new genus vs. its presence on at least the carapace, all pereiopods and parts of the tail fan in Bruceonia; the rostrum being distally pointed and dorsally carinate in the new genus vs. distally truncate and dorsally not carinate in Bruceonia; the ventral lobe of the maxillular palp with a long stiff seta dorsal to a small hook-like process in Opaepupu gen. nov. vs. without such a seta in Bruceonia; and the first maxilliped with two distinct endites and lacking a palp in Opaepupu gen. nov. vs. with the endites completely fused and with a reduced palp in Bruceonia (cf. Fransen 2002). In addition, Opaepupu gen. nov. can be separated from Bruceonia by the presence of a stout sharp antennal tooth and the infraorbital angle not produced anteriorly vs. with a greatly reduced antennal tooth and infraorbital angle anteriorly protruding in Bruceonia; the posterior margin of telson with two pairs of spiniform setae vs. three in Bruceonia; the distolateral tooth of the scaphocerite greatly reduced vs. well-developed, sharp in Bruceonia; and the dactylus of the ambulatory pereiopods elongate, biunguiculate, with the secondary unguis almost parallel to the main unguis in Opaepupu gen. nov. vs. more robust, biunguiculate, with the secondary unguis being distinctly divergent from the axis of the main unguis, the latter also with micro-ornamentation in Bruceonia (cf. Bruce 1981; Fransen 2002). Opaepupu gen. nov. can also be distinguished from the monotypic Cainonia, for instance, by the general shape and length of the rostrum; the development of the distolateral tooth of the antennal scaphocerite; several features of the mouthparts, including the same distinguishing features on the maxillule, maxilla and first maxilliped used in the comparisons above; and the general shape and armature of the dactylus of the ambulatory pereiopods (cf. Bruce 1980, 2005; Fransen 2002). The single species of this genus, C. medipacifica (Edmondson, 1935), is associated with spiny oysters, Spondylus spp. (Fransen 2002, as Dactylonia medipacifica), morphologically and ecologically very different from the trapezid clam host of the type species of Opaepupu gen. nov. (see below). Pontonia, as redefined by Fransen (2002), presently contains 11 species (Fransen 2002; Marin & Anker 2008), all of them distributed in the Atlantic and East Pacific. All but one species are associated with various bivalves, including spiny and pearl oysters and fan clams (Spondylus, Pinna Linnaeus, 1758, Pinctada R ̂ ding, 1798), the only exception being P. panamica Marin & Anker, 2008, associated with ascidians. Using the diagnosis of Pontonia and species accounts in Fransen (2002), Opaepupu gen. nov. can be distinguished from Pontonia by the slender ambulatory pereiopods, each ending in an elongate biunguiculate dactylus, with the secondary unguis almost parallel to the main unguis vs. the stouter ambulatory pereiopods, each ending in a stout biunguiculate dactylus, with the secondary unguis distinctly divergent from the axis of the main unguis in Pontonia (cf. Fransen 2002). Additional distinguishing features between the two genera are the dorsally carinate rostrum in Opaepupu gen. nov. vs. not carinate in Pontonia; the coxal endite of the maxillule not particularly enlarged vs. more or less expanded, sometimes hypertrophied, in Pontonia; the ventral lobe of the maxillular palp with a long stiff seta dorsal to a small hook-like process vs. typically with a small hook-like process, but never with a stiff seta, in Pontonia; and the first maxilliped without a palp and with the coxal and basial endites separate vs. with a well-developed palp and with the two endites completely fused in Pontonia (cf. Fransen 2002). Furthermore, in most species of Pontonia, the second pereiopods are unequal in size, more or less asymmetrical in shape and very robust, especially in males, contrasting with the subequal and more slender chelipeds in both sexes in the new genus. Opaepupu gen. nov. clearly differs from Conchodytes, currently represented by 10 species, by a series of morphological characters, involving the rostrum, telson, antenna, maxillule, maxilla, first and third maxillipeds, first and second pereiopods, as well as ambulatory pereiopods, especially the strikingly different configurations of the dactylus (cf. Johnson 1967, as Chernocaris; Bruce 1977, 1989, 2005; Marin 2011, as Lacertopontonia; Fransen & Reijnen 2012). Remarkably, members of the genus Conchodytes have been reported from an array of bivalve hosts from eight bivalve families (Fransen & Reijnen 2013 and references therein), indicating a relatively low specificity and tendency to host switching (see also Horká et al. 2016). Finally, Opaepupu gen. nov. can be easily distinguished, using many morphological criteria, from the remaining three genera containing exclusively bivalve symbionts, viz. Anchistus (7 species), Neoanchistus (2 species) and Paranchistus (6 species) (De Grave & Fransen 2011; Fransen & Reijnen 2012). The new genus differs from all of them by the posterior margin of the telson furnished with two pairs of spiniform setae inserted at the same level vs. with three pairs of spiniform setae, the most lateral pair located distinctly above the two other pairs, in Anchistus, Neoanchistus and Paranchistus; the distolateral tooth of the scaphocerite greatly reduced vs. moderately developed to very strong in the other three genera; several features on the mouthparts (e. g., maxillule, first maxilliped, see above); the configuration of the dactyli of the ambulatory pereiopods, which are very different in the other three genera; and the lateral lobe of the uropodal protopod ending bluntly vs. ending in sharp tooth in the other three genera (cf. Kemp 1922; Holthuis 1952; Bruce 1975, 1978). The new genus differs specifically from Neoanchistus by the antennal tooth being strong and sharp in the new genus vs. reduced to a blunt lobe in Neoanchistus and the first pereiopod chela with the finger cutting edges simple, blade-like in the new genus vs. excavated and subspatulate in Neoanchistus (cf. Bruce 1975). It differs specifically from Paranchistus by the rostrum being dorsally non-dentate vs. distally toothed in Paranchistus and the carapace lacking an articulate hepatic tooth, as well as a protruding infraorbital lobe, both of which are present in Paranchistus (cf. Bruce 1978). The members of these three genera associate with a variety of bivalve hosts, both sessile and burrowing, including giant clams of the genus Tridacna Bruguière, 1797 (cf. Bruce 1975, 1978; Fransen 1995; Britayev & Marin 2011; Fransen & Reijnen 2012). In view of numerous host switches reported in the Palaemonidae (e. g., Marin & Anker 2008; Horká et al. 2016) and no morphological feature indicating a potential close phylogenetic proximity of Opaepupu gen. nov. to any of the other bivalve-associated palaemonid genera, it seems quite possible that the new genus may be part of a different lineage otherwise containing taxa not (or not only) associated with bivalves. For instance, the general shape of the second pereiopods (chelipeds) is somewhat reminiscent of several coral-associated genera, such as Philarius Holthuis, 1952 or Harpilius Dana, 1852. Of course, these might be only superficial resemblances, plesiomorphies or distant convergences. Although the status of Opaepupu gen. nov. as a new genus is not in doubt, a cladistic analysis of the entire family is needed to elucidate its phylogenetic position.	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20A6B75FF0C9F9CFDA55BD3.taxon	description	(Figs. 1 – 8)	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20A6B75FF0C9F9CFDA55BD3.taxon	materials_examined	Type material. Holotype: ovigerous female (pocl 3.6 mm, cl 4.3 mm), FLMNH UF 45826, Hawaiian Archipelago, Hawai’i, Kâne’ohe Bay, outside reef at northern side of bay, 21.504 ° - 157.802 °, outer reef slope, depth: 14 m, inside Trapezium oblongum, leg. D. Uyeno and IZ team, 28 May 2017 [fcn BKON 1492 - A]. Allotype: male (pocl 2.3 mm, cl 2.9 mm), FLMNH UF 51717, same collection data as for the holotype [fcn BKON 1492 - B].	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20A6B75FF0C9F9CFDA55BD3.taxon	description	Description. Small-sized palaemonid shrimp with subcylindrical, dorsoventrally somewhat flattened body; female with greatly enlarged brood chamber (Figs. 1, 2 C). Carapace smooth, glabrous, neither densely setose nor pubescent, with few erect setae (Fig. 2 B). Rostrum moderately developed, subtriangular in dorsal view, as long as wide (female) to 1.2 times longer than wide (male), reaching to about 0.7 of distal margin of first article of antennular peduncle, slender in lateral view; mid-dorsal carina present, non-dentate, flattening slightly posterior to level of post-orbital margin; ventral carina non-dentate; lateral carinae proximally expanded; tip slightly descendant in lateral view, with several apical and subapical setae (Figs. 2 A, B, 6 A – C). Supraorbital, hepatic and epigastric teeth absent (Figs. 2 A, B, 6 A). Orbit well developed, infraorbital angle distinct. Antennal tooth well developed, acute (Figs. 2 A, B, 6 A). Pterygostomial angle well developed, protruding, rounded, almost reaching to level of rostral tip (Fig. 2 B). Thoracic sternites without strong teeth, including fourth sternite. Pleon smooth, glabrous; brood chamber of female strongly expanded laterally; posteroventral angle of first five pleura rounded, that of sixth pleonite quadrate, posteriorly unarmed (Figs. 1, 2 D, 6 D, E). Telson broad, distally tapering, about 1.6 and 2.2 times as long as proximal width in female and male, respectively; dorsal surface with wide median depression and two pairs of submarginally inserted, stout cuspidate setae, latter noticeably stouter in male; anterior pair situated at about 0.25 and 0.18 of telson length in female and male, respectively; posterior pair situated at about 0.55 of telson length in both female and male; posterior margin slightly rounded, with two pairs of small, elongate spiniform setae and three (female) or one pair (male) of longer, submedian, sparsely plumose setae; mesial spiniform setae about 1.5 times to twice as long as lateral spiniform setae (Figs. 2 D – F, 6 E, F). Eye peduncle robust; cornea well developed, globular, about 0.6 times as wide as maximal peduncle width; accessory eye (nebenauge) absent (Figs. 2 A, B, 6 A). Antennule robust, short; first article longest and widest, ending in subtriangular, blunt distolateral lobe, without sharp tooth; stylocerite small, distally obtusely triangular; second and third article about 0.35 times as long as visible portion of first article; lateral flagellum with short accessory ramus, comprised of single unit (Figs. 2 A, B, G, 6 A). Antenna with basicerite unarmed; carpocerite reaching to distal end of second article of antennular peduncle; scaphocerite well developed, broad, ovoid, reaching far beyond antennular peduncle; blade about 1.8 times as long as maximal width; distolateral tooth vestigial (Figs. 2 A, B, 6 A). Epistome unarmed. Mandible with incisor and molar processes widely separated; palp absent; incisor process well developed, with six sharp teeth; molar process distally with micro-serration and low bosses (Fig. 3 A, B). Maxillule with both laciniae well developed; palp bilobed distally, ventral lobe more developed, with small, acute, tooth-like ventral extension and long, simple, dorsal seta; dorsal lobe short, unadorned (Fig. 3 C). Maxilla with palp (endopod) well developed, not subdivided; basial endite with deep cleft; coxal endite reduced; scaphognathite well developed (Fig. 3 D). First maxilliped with basial and coxal endites broad, clearly separated; palp (endopod) absent [or at least reduced, not visible in lateral view]; exopod short, with well-developed caridean lobe and moderately long flagellum; epipod well developed, bilobed (Fig. 3 E). Second maxilliped with endopod typical for family; exopod moderately long; epipod well developed, without podobranch (Fig. 3 F). Third maxilliped pediform, moderately stout; lateral plate of coxa rounded; ischiomerus and basis fused into single, broad antepenultimate article; ultimate article about 0.8 times as long as penultimate article, distally tapering; exopod well developed, overreaching distal margin of antepenultimate article; arthrobranch absent (Fig. 3 G). Epistome with median carina; paragnaths rounded, unarmed (Fig. 3 H). First pereiopods symmetrical in shape, equal in size; relatively short, stout; ischium stout, unarmed, somewhat widening distally; merus unarmed, about 6.5 times as long as maximal width; carpus slightly longer than merus, distally widening; chela about 0.75 times as long as carpus; fingers about as long as palm, non-spatulate, with entire, non-dentate cutting edges; fingertips not strongly curved, simple (Fig. 5 A, B). Second pereiopods (chelipeds) of female holotype subequal in size, symmetrical, i. e. similar in shape, not particularly enlarged, moderately setose; all articles with smooth surface; ischium rectangular, about 2.5 times as long as distal width; merus rectangular, slightly longer than ischium, about 2.3 times as wide as long; carpus vase-shaped, about as long as merus; chela not swollen, smooth, without grooves or superficial sculpture; fingers about 0.7 times as long as palm; cutting edge of fingers proximally dentate, distally unarmed (pollex) or with very low tooth and broad convexity (dactylus); proximal tooth of pollex distally truncate, with minute rugosities; distal tooth of pollex subtriangular; dactylus with two subtriangular teeth, proximal much stronger than distal; fingertips strongly crossing (Fig. 4 A – H). Second pereiopods (chelipeds) of male paratype generally similar to those of female, somewhat stouter; fingers slightly gaping between 0.3 and 0.5 of their length; dactylus with very stout, distally truncate and rugose proximal tooth, and much smaller subtriangular distal tooth (Fig. 7 A – E). Third to fifth (ambulatory) pereiopods relatively slender, generally similar in shape and size. Third pereiopod with ischium about 3.5 times as long as wide, unarmed; merus about 1.6 times as long as ischium, about 5.5 times as long as wide, unarmed; carpus more slender than merus, about 0.45 times as long as merus, about 3.5 times as long as wide, unarmed; propodus about twice as long as carpus, about 7.5 times as long as wide, distally somewhat curved, unarmed; dactylus relatively slender, about 0.3 times as long as propodus, compressed, biunguiculate; main (terminal) unguis slender, acute; corpus with single sharp subdistal tooth (= secondary or accessory unguis), latter stouter but shorter than terminal unguis and running almost parallel to it (Fig. 5 C, D). Fourth pereiopod (not illustrated) generally very similar to third. Fifth pereiopod generally similar to third; propodal cleaning brush absent (Fig. 5 E, F). First pleopod sexually dimorphic in proportions, lacking appendix interna in both sexes; in female, endopod short, approximately 0.3 length of exopod (Fig. 2 G); in male, endopod about 0.53 length of exopod, fringed with stiff, slender, subspiniform setae; exopod fringed with plumose setae (Fig. 6 F). Second to fifth pleopods broadly similar with endopod being slightly shorter than exopod, bearing appendix interna (Fig. 2 I). Second pleopod of male with appendix masculina slightly shorter than appendix interna, furnished with stiff, slender, subspiniform setae on apex and subapically (Fig. 6 G). Uropod with lateral lobe of protopod ending bluntly; exopod and endopod subequal, distinctly overreaching telson; exopod broadly ovoid; distolateral tooth reduced; adjacent distolateral spiniform seta minute in female, slightly larger but still small, in male; diaeresis feebly developed, unarmed, visible only in its lateral section in female, visible over its entire length in male, sinuous; endopod narrowly ovoid, without specific features (Figs. 2 D, J, 6 D, E). Colour pattern. Female: Body translucent with numerous rounded to star-shaped groups of white chromatophores, especially numerous on dorsal surface of carapace and pleon, less numerous on tail fan and antennae; chelipeds and walking legs colourless, with some slight iridescence; ovaries and fresh eggs bright yellow (Fig. 8 A). Male: Body translucent with numerous groups of white chromatophores, mostly concentrated on dorsal surface of carapace and pleon, with some isolated white chromatophores on pleon; tail fan, antennae, chelipeds and walking legs mostly colourless, with some slight iridescence (Fig 8 B).	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20A6B75FF0C9F9CFDA55BD3.taxon	etymology	Etymology. The species name (huna) is derived from the Hawaiian word hűnâ, meaning secretive, hidden, in reference to the shrimps’ highly cryptic life style inside bivalves; used as a noun in apposition.	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20A6B75FF0C9F9CFDA55BD3.taxon	materials_examined	Type locality. Kâne’ohe Bay, Hawai’i Island, Hawaiian Islands.	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20A6B75FF0C9F9CFDA55BD3.taxon	distribution	Distribution. Presently known only from the type locality in Hawai’i.	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20A6B75FF0C9F9CFDA55BD3.taxon	biology_ecology	Ecology. The single known pair of the new species was found on a reef slope, inside the mantle cavity of the bivalve Trapezium oblongum (Linnaeus, 1758) (Trapezidae).	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
073A87C8E20A6B75FF0C9F9CFDA55BD3.taxon	discussion	Remarks. See generic account.	en	Anker, Arthur, Grave, Sammy De (2021): Opaepupu, a new genus and species of bivalve-associated shrimp (Decapoda Caridea: Palaemonidae) from Hawai’i. Zootaxa 4903 (1): 55-70, DOI: 10.11646/zootaxa.4903.1.3
