taxonID	type	description	language	source
81AC473ECAF1576BA55364283874D1B0.taxon	description	Lineage E Figures 6, 13	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
81AC473ECAF1576BA55364283874D1B0.taxon	diagnosis	Diagnosis. Within the B. blommersae group, distinguished from B. blommersae by calls consisting of a series of whistles (vs. pulsed trills); and from B. vittatus by calls consisting of a series of whistles (vs. series of short clicks), and absence of dorsolateral stripes (vs. presence). Furthermore, distinguished from B. marojezensis by advertisement calls consisting of notes of rather similar duration (vs. clear distinction of short and long notes), and minimum note duration of 158 ms (vs. 15 ms); from B. kirki sp. nov. by advertisement calls consisting of 3 – 6 notes (vs. 9 – 19 notes) and longer note duration (158 – 308 ms vs. 54 – 105 ms), and absence of red color in outer iris area (vs. presence in some specimens); from B. picardi sp. nov. by advertisement calls consisting of notes of rather similar duration (vs. clear distinction of short and long notes), consisting of 3 – 6 notes (vs. 17 – 25 notes), minimum note duration 158 ms (vs. 19 ms), and absence of red color in outer iris area (vs. distinct in many specimens); from B. pikei sp. nov. by advertisement calls consisting of 3 – 6 notes (vs. 25 – 33 notes) of 158 – 308 ms note duration (vs. max. duration of 98 ms); from B. siskoi sp. nov. by advertisement calls consisting of 3 – 6 notes (vs. 7 – 12 notes), with each note strongly frequency-modulated with an initial ascent and final descent of frequency (vs. regularly ascending frequency); and from B. janewayae sp. nov. by advertisement calls with each note strongly frequency-modulated with an initial ascent and final descent of frequency (vs. regularly ascending frequency), and higher dominant frequency (4130 – 4799 vs. 2687 – 3404 Hz). For a distinction from other species of the B. marojezensis complex described herein, see accounts of these new species below.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
81AC473ECAF1576BA55364283874D1B0.taxon	description	Description of the holotype. Adult male, in excellent state of preservation, SVL 25.3 mm, muscle tissue removed from left thigh for molecular analysis. Body moderately slender; head slightly wider than long, as wide as body; snout rounded in dorsal view, sloped to rounded in lateral view; nostrils directed laterally, nearer to eye than to tip of snout; canthus rostralis indistinct, slightly concave in dorsal view, loreal region slightly concave; tympanum very indistinct, estimated TD about 43 % of ED; supratympanic fold poorly recognizable, slightly curved in its anterior and more straight in its posterior half; vomerine odontophores weakly developed, well-separated in two small rounded aggregations, positioned posteromedial to choanae; choanae medium-sized, rounded to ovoid; maxillary teeth present. Tongue ovoid, posteriorly bifid, free. Arms slender, forearms of slightly larger diameter, subarticular tubercles single, round; metacarpal tubercles not recognizable; fingers weakly webbed and with lateral dermal fringes; webbing formula 1 (traces), 2 i (traces), 2 e (traces), 3 i (traces), 3 e (2), 4 (1.5); relative length of fingers 1 <2 <4 <3 (finger 2 distinctly shorter than finger 4); finger discs enlarged, rounded; nuptial pads recognizable as unpigmented swelling on first finger. Hindlimbs slender; tibiotarsal articulation reaching anterior edge of eye when hindlimb is adpressed along body; lateral metatarsalia separated by webbing; inner metatarsal tubercle small, distinct, elongated; no outer metatarsal tubercle; toes broadly webbed; webbing formula 1 (0.25), 2 i (0.75), 2 e (0), 3 i (1.25), 3 e (0), 4 i (1.75), 4 e (1.75), 5 (0.25); relative length of toes 1 <2 <3 <5 <4; toe discs enlarged, rounded. Skin smooth on dorsal surfaces, throat, chest, and ventral surface of thighs, finely granular on belly; cloacal region surrounded by an area of distinct, large, white-colored granules. In preservative, 7 years after collection (Fig. 6), dorsally light reddish brown with a moderately contrasted but incomplete and somewhat discontinuous brown hourglass marking on anterior part of the dorsum, a broad brown transverse bar on the posterior part of the dorsum, and a narrow dark transverse bar between the eyes. Dorsum densely spotted with poorly contrasted small brown spots. Limbs light brown with darker brown crossbands: about 4 poorly marked crossbands on forearm, 5 on shank, 7 – 8 on thigh. Ventrally cream, white on belly and with dark pigment on ventral side of feet. In life (Fig. 13), dorsally cream and all dark dorsal elements poorly contrasted. Iris rather uniformly beige, iris periphery turquoise.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
81AC473ECAF1576BA55364283874D1B0.taxon	etymology	Etymology. Named after the fictional character Captain Jonathan Archer, first portrayed by Scott Bakula in Rick Berman and Brannon Braga’s Star Trek: Enterprise.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
81AC473ECAF1576BA55364283874D1B0.taxon	distribution	Distribution. According to molecular data summarized herein, the species is known from (1) the type locality, Bevitagnono forest, and (2) Andranonafindra forest. The known elevational range of the species spans from 1016 – 1169 m a. s. l.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
261A57EE474F5C9CA570F3EC2C66D28E.taxon	description	Lineage H Figures 6, 14	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
261A57EE474F5C9CA570F3EC2C66D28E.taxon	diagnosis	Diagnosis. Within the B. blommersae group, distinguished from B. vittatus by absence of dorsolateral stripes (vs. presence). Because calls of this species are unknown, a bioacoustic pairwise diagnosis with other species of the group is not possible and therefore, the species can only be distinguished from some of its close relatives (B. archeri sp. nov., B. blommersae, B. janewayae sp. nov., B. siskoi sp. nov.) by molecular diagnostic sites (see Definition above). The species can be distinguished from B. kirki sp. nov. by absence of red color in outer iris area (vs. presence in some specimens), and presence of a lateral transparent area of the integument of tadpoles (vs. absence); from B. picardi sp. nov. by larger body size (male SVL 25.0 – 27.2 vs. 21.3 – 23.2 mm), and absence of red color in outer iris area (vs. distinct in many); from B. pikei sp. nov. by larger body size (male SVL 25.0 – 27.2 vs. 21.4 – 25.0 mm); and from B. marojezensis by absence of rounded patches on the posterior half of the tail musculature of tadpoles (vs. presence).	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
261A57EE474F5C9CA570F3EC2C66D28E.taxon	description	Description of the holotype. Adult male, in excellent state of preservation, SVL 25.5 mm, muscle tissue removed from left thigh for molecular analysis. Body moderately slender; head slightly wider than long, of similar width as body; snout rounded in dorsal view, moderately rounded to sloping in lateral view; nostrils directed laterally, about equidistant between tip of snout and eye; canthus rostralis distinct and concave in dorsal view, loreal region slightly concave; tympanum indistinct, difficult to recognize, somewhat ovoid (higher than wide), TD 43 % of ED; supratympanic fold not recognizable anteriorly and dorsally of tympanum, weakly recognizable and regularly curved posterior of tympanum; vomerine odontophores weakly developed, well-separated in two small rounded aggregations, positioned posteromedial to choanae; choanae medium-sized, rounded; maxillary teeth present. Tongue ovoid, posteriorly bifid, free. Arms slender, forearms of slightly larger diameter, subarticular tubercles single, round; metacarpal tubercles not recognizable; fingers moderately webbed and with lateral dermal fringes; webbing formula 1 (traces), 2 i (traces), 2 e (1), 3 i (2.5), 3 e (1.75), 4 (1.25); relative length of fingers 1 <2 <4 <3 (finger 2 distinctly shorter than finger 4); finger discs enlarged, rounded; nuptial pads recognizable as unpigmented swelling on first finger. Hindlimbs slender; tibiotarsal articulation reaching tip of snout when hindlimb is adpressed along body; lateral metatarsalia separated by webbing; inner metatarsal tubercle small, distinct, elongated; no outer metatarsal tubercle; toes broadly webbed; webbing formula 1 (0.25), 2 i (1), 2 e (0), 3 i (1.25), 3 e (0.25), 4 i (1.75), 4 e (1.75), 5 (0.5); relative length of toes 1 <2 <3 <5 <4; toe discs enlarged, rounded. Skin smooth on dorsal surfaces, throat, chest, and ventral surface of thighs, finely granular on belly; cloacal region surrounded by an area of distinct, large, white-colored granules. In preservative, 13 years after collection (Fig. 6), dorsally light reddish brown with a moderately contrasted but distinct and complete brown hourglass marking on anterior part of the dorsum, a discontinuous broad brown transverse bar on the posterior part of the dorsum, and a narrow dark transverse bar between the eyes. Dorsum densely spotted with poorly contrasted small brown spots which partly fuse to larger markings. Limbs light brown with darker brown crossbands: about 3 – 5 poorly marked crossbands on forearm, 5 on shank, 7 – 8 on thigh. Ventrally cream, white on belly and with some dark pigment on ventral side of feet. Color of holotype in life not recorded.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
261A57EE474F5C9CA570F3EC2C66D28E.taxon	etymology	Etymology. Named after the fictional character Captain Michael Burnham, first portrayed by Sonequa Martin-Green in Bryan Fuller and Alex Kurtman’s Star Trek: Discovery.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
261A57EE474F5C9CA570F3EC2C66D28E.taxon	distribution	Distribution. According to molecular data summarized herein, the species is reliably known from: (1) the type locality, Andrevorevo, and (2) Marojejy at mid-elevation near Camp Marojejia. The elevational range spans between 616 – 1717 m a. s. l.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
D716B4CFC07459C4B65B6D45B97D698B.taxon	description	Lineage B Figures 6, 12	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
D716B4CFC07459C4B65B6D45B97D698B.taxon	diagnosis	Diagnosis. Within the B. blommersae group, distinguished from B. blommersae by calls consisting of a series of whistles (vs. pulsed trills); and from B. vittatus by calls consisting of a series of whistles (vs. series of short clicks), and absence of dorsolateral stripes (vs. presence). Furthermore, distinguished from B. marojezensis by advertisement calls with lower dominant frequency (2687 – 3404 vs. 4118 – 4441 Hz), with notes emitted at longer maximum inter-note intervals (639 vs. 179 ms), and larger body size (male SVL 25.2 – 28.8 vs. 20.0 – 25.7 mm); from B. kirki sp. nov. by advertisement calls with lower dominant frequency (2687 – 3404 Hz vs. 3499 – 5604 Hz), longer note duration (238 – 604 ms vs. 54 – 105 ms), larger male body size (SVL 25.2 – 28.8 vs. 20.0 – 23.4 mm), and absence of red color in outer iris area (vs. presence in some specimens); from B. picardi sp. nov. by advertisement calls with lower dominant frequency (2687 – 3404 Hz vs. 4903 – 5819 Hz), consisting of 3 ‒ 5 notes (vs. 17 ‒ 25 notes), longer note duration (238 – 604 ms vs. 19 – 225 ms), larger body size (male SVL 25.2 – 28.8 vs. 21.3 – 23.2 mm), and absence of red color in outer iris area (vs. distinct in many specimens); from B. pikei sp. nov. by advertisement calls consisting of 3 – 5 notes (vs. 25 – 33 notes) of 238 – 604 ms note duration (vs. max. duration of 98 ms), with lower dominant frequency (2687 – 3404 vs. 5174 – 5507 Hz), and larger body size (male SVL 25.2 – 28.8 vs. 21.4 – 25.0 mm); and from B. siskoi sp. nov. by advertisement calls consisting of 3 – 5 notes (vs. 7 – 12 notes) of 238 – 604 ms note duration (vs. max. duration of 220 ms) and with lower dominant frequency (2687 – 3404 vs. 4688 – 5332 Hz). For a distinction from other species of the B. marojezensis complex described herein, see accounts of these new species below.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
D716B4CFC07459C4B65B6D45B97D698B.taxon	description	Description of the holotype. Adult male, in excellent state of preservation, SVL 27.4 mm, muscle tissue removed from right thigh for molecular analysis. Body moderately slender; head slightly wider than long and slightly wider than body; snout rounded in dorsal and lateral views; nostrils directed laterally, about equidistant between tip of snout and eye; canthus rostralis distinct, concave in dorsal view, loreal region slightly concave; tympanum indistinct, round, TD about 54 % of ED; supratympanic fold not recognizable (traces posterior to tympanum); vomerine odontophores weakly developed, well-separated in two very small rounded aggregations, positioned posteromedial to choanae; choanae medium-sized, rounded; maxillary teeth present. Tongue ovoid, posteriorly bifid, free. Arms slender, forearms of slightly larger diameter, subarticular tubercles single, round; metacarpal tubercles not recognizable; fingers weakly webbed and with lateral dermal fringes; webbing formula 1 (traces), 2 i (traces), 2 e (traces), 3 i (traces), 3 e (2), 4 (1.5); relative length of fingers 1 <2 <4 <3 (finger 2 distinctly shorter than finger 4); finger discs enlarged, rounded; nuptial pads recognizable as unpigmented swelling on first finger. Hindlimbs slender; tibiotarsal articulation reaching between nostril and tip of snout when hindlimb is adpressed along body; lateral metatarsalia separated by webbing; inner metatarsal tubercle small, distinct, elongated; no outer metatarsal tubercle; toes broadly webbed; webbing formula 1 (0), 2 i (0.5), 2 e (0.25), 3 i (1.25), 3 e (0.25), 4 i (1.75), 4 e (1.75), 5 (0.5); relative length of toes 1 <2 <3 <5 <4; toe discs enlarged, rounded. Skin smooth on dorsal surfaces, throat, chest, and ventral surface of thighs, finely granular on belly; cloacal region surrounded by an area of distinct, large, white-colored granules. In preservative, 14 years after collection (Fig. 6), dorsally light brown with an indistinct and poorly contrasted brown hourglass marking on anterior part of the dorsum. No dark transverse bar is visible on the posterior part of the dorsum, but a comparatively wide dark transverse bar is present between the eyes. Dorsum with an irregular pattern of dark brown, black and whitish small spots, many of which are poorly contrasted. Limbs light brown with darker brown crossbands: 3 – 4 on forearm, about 5 on shank, about 5 on thigh. Ventrally cream, white on belly and with dark pigment on ventral side of feet. In life (Fig. 12), similar but the dorsal hourglass pattern almost not recognizable, crossbands on limbs poorly contrasted, but a few black and white dorsal spots well visible and contrasted. Outer iris color yellowish, inner iris color beige, iris periphery turquoise.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
D716B4CFC07459C4B65B6D45B97D698B.taxon	etymology	Etymology. Named after the fictional character Captain Kathryn Janeway, first portrayed by Kate Mulgrew in Rick Berman, Michael Piller, and Jeri Taylor’s Star Trek: Voyager.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
D716B4CFC07459C4B65B6D45B97D698B.taxon	distribution	Distribution. According to molecular data summarized herein, the species is known from (1) the type locality, western side of the Makira Reserve (Angozongahy and Fotsialanana), (2) the eastern side of the Makira Reserve (around “ Simpona Lodge ”), and (3) Betampona Reserve (sites: Sahambendrana, Sahabefoza, Sahaindrana, Vohitsivalana; Rosa et al. 2012). The elevational occurrence of the species spans from 349 m a. s. l. (Sahaindrana site in Betampona; Rosa et al. 2012) to 1067 m a. s. l. (Fotsialanana source, Makira).	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
41D7C9C94CF65656A74782F18476AEC9.taxon	description	Lineage A = Ca 51 Figures 6, 8	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
41D7C9C94CF65656A74782F18476AEC9.taxon	diagnosis	Diagnosis. Within the B. blommersae group, distinguished from B. blommersae by calls consisting of frequency-modulated whistles (vs. pulsed trills); and from B. vittatus by calls consisting of frequency-modulated whistles (vs. series of short clicks), and absence of dorsolateral stripes (vs. presence). Furthermore, distinguished from B. marojezensis by presence of red color in outer iris area in many specimens (vs. absence), and calls consisting of 9 ‒ 19 whistling notes of successively increasing durations of 54 – 105 ms (vs. 7 – 8 notes, with short and long notes distinguishable). For a distinction from other species of the B. marojezensis complex described herein, see accounts of these new species below.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
41D7C9C94CF65656A74782F18476AEC9.taxon	description	Description of the holotype. Adult male, in good state of preservation, SVL 23.4 mm, tongue and right forelimb removed as tissue samples for molecular analysis. Body slender; head wider than long, wider than body; snout rounded in dorsal view, truncate in lateral view; nostrils directed laterally, nearer to tip of snout than to eye; canthus rostralis indistinct, slightly concave in dorsal view, loreal region slightly concave; tympanum indistinct but recognizable, round, TD 50 % of ED; supratympanic fold not recognizable; vomerine odontophores weakly developed, well-separated in two very small rounded aggregations, positioned posteromedial to choanae; choanae medium-sized, rounded; maxillary teeth present. Tongue removed. Arms slender, subarticular tubercles single, round; metacarpal tubercles not recognizable; fingers weakly webbed and with lateral dermal fringes; webbing formula 1 (traces), 2 i (traces), 2 e (traces), 3 i (traces), 3 e (1.75), 4 (1); relative length of fingers 1 <2 <4 <3 (finger 2 distinctly shorter than finger 4); finger discs enlarged, rounded; nuptial pads recognizable as unpigmented swelling on first finger. Hindlimbs slender; tibiotarsal articulation reaching nostril when hindlimb is adpressed along body; lateral metatarsalia separated by webbing; inner metatarsal tubercle small, distinct, elongated; no outer metatarsal tubercle; toes broadly webbed; webbing formula 1 (0.5), 2 i (1), 2 e (0), 3 i (1), 3 e (0.25), 4 i (1.5), 4 e (1.5), 5 (0.25); relative length of toes 1 <2 <3 <5 <4; toe discs enlarged, rounded. Skin smooth on dorsal surfaces, throat, chest, and ventral surface of thighs, finely granular on belly; cloacal region surrounded by an area of distinct, large, white-colored granules. In preservative, 20 years after collection (Fig. 6), dorsally reddish brown with a distinct and strongly contrasted dark brown hourglass-shaped marking on anterior part of the dorsum, and a dark brown transverse bar, somewhat chevron-shaped, on the posterior part of the dorsum. In addition, a few small dark brown and cream spots are scattered across the dorsum. A distinct light beige vertebral stripe runs from snout to cloaca and interrupts the dark markings. Limbs light brown with darker brown crossbands: 4 – 5 on forearm, 6 – 7 on shank, 6 on thigh. Ventrally cream, white on belly, with some dark pigment only on ventral side of feet. Color of holotype in life not recorded.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
41D7C9C94CF65656A74782F18476AEC9.taxon	etymology	Etymology. Named after the fictional character Captain James T. Kirk, first portrayed by William Shatner in Gene Roddenberry’s Star Trek (The Original Series), and also portrayed by Chris Pine in J. J. Abrams’ Star Trek films.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
41D7C9C94CF65656A74782F18476AEC9.taxon	distribution	Distribution. According to the molecular data summarized herein, the species is known from (1) the type locality, Vohiparara, and many other sites in Ranomafana National Park (Ambatolahy, Ambatovory, Fompohonina, Imaloka, Maharira, Mariavaratra, Ranomena, Sahateza, Samahalaotra, Valohoaka, Vatoharanana, Vohimanara), (2) from Antoetra (Farihimazava), based on barcoded specimen MRSN A 2245 (Andreone et al. 2007); and (3) from Vohidrazana in the Andasibe region. The elevational range spans from mid elevations (e. g., 915 m a. s. l. at Ambatolahy) to relatively high elevations (1272 m a. s. l. at Maharira, up to 1420 m a. s. l. at Antoetra).	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
CE46A5926D5453FDA6C4E36E70517C10.taxon	description	Figures 6, 7	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
CE46A5926D5453FDA6C4E36E70517C10.taxon	diagnosis	Diagnosis. Within the B. blommersae group, distinguished from B. blommersae by calls containing multiple frequency-modulated whistles (vs. pulsed trills), and from B. vittatus by calls containing multiple frequency-modulated whistles (vs. series of short clicks), and absence of dorsolateral stripes (vs. presence). For a distinction from other species of the B. marojezensis complex described herein, see accounts of these new species below.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
CE46A5926D5453FDA6C4E36E70517C10.taxon	distribution	Distribution. According to the molecular data summarized herein, the species is known from (1) the type locality, the Marojejy Massif at low elevation (close to “ Camp Mantella ”), (2) a second site at Marojejy (sample THC 302, sequence downloaded from GenBank; collected at 14.4467 ° S, 49.8251 ° E, 225 m a. s. l. by T. R. Fulgence), (3) the Masoala Peninsula near the Eco-Lodge “ Chez Arol ”, and (4) the Makira Reserve, at Analanjirofo near Simpona Lodge. Boophis marojezensis is a low-elevation species, known from 21 – 410 m a. s. l.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
D197623F8F915E3B95F7D13786C35173.taxon	description	Lineage D Figures 6, 9	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
D197623F8F915E3B95F7D13786C35173.taxon	diagnosis	Diagnosis. Within the B. blommersae group, distinguished from B. blommersae by calls mainly consisting of frequency-modulated whistles (vs. pulsed trills); and from B. vittatus by calls mainly consisting of frequency-modulated whistles (vs. series of short clicks), and absence of dorsolateral stripes (vs. presence). Furthermore, distinguished from B. marojezensis by presence of red color in outer iris area in most specimens (vs. absence), and advertisement calls consisting of 17 – 25 notes (vs. 7 – 8 notes), and from B. kirki sp. nov. by advertisement calls consisting of two types of (short and long) notes (vs. notes of successively increasing duration). For a distinction from other species of the B. marojezensis complex described herein, see accounts of these new species below.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
D197623F8F915E3B95F7D13786C35173.taxon	description	Description of the holotype. Adult male, in good state of preservation, SVL 23.0 mm, tissue from right thigh removed as tissue samples for molecular analysis and posterior venter cut open for parasitological examination. Body slender; head wider than long, wider than body; snout rounded in dorsal view, truncate in lateral view; nostrils directed laterally, about equidistant to tip of snout and eye; canthus rostralis indistinct, slightly concave in dorsal view, loreal region slightly concave; tympanum indistinct but recognizable, round, TD about 44 % of ED; supratympanic fold very indistinct, largely straight; vomerine odontophores weakly developed, well-separated in two very small rounded aggregations, positioned posteromedial to choanae; choanae medium-sized, rounded; maxillary teeth present. Tongue ovoid, posteriorly bifid, free. Arms slender, forearms of slightly larger diameter, subarticular tubercles single, round; metacarpal tubercles not recognizable; fingers weakly webbed and with lateral dermal fringes; webbing formula 1 (traces), 2 i (traces), 2 e (traces), 3 i (traces), 3 e (2), 4 (1.5); relative length of fingers 1 <2 <4 <3 (finger 2 distinctly shorter than finger 4); finger discs enlarged, rounded; nuptial pads indistinct, recognizable as unpigmented weak swelling on first finger. Hindlimbs slender; tibiotarsal articulation reaching nostril when hindlimb is adpressed along body; lateral metatarsalia separated by webbing; inner metatarsal tubercle small, distinct, elongated; no outer metatarsal tubercle; toes broadly webbed; webbing formula 1 (0.5), 2 i (0.75), 2 e (0.25), 3 i (1), 3 e (0), 4 i (1.75), 4 e (1.5), 5 (0.5); relative length of toes 1 <2 <3 <5 <4; toe discs enlarged, rounded. Skin smooth on dorsal surfaces, throat, chest, and ventral surface of thighs, finely granular on belly; cloacal region surrounded by an area of distinct, large, white-colored granules. In preservative, 17 years after collection (Fig. 6), dorsally reddish brown with a distinct and moderately contrasted dark brown hourglass marking on anterior part of the dorsum, and a dark brown transverse bar on the posterior part of the dorsum. Many dark spots of different sizes are scattered across the dorsum. Limbs light brown with a few rather poorly contrasted darker brown crossbands: 1 – 2 on forearm, 2 – 3 on shank, 2 – 3 on thigh. Ventrally cream, white on belly, with some dark pigment only on ventral side of feet. Color of holotype in life not recorded.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
D197623F8F915E3B95F7D13786C35173.taxon	etymology	Etymology. Named after the fictional character Captain Jean-Luc Picard, first portrayed by Sir Patrick Stewart in Gene Roddenberry’s Star Trek: The Next Generation, and later in Akiva Goldsman, Michael Chabon, Kirsten Beyer, and Alex Kurtzman’s Star Trek: Picard.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
D197623F8F915E3B95F7D13786C35173.taxon	distribution	Distribution. According to the molecular data summarized herein, the species is known from several sites in the wider area around the village of Andasibe, i. e., (1) the type locality, An’Ala, (2) Vohidrazana, (3) Vohimana, and it also has been recorded from (4) Betampona (Sahambendrana and Sahabefoza sites, according to Rosa et al. 2012). The elevational range spans from 349 (Betampona, Sahabefoza) to 880 m a. s. l. (An’Ala).	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
01E0667E424F50BF8CF726ED5DC5E3F3.taxon	description	Lineage C Figures 6, 10	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
01E0667E424F50BF8CF726ED5DC5E3F3.taxon	diagnosis	Diagnosis. Within the B. blommersae group, distinguished from B. blommersae by calls consisting of a fast series of short whistles (vs. pulsed trills); and from B. vittatus by calls consisting of a fast series of short whistles (vs. series of short clicks), and absence of dorsolateral stripes (vs. presence). Furthermore, distinguished from B. marojezensis by advertisement calls consisting of 25 – 33 notes (vs. 7 – 8 notes); from B. kirki sp. nov. by advertisement calls consisting of 25 – 33 notes (vs. 9 – 19 notes) and absence of red color in outer iris area (vs. presence in some specimens); and from B. picardi sp. nov. by advertisement calls consisting of 25 – 33 notes (vs. 17 – 25 notes) of maximum duration of 98 ms (vs. max. duration of 225 ms), and absence of red color in outer iris area (vs. distinct in many specimens). For a distinction from other species of the B. marojezensis complex described herein, see accounts of these new species below.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
01E0667E424F50BF8CF726ED5DC5E3F3.taxon	description	Description of the holotype. Adult male, in excellent state of preservation, SVL 22.3 mm, muscle tissue removed from right thigh for molecular analysis. Body slender; head slightly wider than long, much wider than body; snout rounded in dorsal view and rounded to slightly sloped in lateral view; nostrils directed laterally, equidistant to eye and tip of snout; canthus rostralis indistinct, slightly concave in dorsal view, loreal region slightly concave; tympanum indistinct but recognizable, round, TD 50 % of ED; supratympanic fold poorly recognizable, mostly straight; vomerine odontophores weakly developed, well-separated in two very small rounded aggregations, positioned posteromedial to choanae; choanae medium-sized, rounded; maxillary teeth present. Tongue ovoid, posteriorly bifid, free. Arms slender, forearms of slightly larger diameter, subarticular tubercles single, round; metacarpal tubercles not recognizable; fingers weakly webbed and with lateral dermal fringes; webbing formula 1 (traces), 2 i (traces), 2 e (traces), 3 i (traces), 3 e (2), 4 (1.25); relative length of fingers 1 <2 <4 <3 (finger 2 distinctly shorter than finger 4); finger discs enlarged, rounded; nuptial pads recognizable as unpigmented swelling on first finger. Hindlimbs slender; tibiotarsal articulation reaching beyond tip of snout when hindlimb is adpressed along body; lateral metatarsalia separated by webbing; inner metatarsal tubercle small, distinct, elongated; no outer metatarsal tubercle; toes broadly webbed; webbing formula 1 (1), 2 i (1.5), 2 e (0.5), 3 i (1.75), 3 e (0.5), 4 i (1.75), 4 e (1.75), 5 (1); relative length of toes 1 <2 <3 ≤ 5 <4; toe discs enlarged, rounded. Skin smooth on dorsal surfaces, throat, chest, and ventral surface of thighs, finely granular on belly; cloacal region surrounded by an area of distinct, large, white-colored granules. In preservative, 7 years after collection (Fig. 6), dorsally brown with a distinct and moderately contrasted dark brown hourglass marking on anterior part of the dorsum, and a dark brown chevron-like transverse marking on the posterior part of the dorsum. An indistinct narrow dark transverse bar is visible between the eyes. Very few poorly contrasted light spots are scattered across the dorsum. Limbs light brown with distinct darker brown crossbands: about 3 on forearm, 3 – 5 on shank, 5 on thigh. Ventrally cream, white on belly, with dark pigment on ventral side of feet. In life (Fig. 10), similar but overall lighter and dorsal pattern less contrasted. Iris yellowish to beige, iris periphery turquoise.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
01E0667E424F50BF8CF726ED5DC5E3F3.taxon	etymology	Etymology. Named after the fictional character Captain Christopher Pike, first portrayed by Sean Kenney and Jeffrey Hunter in Gene Roddenberry’s Star Trek (The Original Series), and later portrayed by Anson Mount in Akiva Goldsman, Alex Kurtzman, and Jenny Lumet’s Star Trek: Strange New Worlds.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
01E0667E424F50BF8CF726ED5DC5E3F3.taxon	distribution	Distribution. According to molecular data summarized herein, the species is known from (1) the type locality, higher elevations at the Marojejy Massif (around Camp Simpona), and (2) the Sorata Massif. Based on tadpoles, it also has been recorded (3) on the Tsaratanana Massif at a site called Antevialambazaha (14.1743 ° S, 48.9452 ° E, 1699 m a. s. l.), by Randrianiaina et al. (2012). The elevational range of the species spans between 1325 – 1699 m a. s. l.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
687EFDBF5FE95286A63413D560D70E96.taxon	description	Lineage G Figures 6, 11	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
687EFDBF5FE95286A63413D560D70E96.taxon	diagnosis	Diagnosis. Within the B. blommersae group, distinguished from B. blommersae by calls consisting of a series of whistles (vs. pulsed trills); and from B. vittatus by calls consisting of a series of whistles (vs. series of short clicks), and absence of dorsolateral stripes (vs. presence). Furthermore, distinguished from B. marojezensis by somewhat larger body size (male SVL 25.0 – 27.2 vs. 20.0 – 25.7 mm), advertisement calls consisting of notes of rather similar duration (vs. clear distinction of short and long notes); from B. kirki sp. nov. by larger body size (male SVL 25.0 – 27.2 vs. 20.0 – 23.4 mm), advertisement calls with notes of maximum duration of 220 ms (vs. max. duration of 105 ms) and with regularly ascending frequency modulation (vs. very steep initial frequency ascent in the beginning of each note, slowing down towards end of note), and absence of red color in outer iris area (vs. presence in some specimens); from B. picardi sp. nov. by larger body size (male SVL 25.0 – 27.2 vs. 21.3 – 23.2 mm), advertisement calls consisting of notes of rather similar duration (vs. clear distinction of short and long notes), and absence of red color in outer iris area (vs. distinct in many specimens); and from B. pikei sp. nov. by larger body size (male SVL 25.0 – 27.2 vs. 21.4 – 25.0 mm), and advertisement calls consisting of 7 – 12 notes (vs. 25 – 33 notes) of maximum duration of 220 ms (vs. max. duration of 98 ms). For a distinction from other species of the B. marojezensis complex described herein, see accounts of these new species below.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
687EFDBF5FE95286A63413D560D70E96.taxon	description	Description of the holotype. Adult male, in excellent state of preservation, SVL 25.8 mm, muscle tissue removed from left thigh for molecular analysis. Body moderately slender; head slightly wider than long and slightly wider than body; snout rounded to truncate in dorsal view, moderately rounded to sloping in lateral view; nostrils directed laterally, nearer to tip of snout than to eye; canthus rostralis rather weakly expressed, concave in dorsal view, loreal region slightly concave; tympanum rather distinct, round, TD 39 % of ED; supratympanic fold distinct, slightly curved in its anterior and straight in its posterior half; vomerine odontophores weakly developed, well-separated in two very small rounded aggregations, positioned posteromedial to choanae; choanae small to medium-sized, rounded; maxillary teeth present. Tongue ovoid, posteriorly bifid, free. Arms slender, forearms of slightly larger diameter, subarticular tubercles single, round; metacarpal tubercles not recognizable; fingers weakly webbed and with lateral dermal fringes; webbing formula 1 (traces), 2 i (traces), 2 e (traces), 3 i (traces), 3 e (2), 4 (1); relative length of fingers 1 <2 <4 <3 (finger 2 distinctly shorter than finger 4); finger discs enlarged, rounded; nuptial pads recognizable as unpigmented swelling on first finger. Hindlimbs slender; tibiotarsal articulation reaching beyond tip of snout when hindlimb is adpressed along body; lateral metatarsalia separated by webbing; inner metatarsal tubercle small, distinct, elongated; no outer metatarsal tubercle; toes broadly webbed; webbing formula 1 (0.5), 2 i (0.75), 2 e (0.5), 3 i (1.25), 3 e (0.5), 4 i (2), 4 e (2), 5 (0.5); relative length of toes 1 <2 <3 <5 <4; toe discs enlarged, rounded. Skin smooth on dorsal surfaces, throat, chest, and ventral surface of thighs, finely granular on belly; cloacal region surrounded by an area of distinct, large, white-colored granules. In preservative, 22 years after collection (Fig. 6), dorsally red brown with a distinct and moderately contrasted dark brown hourglass marking on anterior part of the dorsum, and a dark brown curved transverse bar on the posterior part of the dorsum. A narrow dark transverse bar is visible between the eyes. Dorsum with many poorly delimited and indistinct small dark spots. Limbs light brown with distinct darker brown crossbands: about 3 on forearm, 3 – 5 on shank, 5 on thigh. Ventrally cream, white on belly and with dark pigment on ventral side of feet. In life (Fig. 11), similar but dorsal color light brown rather than reddish brown. Outer iris color yellowish, inner iris color light brown, iris periphery turquoise.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
687EFDBF5FE95286A63413D560D70E96.taxon	etymology	Etymology. Named after the fictional character Captain Benjamin Sisko, first portrayed by Avery Brooks in Rick Berman and Michael Piller’s Star Trek: Deep Space Nine.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
687EFDBF5FE95286A63413D560D70E96.taxon	distribution	Distribution. According to molecular data summarized herein, the species is known from (1) the type locality, Antsahamanara campsite on the western versant of the Tsaratanana Massif, (2) Ambinanitelo forest (14.2254 ° S, 48.9635 ° E, 1182 m a. s. l.; Randrianiaina et al. 2012), and (3) Ampotsidy forest. The elevational range of the species spans between ca. 1000 – 1320 m a. s. l.	en	Vences, Miguel, Köhler, Jörn, Hutter, Carl R., Preick, Michaela, Petzold, Alice, Rakotoarison, Andolalao, Ratsoavina, Fanomezana M., Glaw, Frank, Scherz, Mark D. (2024): Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643-681, DOI: 10.3897/vz.74.e121110
