taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650461	https://doi.org/10.3897/jhr.89.79832.figure2	Figure 2. Imagines of the C. cyprius - group in lateral view, proboscis retracted (C. yemenensis female dbM 4944 male dbM 4945, C. osseus female dbM 4617 male dbM 5521, C. ivanovi sp. nov. female dbM 5493 male dbM 5498, C. cagrii sp. nov. female dbM 5607 male dbM 5619, C. clarus female dbM 5674).	Figure 2. Imagines of the C. cyprius - group in lateral view, proboscis retracted (C. yemenensis female dbM 4944 male dbM 4945, C. osseus female dbM 4617 male dbM 5521, C. ivanovi sp. nov. female dbM 5493 male dbM 5498, C. cagrii sp. nov. female dbM 5607 male dbM 5619, C. clarus female dbM 5674).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650463	https://doi.org/10.3897/jhr.89.79832.figure4	Figure 4. Female head in frontal view a C. yemenensis (dbM 5528) b C. clarus (dbM 4616) c C. cyprius (dbM 5431) d C. osseus (dbM 5678) e C. ivanovi sp. nov. (dbM 5490) f C. cagrii sp. nov. (dbM 5609).	Figure 4. Female head in frontal view a C. yemenensis (dbM 5528) b C. clarus (dbM 4616) c C. cyprius (dbM 5431) d C. osseus (dbM 5678) e C. ivanovi sp. nov. (dbM 5490) f C. cagrii sp. nov. (dbM 5609).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650464	https://doi.org/10.3897/jhr.89.79832.figure5	Figure 5. a, b Female clypeus in frontal view a C. cyprius (dbM 5431) b C. ivanovi sp. nov. (dbM 5490) c, d female mandible outside c C. cyprius (dbM 5431) d C. ivanovi sp. nov. (dbM 5490) e, f female ocelli and vertex in dorsal view e C. ivanovi sp. nov. (dbM 5492) f C. cagrii sp. nov. (dbM 5609).	Figure 5. a, b Female clypeus in frontal view a C. cyprius (dbM 5431) b C. ivanovi sp. nov. (dbM 5490) c, d female mandible outside c C. cyprius (dbM 5431) d C. ivanovi sp. nov. (dbM 5490) e, f female ocelli and vertex in dorsal view e C. ivanovi sp. nov. (dbM 5492) f C. cagrii sp. nov. (dbM 5609).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650466	https://doi.org/10.3897/jhr.89.79832.figure7	Figure 7. Right side of postero-lateral part of the propodeum of females in dorso-posterior view a C. yemenensis (dbM 4759) b C. clarus (dbM 4616) c C. cagrii sp. nov. (dbM 5611) d C. rugiceps (dbM 5446) e C. cyprius (dbM 5433) f C. osseus (dbM 5677) g C. ivanovi sp. nov. (dbM 5491) (ps = posterior surface of propodeum fp = postero-medial flange of propodeum pp = postero-lateral process ll = lateral lamella).	Figure 7. Right side of postero-lateral part of the propodeum of females in dorso-posterior view a C. yemenensis (dbM 4759) b C. clarus (dbM 4616) c C. cagrii sp. nov. (dbM 5611) d C. rugiceps (dbM 5446) e C. cyprius (dbM 5433) f C. osseus (dbM 5677) g C. ivanovi sp. nov. (dbM 5491) (ps = posterior surface of propodeum fp = postero-medial flange of propodeum pp = postero-lateral process ll = lateral lamella).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650467	https://doi.org/10.3897/jhr.89.79832.figure8	Figure 8. a-c Female sternum VI in ventral view a C. yemenensis (dbM 4759) b C. ivanovi sp. nov. (dbM 5491) c C. cagrii sp. nov. (dbM 5611) d-f female fore femur in posterior view d C. cyprius (dbM 5431) e C. osseus (dbM 5678) f C. ivanovi sp. nov. (dbM 5493) g-i female sternum VI posterior margin in dorso-posterior view g C. ivanovi sp. nov. (dbM 5493) h C. cagrii sp. nov. (dbM 5607) i C. cagrii sp. nov. (dbM 5618).	Figure 8. a-c Female sternum VI in ventral view a C. yemenensis (dbM 4759) b C. ivanovi sp. nov. (dbM 5491) c C. cagrii sp. nov. (dbM 5611) d-f female fore femur in posterior view d C. cyprius (dbM 5431) e C. osseus (dbM 5678) f C. ivanovi sp. nov. (dbM 5493) g-i female sternum VI posterior margin in dorso-posterior view g C. ivanovi sp. nov. (dbM 5493) h C. cagrii sp. nov. (dbM 5607) i C. cagrii sp. nov. (dbM 5618).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650468	https://doi.org/10.3897/jhr.89.79832.figure9	Figure 9. a, b Male head in frontal view a C. ivanovi sp. nov. (dbM 5498) b C. cagrii sp. nov. (dbM 5619) c-e male tergum VII in dorso-posterior view c C. osseus (dbM 5521) d C. ivanovi sp. nov. (dbM 5498) e C. cagrii sp. nov. (dbM 5619) f, g male antennal club in dorsal view f C. ivanovi sp. nov. (dbM 5499) g C. cagrii sp. nov. (dbM 5619) h, i male head in dorsal view h C. ivanovi sp. nov. (dbM 5498) i C. cagrii sp. nov. (dbM 5619).	Figure 9. a, b Male head in frontal view a C. ivanovi sp. nov. (dbM 5498) b C. cagrii sp. nov. (dbM 5619) c-e male tergum VII in dorso-posterior view c C. osseus (dbM 5521) d C. ivanovi sp. nov. (dbM 5498) e C. cagrii sp. nov. (dbM 5619) f, g male antennal club in dorsal view f C. ivanovi sp. nov. (dbM 5499) g C. cagrii sp. nov. (dbM 5619) h, i male head in dorsal view h C. ivanovi sp. nov. (dbM 5498) i C. cagrii sp. nov. (dbM 5619).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650473	https://doi.org/10.3897/jhr.89.79832.figure14	Figure 14. Male fused sterna VII + VIII in ventral view a C. rugiceps (dbM 5289) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619).	Figure 14. Male fused sterna VII + VIII in ventral view a C. rugiceps (dbM 5289) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650469	https://doi.org/10.3897/jhr.89.79832.figure10	Figure 10. Male genitalia in dorsal view a C. rugiceps (dbM 2834) b C. cyprius (dbM 5452) c C. yemenensis (dbM 5531) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5497) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus at = apodema thyrsos bs = basal sclerite cu = cupula ha = harpide sp = stipes).	Figure 10. Male genitalia in dorsal view a C. rugiceps (dbM 2834) b C. cyprius (dbM 5452) c C. yemenensis (dbM 5531) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5497) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus at = apodema thyrsos bs = basal sclerite cu = cupula ha = harpide sp = stipes).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650470	https://doi.org/10.3897/jhr.89.79832.figure11	Figure 11. Microphotos of male genitalia in dorsal view a C. rugiceps (dbM 5450) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus at = apodema thyrsos bs = basal sclerite cu = cupula ha = harpide sp = stipes).	Figure 11. Microphotos of male genitalia in dorsal view a C. rugiceps (dbM 5450) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus at = apodema thyrsos bs = basal sclerite cu = cupula ha = harpide sp = stipes).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650471	https://doi.org/10.3897/jhr.89.79832.figure12	Figure 12. Male genitalia in ventral view a C. rugiceps (dbM 2834) b C. cyprius (dbM 5452) c C. yemenensis (dbM 5531) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5497) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus bs = basal sclerite cu = cupula ha = harpide sp = stipes vo = volsella ut = uncus thyrsos).	Figure 12. Male genitalia in ventral view a C. rugiceps (dbM 2834) b C. cyprius (dbM 5452) c C. yemenensis (dbM 5531) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5497) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus bs = basal sclerite cu = cupula ha = harpide sp = stipes vo = volsella ut = uncus thyrsos).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650472	https://doi.org/10.3897/jhr.89.79832.figure13	Figure 13. Microphotos of male genitalia in ventral view a C. rugiceps (dbM 5450) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus bs = basal sclerite cu = cupula ha = harpide sp = stipes vo = volsella ut = uncus thyrsos).	Figure 13. Microphotos of male genitalia in ventral view a C. rugiceps (dbM 5450) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus bs = basal sclerite cu = cupula ha = harpide sp = stipes vo = volsella ut = uncus thyrsos).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650475	https://doi.org/10.3897/jhr.89.79832.figure16	Figure 16. Neighbour joining BOLD taxon ID tree of 18 specimens of the C. cyprius - group based on COI- 5 P gene sequences (individually marked by BOLD process ID and assigned to species a priori by morphological characters; COI- 5 P gene sequence of C. osseus and C. clarus unknown).	Figure 16. Neighbour joining BOLD taxon ID tree of 18 specimens of the C. cyprius - group based on COI- 5 P gene sequences (individually marked by BOLD process ID and assigned to species a priori by morphological characters; COI- 5 P gene sequence of C. osseus and C. clarus unknown).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650476	https://doi.org/10.3897/jhr.89.79832.figure17	Figure 17. Geographic distribution of members of the C. cyprius - group (full circles specimens investigated, open circles records from literature; previously published records of C. cyprius from Iran and Armenia excluded; made with Natural Earth, www. naturalearthdata. com).	Figure 17. Geographic distribution of members of the C. cyprius - group (full circles specimens investigated, open circles records from literature; previously published records of C. cyprius from Iran and Armenia excluded; made with Natural Earth, www. naturalearthdata. com).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
1EDA28F79ADF510DA9BF92DA340A4C7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650477	https://doi.org/10.3897/jhr.89.79832.figure18	Figure 18. a-e, g, h Flower visiting behaviour of Celonites ivanovi sp. nov. females at flowers of Heliotropium styligerum and behaviours associated with it (details see text) f habitat of C. ivanovi sp. nov. at locality 2 in the vicinity of Maydanskoye, Dagestan i habitat of C. cagrii sp. nov. near Aksar, Turkey (aspect in autumn).	Figure 18. a-e, g, h Flower visiting behaviour of Celonites ivanovi sp. nov. females at flowers of Heliotropium styligerum and behaviours associated with it (details see text) f habitat of C. ivanovi sp. nov. at locality 2 in the vicinity of Maydanskoye, Dagestan i habitat of C. cagrii sp. nov. near Aksar, Turkey (aspect in autumn).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650464	https://doi.org/10.3897/jhr.89.79832.figure5	Figure 5. a, b Female clypeus in frontal view a C. cyprius (dbM 5431) b C. ivanovi sp. nov. (dbM 5490) c, d female mandible outside c C. cyprius (dbM 5431) d C. ivanovi sp. nov. (dbM 5490) e, f female ocelli and vertex in dorsal view e C. ivanovi sp. nov. (dbM 5492) f C. cagrii sp. nov. (dbM 5609).	Figure 5. a, b Female clypeus in frontal view a C. cyprius (dbM 5431) b C. ivanovi sp. nov. (dbM 5490) c, d female mandible outside c C. cyprius (dbM 5431) d C. ivanovi sp. nov. (dbM 5490) e, f female ocelli and vertex in dorsal view e C. ivanovi sp. nov. (dbM 5492) f C. cagrii sp. nov. (dbM 5609).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650465	https://doi.org/10.3897/jhr.89.79832.figure6	Figure 6. a, b Metasomal tergum II and III of female in dorsal view a C. rugiceps (dbM 5446) b C. ivanovi sp. nov. (dbM 5481) c-f mesonotum of female in dorsal view c C. ivanovi sp. nov. (dbM 5492) d C. osseus (dbM 4617) e C. yemenensis (dbM 5528) f C. clarus (dbM 4616).	Figure 6. a, b Metasomal tergum II and III of female in dorsal view a C. rugiceps (dbM 5446) b C. ivanovi sp. nov. (dbM 5481) c-f mesonotum of female in dorsal view c C. ivanovi sp. nov. (dbM 5492) d C. osseus (dbM 4617) e C. yemenensis (dbM 5528) f C. clarus (dbM 4616).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650461	https://doi.org/10.3897/jhr.89.79832.figure2	Figure 2. Imagines of the C. cyprius - group in lateral view, proboscis retracted (C. yemenensis female dbM 4944 male dbM 4945, C. osseus female dbM 4617 male dbM 5521, C. ivanovi sp. nov. female dbM 5493 male dbM 5498, C. cagrii sp. nov. female dbM 5607 male dbM 5619, C. clarus female dbM 5674).	Figure 2. Imagines of the C. cyprius - group in lateral view, proboscis retracted (C. yemenensis female dbM 4944 male dbM 4945, C. osseus female dbM 4617 male dbM 5521, C. ivanovi sp. nov. female dbM 5493 male dbM 5498, C. cagrii sp. nov. female dbM 5607 male dbM 5619, C. clarus female dbM 5674).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650463	https://doi.org/10.3897/jhr.89.79832.figure4	Figure 4. Female head in frontal view a C. yemenensis (dbM 5528) b C. clarus (dbM 4616) c C. cyprius (dbM 5431) d C. osseus (dbM 5678) e C. ivanovi sp. nov. (dbM 5490) f C. cagrii sp. nov. (dbM 5609).	Figure 4. Female head in frontal view a C. yemenensis (dbM 5528) b C. clarus (dbM 4616) c C. cyprius (dbM 5431) d C. osseus (dbM 5678) e C. ivanovi sp. nov. (dbM 5490) f C. cagrii sp. nov. (dbM 5609).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650466	https://doi.org/10.3897/jhr.89.79832.figure7	Figure 7. Right side of postero-lateral part of the propodeum of females in dorso-posterior view a C. yemenensis (dbM 4759) b C. clarus (dbM 4616) c C. cagrii sp. nov. (dbM 5611) d C. rugiceps (dbM 5446) e C. cyprius (dbM 5433) f C. osseus (dbM 5677) g C. ivanovi sp. nov. (dbM 5491) (ps = posterior surface of propodeum fp = postero-medial flange of propodeum pp = postero-lateral process ll = lateral lamella).	Figure 7. Right side of postero-lateral part of the propodeum of females in dorso-posterior view a C. yemenensis (dbM 4759) b C. clarus (dbM 4616) c C. cagrii sp. nov. (dbM 5611) d C. rugiceps (dbM 5446) e C. cyprius (dbM 5433) f C. osseus (dbM 5677) g C. ivanovi sp. nov. (dbM 5491) (ps = posterior surface of propodeum fp = postero-medial flange of propodeum pp = postero-lateral process ll = lateral lamella).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650467	https://doi.org/10.3897/jhr.89.79832.figure8	Figure 8. a-c Female sternum VI in ventral view a C. yemenensis (dbM 4759) b C. ivanovi sp. nov. (dbM 5491) c C. cagrii sp. nov. (dbM 5611) d-f female fore femur in posterior view d C. cyprius (dbM 5431) e C. osseus (dbM 5678) f C. ivanovi sp. nov. (dbM 5493) g-i female sternum VI posterior margin in dorso-posterior view g C. ivanovi sp. nov. (dbM 5493) h C. cagrii sp. nov. (dbM 5607) i C. cagrii sp. nov. (dbM 5618).	Figure 8. a-c Female sternum VI in ventral view a C. yemenensis (dbM 4759) b C. ivanovi sp. nov. (dbM 5491) c C. cagrii sp. nov. (dbM 5611) d-f female fore femur in posterior view d C. cyprius (dbM 5431) e C. osseus (dbM 5678) f C. ivanovi sp. nov. (dbM 5493) g-i female sternum VI posterior margin in dorso-posterior view g C. ivanovi sp. nov. (dbM 5493) h C. cagrii sp. nov. (dbM 5607) i C. cagrii sp. nov. (dbM 5618).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650468	https://doi.org/10.3897/jhr.89.79832.figure9	Figure 9. a, b Male head in frontal view a C. ivanovi sp. nov. (dbM 5498) b C. cagrii sp. nov. (dbM 5619) c-e male tergum VII in dorso-posterior view c C. osseus (dbM 5521) d C. ivanovi sp. nov. (dbM 5498) e C. cagrii sp. nov. (dbM 5619) f, g male antennal club in dorsal view f C. ivanovi sp. nov. (dbM 5499) g C. cagrii sp. nov. (dbM 5619) h, i male head in dorsal view h C. ivanovi sp. nov. (dbM 5498) i C. cagrii sp. nov. (dbM 5619).	Figure 9. a, b Male head in frontal view a C. ivanovi sp. nov. (dbM 5498) b C. cagrii sp. nov. (dbM 5619) c-e male tergum VII in dorso-posterior view c C. osseus (dbM 5521) d C. ivanovi sp. nov. (dbM 5498) e C. cagrii sp. nov. (dbM 5619) f, g male antennal club in dorsal view f C. ivanovi sp. nov. (dbM 5499) g C. cagrii sp. nov. (dbM 5619) h, i male head in dorsal view h C. ivanovi sp. nov. (dbM 5498) i C. cagrii sp. nov. (dbM 5619).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650473	https://doi.org/10.3897/jhr.89.79832.figure14	Figure 14. Male fused sterna VII + VIII in ventral view a C. rugiceps (dbM 5289) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619).	Figure 14. Male fused sterna VII + VIII in ventral view a C. rugiceps (dbM 5289) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650469	https://doi.org/10.3897/jhr.89.79832.figure10	Figure 10. Male genitalia in dorsal view a C. rugiceps (dbM 2834) b C. cyprius (dbM 5452) c C. yemenensis (dbM 5531) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5497) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus at = apodema thyrsos bs = basal sclerite cu = cupula ha = harpide sp = stipes).	Figure 10. Male genitalia in dorsal view a C. rugiceps (dbM 2834) b C. cyprius (dbM 5452) c C. yemenensis (dbM 5531) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5497) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus at = apodema thyrsos bs = basal sclerite cu = cupula ha = harpide sp = stipes).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650470	https://doi.org/10.3897/jhr.89.79832.figure11	Figure 11. Microphotos of male genitalia in dorsal view a C. rugiceps (dbM 5450) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus at = apodema thyrsos bs = basal sclerite cu = cupula ha = harpide sp = stipes).	Figure 11. Microphotos of male genitalia in dorsal view a C. rugiceps (dbM 5450) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus at = apodema thyrsos bs = basal sclerite cu = cupula ha = harpide sp = stipes).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650471	https://doi.org/10.3897/jhr.89.79832.figure12	Figure 12. Male genitalia in ventral view a C. rugiceps (dbM 2834) b C. cyprius (dbM 5452) c C. yemenensis (dbM 5531) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5497) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus bs = basal sclerite cu = cupula ha = harpide sp = stipes vo = volsella ut = uncus thyrsos).	Figure 12. Male genitalia in ventral view a C. rugiceps (dbM 2834) b C. cyprius (dbM 5452) c C. yemenensis (dbM 5531) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5497) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus bs = basal sclerite cu = cupula ha = harpide sp = stipes vo = volsella ut = uncus thyrsos).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650472	https://doi.org/10.3897/jhr.89.79832.figure13	Figure 13. Microphotos of male genitalia in ventral view a C. rugiceps (dbM 5450) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus bs = basal sclerite cu = cupula ha = harpide sp = stipes vo = volsella ut = uncus thyrsos).	Figure 13. Microphotos of male genitalia in ventral view a C. rugiceps (dbM 5450) b C. cyprius (dbM 5439) c C. yemenensis (dbM 5532) d C. osseus (dbM 5521) e C. ivanovi sp. nov. (dbM 5498) f C. cagrii sp. nov. (dbM 5619) (ad = aedoeagus bs = basal sclerite cu = cupula ha = harpide sp = stipes vo = volsella ut = uncus thyrsos).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650475	https://doi.org/10.3897/jhr.89.79832.figure16	Figure 16. Neighbour joining BOLD taxon ID tree of 18 specimens of the C. cyprius - group based on COI- 5 P gene sequences (individually marked by BOLD process ID and assigned to species a priori by morphological characters; COI- 5 P gene sequence of C. osseus and C. clarus unknown).	Figure 16. Neighbour joining BOLD taxon ID tree of 18 specimens of the C. cyprius - group based on COI- 5 P gene sequences (individually marked by BOLD process ID and assigned to species a priori by morphological characters; COI- 5 P gene sequence of C. osseus and C. clarus unknown).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650476	https://doi.org/10.3897/jhr.89.79832.figure17	Figure 17. Geographic distribution of members of the C. cyprius - group (full circles specimens investigated, open circles records from literature; previously published records of C. cyprius from Iran and Armenia excluded; made with Natural Earth, www. naturalearthdata. com).	Figure 17. Geographic distribution of members of the C. cyprius - group (full circles specimens investigated, open circles records from literature; previously published records of C. cyprius from Iran and Armenia excluded; made with Natural Earth, www. naturalearthdata. com).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
53CD3F8F5F0552E49E0334E83BE1DE57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650477	https://doi.org/10.3897/jhr.89.79832.figure18	Figure 18. a-e, g, h Flower visiting behaviour of Celonites ivanovi sp. nov. females at flowers of Heliotropium styligerum and behaviours associated with it (details see text) f habitat of C. ivanovi sp. nov. at locality 2 in the vicinity of Maydanskoye, Dagestan i habitat of C. cagrii sp. nov. near Aksar, Turkey (aspect in autumn).	Figure 18. a-e, g, h Flower visiting behaviour of Celonites ivanovi sp. nov. females at flowers of Heliotropium styligerum and behaviours associated with it (details see text) f habitat of C. ivanovi sp. nov. at locality 2 in the vicinity of Maydanskoye, Dagestan i habitat of C. cagrii sp. nov. near Aksar, Turkey (aspect in autumn).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
D398D5B0286E584699241B85F1899D01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650462	https://doi.org/10.3897/jhr.89.79832.figure3	Figure 3. Lectotype of Celonites osseus a lateral view b head in frontal view c labels.	Figure 3. Lectotype of Celonites osseus a lateral view b head in frontal view c labels.	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
D398D5B0286E584699241B85F1899D01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/650476	https://doi.org/10.3897/jhr.89.79832.figure17	Figure 17. Geographic distribution of members of the C. cyprius - group (full circles specimens investigated, open circles records from literature; previously published records of C. cyprius from Iran and Armenia excluded; made with Natural Earth, www. naturalearthdata. com).	Figure 17. Geographic distribution of members of the C. cyprius - group (full circles specimens investigated, open circles records from literature; previously published records of C. cyprius from Iran and Armenia excluded; made with Natural Earth, www. naturalearthdata. com).	2022-02-28	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.		Zenodo	biologists	Mauss, Volker;Fateryga, Alexander V.;Yildirim, Erol;Carpenter, James M.			
