taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
6870E7405A5651009774107EC7CA6B44.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439182	https://doi.org/10.3897/zookeys.955.51983.plate16	Plate 16. 1 - 6 Caucasogeyeria chrysomallos sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring 1 holotype 2 - 6 paratypes 7, 8 Caucasogeyeria chrysomallos sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring 9 Caucasogeyeria cf. gloeri, Samegrelo, Mukhuri, Mapeli Cave, paratype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 16. 1 - 6 Caucasogeyeria chrysomallos sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring 1 holotype 2 - 6 paratypes 7, 8 Caucasogeyeria chrysomallos sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring 9 Caucasogeyeria cf. gloeri, Samegrelo, Mukhuri, Mapeli Cave, paratype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
6870E7405A5651009774107EC7CA6B44.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439181	https://doi.org/10.3897/zookeys.955.51983.figure14	Figure 14. A-C Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave D C. chrysomallos sp. nov., Kanti, Mapeli Spring, morphology of the penis. Photograph A. Falniowski and A. Rysiewska.	Figure 14. A-C Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave D C. chrysomallos sp. nov., Kanti, Mapeli Spring, morphology of the penis. Photograph A. Falniowski and A. Rysiewska.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
1F98E04660265385A465B8ED3A38FFBF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439180	https://doi.org/10.3897/zookeys.955.51983.plate15	Plate 15. 1 Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 2 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, Motena Cave, holotype 3 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, spring in village, holotype 4 Caucasogeyeria colchis sp. nov., Imereti, Nakhriduri 2 left side spring in Turchu Gamosadivari Basin above small ford, paratype 5 - 8 Caucasogeyeria pseudocolchis sp. nov., Samegrelo, Mukhuri, Shisha Spring 5 holotype 6 - 8 paratypes. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 15. 1 Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 2 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, Motena Cave, holotype 3 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, spring in village, holotype 4 Caucasogeyeria colchis sp. nov., Imereti, Nakhriduri 2 left side spring in Turchu Gamosadivari Basin above small ford, paratype 5 - 8 Caucasogeyeria pseudocolchis sp. nov., Samegrelo, Mukhuri, Shisha Spring 5 holotype 6 - 8 paratypes. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
1F98E04660265385A465B8ED3A38FFBF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439176	https://doi.org/10.3897/zookeys.955.51983.plate12	Plate 12. Caucasogeyeria specimens used for molecular and anatomical studies 1 C. chrysomallos sp. nov. 2 - 4 C. ignidona sp. nov. 5 - 8 C. colchis sp. nov., live specimens Pirveli Balda, spring in the village 9 - 12 C. chrysomallos sp. nov. The numbers correspond to individuals. Photograph A. Falniowski, J. Grego, A. Rysiewska.	Plate 12. Caucasogeyeria specimens used for molecular and anatomical studies 1 C. chrysomallos sp. nov. 2 - 4 C. ignidona sp. nov. 5 - 8 C. colchis sp. nov., live specimens Pirveli Balda, spring in the village 9 - 12 C. chrysomallos sp. nov. The numbers correspond to individuals. Photograph A. Falniowski, J. Grego, A. Rysiewska.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
9261A6B3F3C551128E975D055C59F644.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439178	https://doi.org/10.3897/zookeys.955.51983.plate13	Plate 13. 1 - 7 Caucasogeyeria gloeri sp. nov. 1, 2 Imereti, Satsiskvilo, Turchusmtha 1 holotype 2 paratype 4 Imereti, Nakhriduri, Spring at Turchu Gamosadivari Basin 5 Imereti, Turchusmtha, spring of Okatse and cave above Kinchkha waterfall 6 Imereti, Nakhriduri, Turchu Gamosadivari Cave Spring 7 Imereti, Nakhriduri, left tributary spring in Turchu Gamosadivari Basin 3 Caucasogeyeria cf. gloeri 2, Samegrelo, Shurubumu, spring on left bank of Khobistskali River. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 13. 1 - 7 Caucasogeyeria gloeri sp. nov. 1, 2 Imereti, Satsiskvilo, Turchusmtha 1 holotype 2 paratype 4 Imereti, Nakhriduri, Spring at Turchu Gamosadivari Basin 5 Imereti, Turchusmtha, spring of Okatse and cave above Kinchkha waterfall 6 Imereti, Nakhriduri, Turchu Gamosadivari Cave Spring 7 Imereti, Nakhriduri, left tributary spring in Turchu Gamosadivari Basin 3 Caucasogeyeria cf. gloeri 2, Samegrelo, Shurubumu, spring on left bank of Khobistskali River. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
A902B762A52C56AD92E2262D1D0542A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439178	https://doi.org/10.3897/zookeys.955.51983.plate13	Plate 13. 1 - 7 Caucasogeyeria gloeri sp. nov. 1, 2 Imereti, Satsiskvilo, Turchusmtha 1 holotype 2 paratype 4 Imereti, Nakhriduri, Spring at Turchu Gamosadivari Basin 5 Imereti, Turchusmtha, spring of Okatse and cave above Kinchkha waterfall 6 Imereti, Nakhriduri, Turchu Gamosadivari Cave Spring 7 Imereti, Nakhriduri, left tributary spring in Turchu Gamosadivari Basin 3 Caucasogeyeria cf. gloeri 2, Samegrelo, Shurubumu, spring on left bank of Khobistskali River. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 13. 1 - 7 Caucasogeyeria gloeri sp. nov. 1, 2 Imereti, Satsiskvilo, Turchusmtha 1 holotype 2 paratype 4 Imereti, Nakhriduri, Spring at Turchu Gamosadivari Basin 5 Imereti, Turchusmtha, spring of Okatse and cave above Kinchkha waterfall 6 Imereti, Nakhriduri, Turchu Gamosadivari Cave Spring 7 Imereti, Nakhriduri, left tributary spring in Turchu Gamosadivari Basin 3 Caucasogeyeria cf. gloeri 2, Samegrelo, Shurubumu, spring on left bank of Khobistskali River. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
901A57E20B8D5DD194CC7BA4E80B9BE9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439177	https://doi.org/10.3897/zookeys.955.51983.figure13	Figure 13. Distribution map of Caucasogeyeria gen. nov. 0 C. valvataeformis (Starobogatov, 1962) (magenta dot) 4. C. horatiaeformis (Starobogatov, 1962) (turquoise dot) 16 - 22 C. gloeri sp. nov. (blue dots) 27, 28 C. cf. gloeri (light blue dots) 7 C. ignidona sp. nov. (purple dot) 20, 22, 23 C. colchis sp. nov. (red dots) 26, 28 C. pseudocolchis sp. nov. (black dots) 27, 29 C. chrysomallos sp. nov. (green dots).	Figure 13. Distribution map of Caucasogeyeria gen. nov. 0 C. valvataeformis (Starobogatov, 1962) (magenta dot) 4. C. horatiaeformis (Starobogatov, 1962) (turquoise dot) 16 - 22 C. gloeri sp. nov. (blue dots) 27, 28 C. cf. gloeri (light blue dots) 7 C. ignidona sp. nov. (purple dot) 20, 22, 23 C. colchis sp. nov. (red dots) 26, 28 C. pseudocolchis sp. nov. (black dots) 27, 29 C. chrysomallos sp. nov. (green dots).	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
16F422A81C675FFF9B361C753D667574.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439180	https://doi.org/10.3897/zookeys.955.51983.plate15	Plate 15. 1 Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 2 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, Motena Cave, holotype 3 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, spring in village, holotype 4 Caucasogeyeria colchis sp. nov., Imereti, Nakhriduri 2 left side spring in Turchu Gamosadivari Basin above small ford, paratype 5 - 8 Caucasogeyeria pseudocolchis sp. nov., Samegrelo, Mukhuri, Shisha Spring 5 holotype 6 - 8 paratypes. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 15. 1 Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 2 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, Motena Cave, holotype 3 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, spring in village, holotype 4 Caucasogeyeria colchis sp. nov., Imereti, Nakhriduri 2 left side spring in Turchu Gamosadivari Basin above small ford, paratype 5 - 8 Caucasogeyeria pseudocolchis sp. nov., Samegrelo, Mukhuri, Shisha Spring 5 holotype 6 - 8 paratypes. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
16F422A81C675FFF9B361C753D667574.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439181	https://doi.org/10.3897/zookeys.955.51983.figure14	Figure 14. A-C Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave D C. chrysomallos sp. nov., Kanti, Mapeli Spring, morphology of the penis. Photograph A. Falniowski and A. Rysiewska.	Figure 14. A-C Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave D C. chrysomallos sp. nov., Kanti, Mapeli Spring, morphology of the penis. Photograph A. Falniowski and A. Rysiewska.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
2939812818C65456B1045F97E563E178.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439180	https://doi.org/10.3897/zookeys.955.51983.plate15	Plate 15. 1 Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 2 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, Motena Cave, holotype 3 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, spring in village, holotype 4 Caucasogeyeria colchis sp. nov., Imereti, Nakhriduri 2 left side spring in Turchu Gamosadivari Basin above small ford, paratype 5 - 8 Caucasogeyeria pseudocolchis sp. nov., Samegrelo, Mukhuri, Shisha Spring 5 holotype 6 - 8 paratypes. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 15. 1 Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 2 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, Motena Cave, holotype 3 Caucasogeyeria colchis sp. nov., Samegrelo, Pirveli Balda, spring in village, holotype 4 Caucasogeyeria colchis sp. nov., Imereti, Nakhriduri 2 left side spring in Turchu Gamosadivari Basin above small ford, paratype 5 - 8 Caucasogeyeria pseudocolchis sp. nov., Samegrelo, Mukhuri, Shisha Spring 5 holotype 6 - 8 paratypes. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
9BAD2D702C5E5B47A977481EA43A770E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439172	https://doi.org/10.3897/zookeys.955.51983.plate9	Plate 9. 1 Caucasopsis olsavskyi sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype 2 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype 3 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Kachara Cave 4 C. egrisi sp. nov., Samegrelo, Pirveli Balda, Motena Cave form 5, 6 C. egrisi sp. nov., Samegrelo, Pirveli Balda, spring in village: 5 holotype 6 paratype 7 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype 9 I. nakeralaensis sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype 10 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 9. 1 Caucasopsis olsavskyi sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype 2 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype 3 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Kachara Cave 4 C. egrisi sp. nov., Samegrelo, Pirveli Balda, Motena Cave form 5, 6 C. egrisi sp. nov., Samegrelo, Pirveli Balda, spring in village: 5 holotype 6 paratype 7 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype 9 I. nakeralaensis sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype 10 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
65E19D736DF65BBC8652527192705777.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439171	https://doi.org/10.3897/zookeys.955.51983.figure11	Figure 11. Morphology of the penis A, B Caucasopsis letsurtsume sp. nov., Samegrelo: A ventrally B dorsally C Imeretiopsis gorgoleti sp. nov. Photograph A. Falniowski and A. Rysiewska.	Figure 11. Morphology of the penis A, B Caucasopsis letsurtsume sp. nov., Samegrelo: A ventrally B dorsally C Imeretiopsis gorgoleti sp. nov. Photograph A. Falniowski and A. Rysiewska.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
65E19D736DF65BBC8652527192705777.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439167	https://doi.org/10.3897/zookeys.955.51983.figure10	Figure 10. Distribution map of Imeretiopsis gen. nov.: (magenta dots) and Caucasopsis gen. nov. (blue dots) 6 I. cameroni sp. nov. and I. iazoni sp. nov. 7 Imeretiopsis prometheus sp. nov. 8 I. nakeralaensis sp. nov. 9 I. gorgoleti sp. nov. 0 Caucasopsis orientalis (Starobogatov, 1962), C. subovata (Starobogatov, 1962) and C. pulcherrima (Starobogatov, 1962) 3 C. shadini (Starobogatov, 1962) 4 C. aculeus (Starobogatov, 1962) and C. schakuranica (Starobogatov, 1962) 22, 23 C. egrisi sp. nov. 24 C. olsavskyi sp. nov. 25, 30, 31 C. letsurtsume sp. nov.	Figure 10. Distribution map of Imeretiopsis gen. nov.: (magenta dots) and Caucasopsis gen. nov. (blue dots) 6 I. cameroni sp. nov. and I. iazoni sp. nov. 7 Imeretiopsis prometheus sp. nov. 8 I. nakeralaensis sp. nov. 9 I. gorgoleti sp. nov. 0 Caucasopsis orientalis (Starobogatov, 1962), C. subovata (Starobogatov, 1962) and C. pulcherrima (Starobogatov, 1962) 3 C. shadini (Starobogatov, 1962) 4 C. aculeus (Starobogatov, 1962) and C. schakuranica (Starobogatov, 1962) 22, 23 C. egrisi sp. nov. 24 C. olsavskyi sp. nov. 25, 30, 31 C. letsurtsume sp. nov.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
A0611C73417855B1B421BABB742B6350.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439172	https://doi.org/10.3897/zookeys.955.51983.plate9	Plate 9. 1 Caucasopsis olsavskyi sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype 2 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype 3 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Kachara Cave 4 C. egrisi sp. nov., Samegrelo, Pirveli Balda, Motena Cave form 5, 6 C. egrisi sp. nov., Samegrelo, Pirveli Balda, spring in village: 5 holotype 6 paratype 7 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype 9 I. nakeralaensis sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype 10 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 9. 1 Caucasopsis olsavskyi sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype 2 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype 3 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Kachara Cave 4 C. egrisi sp. nov., Samegrelo, Pirveli Balda, Motena Cave form 5, 6 C. egrisi sp. nov., Samegrelo, Pirveli Balda, spring in village: 5 holotype 6 paratype 7 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype 9 I. nakeralaensis sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype 10 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
A0611C73417855B1B421BABB742B6350.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439171	https://doi.org/10.3897/zookeys.955.51983.figure11	Figure 11. Morphology of the penis A, B Caucasopsis letsurtsume sp. nov., Samegrelo: A ventrally B dorsally C Imeretiopsis gorgoleti sp. nov. Photograph A. Falniowski and A. Rysiewska.	Figure 11. Morphology of the penis A, B Caucasopsis letsurtsume sp. nov., Samegrelo: A ventrally B dorsally C Imeretiopsis gorgoleti sp. nov. Photograph A. Falniowski and A. Rysiewska.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
A0611C73417855B1B421BABB742B6350.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439175	https://doi.org/10.3897/zookeys.955.51983.figure12	Figure 12. A, B Caucasopsis letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, specimens used for molecular studies. Photograph A. Falniowski.	Figure 12. A, B Caucasopsis letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, specimens used for molecular studies. Photograph A. Falniowski.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
A0611C73417855B1B421BABB742B6350.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439157	https://doi.org/10.3897/zookeys.955.51983.figure5	Figure 5. Maximum Likelihood tree inferred from mitochondrial COI. Bootstrap supports and Bayesian probabilities are given. Newly obtained sequences in bold.	Figure 5. Maximum Likelihood tree inferred from mitochondrial COI. Bootstrap supports and Bayesian probabilities are given. Newly obtained sequences in bold.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
A0611C73417855B1B421BABB742B6350.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439159	https://doi.org/10.3897/zookeys.955.51983.figure7	Figure 7. Maximum Likelihood tree inferred from COI + H 3. Bootstrap supports and Bayesian probabilities are given. Red and green bars indicate results from the ABGD and PTP methods, respectively. Newly obtained sequences in bold.	Figure 7. Maximum Likelihood tree inferred from COI + H 3. Bootstrap supports and Bayesian probabilities are given. Red and green bars indicate results from the ABGD and PTP methods, respectively. Newly obtained sequences in bold.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
204FCF065FC851CBBF6D2424D500690E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439172	https://doi.org/10.3897/zookeys.955.51983.plate9	Plate 9. 1 Caucasopsis olsavskyi sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype 2 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype 3 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Kachara Cave 4 C. egrisi sp. nov., Samegrelo, Pirveli Balda, Motena Cave form 5, 6 C. egrisi sp. nov., Samegrelo, Pirveli Balda, spring in village: 5 holotype 6 paratype 7 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype 9 I. nakeralaensis sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype 10 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 9. 1 Caucasopsis olsavskyi sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype 2 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype 3 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Kachara Cave 4 C. egrisi sp. nov., Samegrelo, Pirveli Balda, Motena Cave form 5, 6 C. egrisi sp. nov., Samegrelo, Pirveli Balda, spring in village: 5 holotype 6 paratype 7 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype 9 I. nakeralaensis sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype 10 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
B16AB4C717385FC284D28FBC76A2DDC8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439185	https://doi.org/10.3897/zookeys.955.51983.figure15	Figure 15. Distribution map of genera Pontohoratia Vinarski, Palatov & Gloeer, 2014; Motsametia Vinarski, Palatov & Gloeer, 2014 and Hausdorfenia gen. nov. 1 P. smyri Vinarski, Palatov & Gloeer, 2014 (black dot) 2 - 4 P. birsteini (Starobogatov, 1962) (grey dots) 6 M. borutzkii (Shadin, 1932) (purple dot) 10 H. shareula sp. nov. (dark red dot) 12 - 15 H. hauffeniaformis sp. nov. (red dots) 27 P. pichkhaiai sp. nov. (light blue dot) 28 P. mapeli sp. nov. (medium blue dot) 30 - 31 P. vinarskii sp. nov. (dark blue dot).	Figure 15. Distribution map of genera Pontohoratia Vinarski, Palatov & Gloeer, 2014; Motsametia Vinarski, Palatov & Gloeer, 2014 and Hausdorfenia gen. nov. 1 P. smyri Vinarski, Palatov & Gloeer, 2014 (black dot) 2 - 4 P. birsteini (Starobogatov, 1962) (grey dots) 6 M. borutzkii (Shadin, 1932) (purple dot) 10 H. shareula sp. nov. (dark red dot) 12 - 15 H. hauffeniaformis sp. nov. (red dots) 27 P. pichkhaiai sp. nov. (light blue dot) 28 P. mapeli sp. nov. (medium blue dot) 30 - 31 P. vinarskii sp. nov. (dark blue dot).	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
2871C1CA50B85997AACBC79D9DB809F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439187	https://doi.org/10.3897/zookeys.955.51983.plate20	Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
2871C1CA50B85997AACBC79D9DB809F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439191	https://doi.org/10.3897/zookeys.955.51983.figure17	Figure 17. Hausdorfenia pseudohauffenia sp. nov. A, B operculum, SBMNH 633086. SEM SBMNH Vanessa Delnavaz.	Figure 17. Hausdorfenia pseudohauffenia sp. nov. A, B operculum, SBMNH 633086. SEM SBMNH Vanessa Delnavaz.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
1C425869E88C5CBFAF75137953446DBA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439187	https://doi.org/10.3897/zookeys.955.51983.plate20	Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
29048650A83252449510B797A1CB57AF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439170	https://doi.org/10.3897/zookeys.955.51983.plate8	Plate 8. 1 - 4 Imeretiopsis gorgoleti sp. nov., Racha, Gorgoleti, small spring above the road to Tsakhi: 1 holotype 2 paratype NHMW 113279 3, 4 paratypes 5, 6 I. iazoni sp. nov., Imereti, Kutaisi, Iazoni Cave: 5 holotype 6 paratype 7, 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni Cave, paratypes 9 - 13 I. gorgoleti sp. nov., specimens used for molecular and anatomical studies 14 I. gorgoleti sp. nov., operculum. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph Sara Schnedl, NHMW, and A. Rysiewska.	Plate 8. 1 - 4 Imeretiopsis gorgoleti sp. nov., Racha, Gorgoleti, small spring above the road to Tsakhi: 1 holotype 2 paratype NHMW 113279 3, 4 paratypes 5, 6 I. iazoni sp. nov., Imereti, Kutaisi, Iazoni Cave: 5 holotype 6 paratype 7, 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni Cave, paratypes 9 - 13 I. gorgoleti sp. nov., specimens used for molecular and anatomical studies 14 I. gorgoleti sp. nov., operculum. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph Sara Schnedl, NHMW, and A. Rysiewska.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
6082F8E275975B299F24A32B5E91899A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439169	https://doi.org/10.3897/zookeys.955.51983.plate7	Plate 7. 1 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, paratype SBMNH 633098 2 I. nakeralaensis sp. nov., Imereti, Tikbuli, spring below Nakerala Pass, paratype SBMNH 633110 3 I. gorgoleti sp. nov., Racha, Gorgoleti, above road to Tsakhi, paratype SBMNH 635910 4 Caucasopsis egrisi sp. nov., Pirveli Blada, spring in village, paratype SBMNH 639559. Scale bars: 1 mm. SEM SBMNH Vanessa Delnavaz.	Plate 7. 1 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, paratype SBMNH 633098 2 I. nakeralaensis sp. nov., Imereti, Tikbuli, spring below Nakerala Pass, paratype SBMNH 633110 3 I. gorgoleti sp. nov., Racha, Gorgoleti, above road to Tsakhi, paratype SBMNH 635910 4 Caucasopsis egrisi sp. nov., Pirveli Blada, spring in village, paratype SBMNH 639559. Scale bars: 1 mm. SEM SBMNH Vanessa Delnavaz.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
6082F8E275975B299F24A32B5E91899A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439171	https://doi.org/10.3897/zookeys.955.51983.figure11	Figure 11. Morphology of the penis A, B Caucasopsis letsurtsume sp. nov., Samegrelo: A ventrally B dorsally C Imeretiopsis gorgoleti sp. nov. Photograph A. Falniowski and A. Rysiewska.	Figure 11. Morphology of the penis A, B Caucasopsis letsurtsume sp. nov., Samegrelo: A ventrally B dorsally C Imeretiopsis gorgoleti sp. nov. Photograph A. Falniowski and A. Rysiewska.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
934DA6999F065F95888E844AC31383DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439167	https://doi.org/10.3897/zookeys.955.51983.figure10	Figure 10. Distribution map of Imeretiopsis gen. nov.: (magenta dots) and Caucasopsis gen. nov. (blue dots) 6 I. cameroni sp. nov. and I. iazoni sp. nov. 7 Imeretiopsis prometheus sp. nov. 8 I. nakeralaensis sp. nov. 9 I. gorgoleti sp. nov. 0 Caucasopsis orientalis (Starobogatov, 1962), C. subovata (Starobogatov, 1962) and C. pulcherrima (Starobogatov, 1962) 3 C. shadini (Starobogatov, 1962) 4 C. aculeus (Starobogatov, 1962) and C. schakuranica (Starobogatov, 1962) 22, 23 C. egrisi sp. nov. 24 C. olsavskyi sp. nov. 25, 30, 31 C. letsurtsume sp. nov.	Figure 10. Distribution map of Imeretiopsis gen. nov.: (magenta dots) and Caucasopsis gen. nov. (blue dots) 6 I. cameroni sp. nov. and I. iazoni sp. nov. 7 Imeretiopsis prometheus sp. nov. 8 I. nakeralaensis sp. nov. 9 I. gorgoleti sp. nov. 0 Caucasopsis orientalis (Starobogatov, 1962), C. subovata (Starobogatov, 1962) and C. pulcherrima (Starobogatov, 1962) 3 C. shadini (Starobogatov, 1962) 4 C. aculeus (Starobogatov, 1962) and C. schakuranica (Starobogatov, 1962) 22, 23 C. egrisi sp. nov. 24 C. olsavskyi sp. nov. 25, 30, 31 C. letsurtsume sp. nov.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
28ACD3799F4557609D11D2E37785C07B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439170	https://doi.org/10.3897/zookeys.955.51983.plate8	Plate 8. 1 - 4 Imeretiopsis gorgoleti sp. nov., Racha, Gorgoleti, small spring above the road to Tsakhi: 1 holotype 2 paratype NHMW 113279 3, 4 paratypes 5, 6 I. iazoni sp. nov., Imereti, Kutaisi, Iazoni Cave: 5 holotype 6 paratype 7, 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni Cave, paratypes 9 - 13 I. gorgoleti sp. nov., specimens used for molecular and anatomical studies 14 I. gorgoleti sp. nov., operculum. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph Sara Schnedl, NHMW, and A. Rysiewska.	Plate 8. 1 - 4 Imeretiopsis gorgoleti sp. nov., Racha, Gorgoleti, small spring above the road to Tsakhi: 1 holotype 2 paratype NHMW 113279 3, 4 paratypes 5, 6 I. iazoni sp. nov., Imereti, Kutaisi, Iazoni Cave: 5 holotype 6 paratype 7, 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni Cave, paratypes 9 - 13 I. gorgoleti sp. nov., specimens used for molecular and anatomical studies 14 I. gorgoleti sp. nov., operculum. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph Sara Schnedl, NHMW, and A. Rysiewska.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
1B2BA0EC1C6452D79A3D1249E5451D1B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439172	https://doi.org/10.3897/zookeys.955.51983.plate9	Plate 9. 1 Caucasopsis olsavskyi sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype 2 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype 3 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Kachara Cave 4 C. egrisi sp. nov., Samegrelo, Pirveli Balda, Motena Cave form 5, 6 C. egrisi sp. nov., Samegrelo, Pirveli Balda, spring in village: 5 holotype 6 paratype 7 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype 9 I. nakeralaensis sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype 10 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 9. 1 Caucasopsis olsavskyi sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype 2 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype 3 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Kachara Cave 4 C. egrisi sp. nov., Samegrelo, Pirveli Balda, Motena Cave form 5, 6 C. egrisi sp. nov., Samegrelo, Pirveli Balda, spring in village: 5 holotype 6 paratype 7 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype 9 I. nakeralaensis sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype 10 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
D5DCFC9B58365627B4FB771A06C5DCE7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439168	https://doi.org/10.3897/zookeys.955.51983.plate6	Plate 6. 1 - 6 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, specimens used for molecular and anatomical studies. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph A. Falniowski.	Plate 6. 1 - 6 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, specimens used for molecular and anatomical studies. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph A. Falniowski.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
2A1F599698F75C50BEBF74AE850C81A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439164	https://doi.org/10.3897/zookeys.955.51983.figure9	Figure 9. Kartvelobia sinuata sp. nov., Imereti, Satsiskvilo, Turchusmtha, morphology of the reproductive organs A-D morphology of the penis E female renal and pallial section of reproductive organs. Bc - bursa copulatrix; cbc - duct of bursa copulatrix; ga - albuminoid gland; gn - nidamental gland; gp - gonoporus; ov - oviduct; ovl - loop of oviduct; rs - receptaculum seminis; vc - ventral channel. Photograph and drawing A. Falniowski.	Figure 9. Kartvelobia sinuata sp. nov., Imereti, Satsiskvilo, Turchusmtha, morphology of the reproductive organs A-D morphology of the penis E female renal and pallial section of reproductive organs. Bc - bursa copulatrix; cbc - duct of bursa copulatrix; ga - albuminoid gland; gn - nidamental gland; gp - gonoporus; ov - oviduct; ovl - loop of oviduct; rs - receptaculum seminis; vc - ventral channel. Photograph and drawing A. Falniowski.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
0BF8373340C25EE4B298C7FBCA54668B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439160	https://doi.org/10.3897/zookeys.955.51983.figure8	Figure 8. Distribution map of Kartvelobia gen. nov. 16 K. kinchkha sp. nov. (blue dot) 17 - 21 K. sinuata sp. nov. (magenta dots) 22, 23 K. cf. sinuata (purple dots) 26, 27 K. shishaensis sp. nov. (red dots).	Figure 8. Distribution map of Kartvelobia gen. nov. 16 K. kinchkha sp. nov. (blue dot) 17 - 21 K. sinuata sp. nov. (magenta dots) 22, 23 K. cf. sinuata (purple dots) 26, 27 K. shishaensis sp. nov. (red dots).	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
4705966665545CD29DB4A566EE90E5FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439165	https://doi.org/10.3897/zookeys.955.51983.plate4	Plate 4. 1 - 4 Kartvelobia shishaensis sp. nov., Georgia, Samegrelo, Mukhuri, Shisha Spring 1 holotype 2 paratype 3, 4 K. kinchkha sp. nov., Imereti, Kinchkha, small travertine waterfall at right bank of Okatse below the large waterfall: 3 holotype 4 paratype 5, 6 K. sinuata sp. nov., Imereti, Kinchkhaperdi, spring along road to Askhi Plateau, dwarf population 7 K. sinuata sp. nov., Imereti, Satsiskvilo, Turchusmtha, paratype - typical form 8, 9 K. cf. shishaensis sp. nov., Georgia, Samegrelo, Mukhuri, Mapeli Cave. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 4. 1 - 4 Kartvelobia shishaensis sp. nov., Georgia, Samegrelo, Mukhuri, Shisha Spring 1 holotype 2 paratype 3, 4 K. kinchkha sp. nov., Imereti, Kinchkha, small travertine waterfall at right bank of Okatse below the large waterfall: 3 holotype 4 paratype 5, 6 K. sinuata sp. nov., Imereti, Kinchkhaperdi, spring along road to Askhi Plateau, dwarf population 7 K. sinuata sp. nov., Imereti, Satsiskvilo, Turchusmtha, paratype - typical form 8, 9 K. cf. shishaensis sp. nov., Georgia, Samegrelo, Mukhuri, Mapeli Cave. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
A372AE2D1BD95AFC9B3BC9240F13178B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439165	https://doi.org/10.3897/zookeys.955.51983.plate4	Plate 4. 1 - 4 Kartvelobia shishaensis sp. nov., Georgia, Samegrelo, Mukhuri, Shisha Spring 1 holotype 2 paratype 3, 4 K. kinchkha sp. nov., Imereti, Kinchkha, small travertine waterfall at right bank of Okatse below the large waterfall: 3 holotype 4 paratype 5, 6 K. sinuata sp. nov., Imereti, Kinchkhaperdi, spring along road to Askhi Plateau, dwarf population 7 K. sinuata sp. nov., Imereti, Satsiskvilo, Turchusmtha, paratype - typical form 8, 9 K. cf. shishaensis sp. nov., Georgia, Samegrelo, Mukhuri, Mapeli Cave. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 4. 1 - 4 Kartvelobia shishaensis sp. nov., Georgia, Samegrelo, Mukhuri, Shisha Spring 1 holotype 2 paratype 3, 4 K. kinchkha sp. nov., Imereti, Kinchkha, small travertine waterfall at right bank of Okatse below the large waterfall: 3 holotype 4 paratype 5, 6 K. sinuata sp. nov., Imereti, Kinchkhaperdi, spring along road to Askhi Plateau, dwarf population 7 K. sinuata sp. nov., Imereti, Satsiskvilo, Turchusmtha, paratype - typical form 8, 9 K. cf. shishaensis sp. nov., Georgia, Samegrelo, Mukhuri, Mapeli Cave. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
E0FF0A3480B75B4EA2C563FDA068149B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439161	https://doi.org/10.3897/zookeys.955.51983.plate1	Plate 1. 1 - 3 Kartvelobia sinuata sp. nov. Imereti, Satsiskvilo, Turchusmtha, specimens used for molecular and anatomical study. Measurement of aperture declination angle (CA): aperture-elongation axis vs. the columella is depicted in green. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph A. Falniowski.	Plate 1. 1 - 3 Kartvelobia sinuata sp. nov. Imereti, Satsiskvilo, Turchusmtha, specimens used for molecular and anatomical study. Measurement of aperture declination angle (CA): aperture-elongation axis vs. the columella is depicted in green. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph A. Falniowski.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
E0FF0A3480B75B4EA2C563FDA068149B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439164	https://doi.org/10.3897/zookeys.955.51983.figure9	Figure 9. Kartvelobia sinuata sp. nov., Imereti, Satsiskvilo, Turchusmtha, morphology of the reproductive organs A-D morphology of the penis E female renal and pallial section of reproductive organs. Bc - bursa copulatrix; cbc - duct of bursa copulatrix; ga - albuminoid gland; gn - nidamental gland; gp - gonoporus; ov - oviduct; ovl - loop of oviduct; rs - receptaculum seminis; vc - ventral channel. Photograph and drawing A. Falniowski.	Figure 9. Kartvelobia sinuata sp. nov., Imereti, Satsiskvilo, Turchusmtha, morphology of the reproductive organs A-D morphology of the penis E female renal and pallial section of reproductive organs. Bc - bursa copulatrix; cbc - duct of bursa copulatrix; ga - albuminoid gland; gn - nidamental gland; gp - gonoporus; ov - oviduct; ovl - loop of oviduct; rs - receptaculum seminis; vc - ventral channel. Photograph and drawing A. Falniowski.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
2AA6C99D3622515BAEC977C931047E85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439192	https://doi.org/10.3897/zookeys.955.51983.figure18	Figure 18. Motsametia borutzkii (Shadin, 1932) A-D Imereti, Kutaisi, Iazoni (Tskal-Tsiteli) Cave. Photograph J. Grego.	Figure 18. Motsametia borutzkii (Shadin, 1932) A-D Imereti, Kutaisi, Iazoni (Tskal-Tsiteli) Cave. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
6F29FFEA9D675B2CAAA738CE7D741348.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439188	https://doi.org/10.3897/zookeys.955.51983.figure16	Figure 16. A, B Pontohoratia vinarskii sp. nov.: Samegrelo, Chkhorotsku, Nazodelavo Cave, morphology of penis C, D P. pichkhaiai sp. nov.: Samegrelo, Chkhorotsku, Shisha Spring, morphology of penis E, F P. mapeli. sp.: Samegrelo, Kanti, Mapeli Spring, morphology of penis. Photograph A. Falniowski and A. Rysiewska; drawing A. Falniowski.	Figure 16. A, B Pontohoratia vinarskii sp. nov.: Samegrelo, Chkhorotsku, Nazodelavo Cave, morphology of penis C, D P. pichkhaiai sp. nov.: Samegrelo, Chkhorotsku, Shisha Spring, morphology of penis E, F P. mapeli. sp.: Samegrelo, Kanti, Mapeli Spring, morphology of penis. Photograph A. Falniowski and A. Rysiewska; drawing A. Falniowski.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
6F29FFEA9D675B2CAAA738CE7D741348.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439187	https://doi.org/10.3897/zookeys.955.51983.plate20	Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
26D3ABD365C057FE97F649AD0323134E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439187	https://doi.org/10.3897/zookeys.955.51983.plate20	Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
26D3ABD365C057FE97F649AD0323134E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439188	https://doi.org/10.3897/zookeys.955.51983.figure16	Figure 16. A, B Pontohoratia vinarskii sp. nov.: Samegrelo, Chkhorotsku, Nazodelavo Cave, morphology of penis C, D P. pichkhaiai sp. nov.: Samegrelo, Chkhorotsku, Shisha Spring, morphology of penis E, F P. mapeli. sp.: Samegrelo, Kanti, Mapeli Spring, morphology of penis. Photograph A. Falniowski and A. Rysiewska; drawing A. Falniowski.	Figure 16. A, B Pontohoratia vinarskii sp. nov.: Samegrelo, Chkhorotsku, Nazodelavo Cave, morphology of penis C, D P. pichkhaiai sp. nov.: Samegrelo, Chkhorotsku, Shisha Spring, morphology of penis E, F P. mapeli. sp.: Samegrelo, Kanti, Mapeli Spring, morphology of penis. Photograph A. Falniowski and A. Rysiewska; drawing A. Falniowski.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
FDA37D44903B56A5BAE425C982C06ADE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439185	https://doi.org/10.3897/zookeys.955.51983.figure15	Figure 15. Distribution map of genera Pontohoratia Vinarski, Palatov & Gloeer, 2014; Motsametia Vinarski, Palatov & Gloeer, 2014 and Hausdorfenia gen. nov. 1 P. smyri Vinarski, Palatov & Gloeer, 2014 (black dot) 2 - 4 P. birsteini (Starobogatov, 1962) (grey dots) 6 M. borutzkii (Shadin, 1932) (purple dot) 10 H. shareula sp. nov. (dark red dot) 12 - 15 H. hauffeniaformis sp. nov. (red dots) 27 P. pichkhaiai sp. nov. (light blue dot) 28 P. mapeli sp. nov. (medium blue dot) 30 - 31 P. vinarskii sp. nov. (dark blue dot).	Figure 15. Distribution map of genera Pontohoratia Vinarski, Palatov & Gloeer, 2014; Motsametia Vinarski, Palatov & Gloeer, 2014 and Hausdorfenia gen. nov. 1 P. smyri Vinarski, Palatov & Gloeer, 2014 (black dot) 2 - 4 P. birsteini (Starobogatov, 1962) (grey dots) 6 M. borutzkii (Shadin, 1932) (purple dot) 10 H. shareula sp. nov. (dark red dot) 12 - 15 H. hauffeniaformis sp. nov. (red dots) 27 P. pichkhaiai sp. nov. (light blue dot) 28 P. mapeli sp. nov. (medium blue dot) 30 - 31 P. vinarskii sp. nov. (dark blue dot).	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
5F0459E14BC65E6AACA4D9890AC9584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439186	https://doi.org/10.3897/zookeys.955.51983.plate19	Plate 19. 1 - 5 Pontohoratia vinarskii sp. nov.: Samegrelo, Chkhorotsku, Letsurtsume Cave, specimens used for molecular and anatomical study. The numbers correspond to individuals. Photograph A. Falniowski.	Plate 19. 1 - 5 Pontohoratia vinarskii sp. nov.: Samegrelo, Chkhorotsku, Letsurtsume Cave, specimens used for molecular and anatomical study. The numbers correspond to individuals. Photograph A. Falniowski.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
5F0459E14BC65E6AACA4D9890AC9584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439188	https://doi.org/10.3897/zookeys.955.51983.figure16	Figure 16. A, B Pontohoratia vinarskii sp. nov.: Samegrelo, Chkhorotsku, Nazodelavo Cave, morphology of penis C, D P. pichkhaiai sp. nov.: Samegrelo, Chkhorotsku, Shisha Spring, morphology of penis E, F P. mapeli. sp.: Samegrelo, Kanti, Mapeli Spring, morphology of penis. Photograph A. Falniowski and A. Rysiewska; drawing A. Falniowski.	Figure 16. A, B Pontohoratia vinarskii sp. nov.: Samegrelo, Chkhorotsku, Nazodelavo Cave, morphology of penis C, D P. pichkhaiai sp. nov.: Samegrelo, Chkhorotsku, Shisha Spring, morphology of penis E, F P. mapeli. sp.: Samegrelo, Kanti, Mapeli Spring, morphology of penis. Photograph A. Falniowski and A. Rysiewska; drawing A. Falniowski.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
5F0459E14BC65E6AACA4D9890AC9584A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/439187	https://doi.org/10.3897/zookeys.955.51983.plate20	Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.	2020-12-31	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian		Zenodo	biologists	Grego, Jozef;Mumladze, Levan;Falniowski, Andrzej;Osikowski, Artur;Rysiewska, Aleksandra;Palatov, Dimitry M.;Hofman, Sebastian			
