identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0A799443FFE05945FF53F8E0B956FC59.text	0A799443FFE05945FF53F8E0B956FC59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zenoria limitrophi Gonzalez & Santos	<div><p>Zenoria limitrophi González &amp; Santos, sp. nov.</p><p>Figs. 1–13</p><p>Diagnosis. Zenoria limitrophi sp. nov. has distinctive male genitalia. The structure differs from all other species in that the penis guide is as wide as long, while in other species it is much longer than wide, except Z. lativerpa González &amp; Honour (2012), which is as wide as long, but with its maximum width at the apex and not at the base.</p><p>Description of holotype, male. Length 3.9 mm; width 3.0 mm. Body oval, elongate, maximum width at basal third of elytra (Fig. 1). Head, antennae, mouthparts, prothorax, legs and abdomen pale yellow; median basal mark on pronotum black, anteriorly rounded. Head with sparse yellowish-white pubescence. Scutellar shield black. Elytra dark metallic green with narrow humeral and apical pale-yellow margin, widened at apex; yellowish-white pubescence, sparse, except discal area with sparse brown pubescence (Figs. 1–4). Margin of elytron broadly explanate, inner carina extending about ¼ distance to outer margin near elytral base. Meso- and metaventrite black. Abdominal postcoxal line curved posterolaterally, closely paralleling posterior margin of ventrite for short distance, then abruptly redirected anteriorly toward base of ventrite in lateral third (Fig. 5). Posterior margin of ventrite V smoothly arched, convex. Posterior margin of ventrite VI with wide and deep semicircular emargination (Fig. 6). Phallobase more than half length of tegminal strut, nearly two times longer than wide (Fig. 7). Penis guide mamilliform, as wide as long at base, first third almost parallel-sided, sharply narrowed at midlength, subtriangular with rounded apex, with tubercules in apical third (Fig. 8); curved in lateral view, basal half with keel on inner side, measuring 2/5 of its length, narrower (Figs. 11–12). Parameres projecting beyond penis guide subtriangular in ventral view, curved inward and narrower in distal third, with short setae at apex (Fig. 9), curved in lateral view in basal half and nearly straight in distal half; apex pointed (Figs. 10–11). Penis normal wide, no more than 40 times longer than wide, semicircular in basal half; apical half straight (Fig. 13); apex bifurcate with outer side short and blunt, inner side long, sharp and slightly curved downward (Fig. 14); penis capsule with inner branch curved slightly inward and ending in a club, outer branch narrowing only near end, with outer membranous projection (Fig. 13).</p><p>Female. Length 3.6 mm; width 3.0 mm. Similar to male except pronotum dark with narrow yellowish anterior and lateral margins, elytra lacking yellow humeral spot, apical spot shorter and narrower than in male (Fig. 15). Ventrite VI with shallow emargination on posterior margin. Spermatheca J-shaped, cornu thickened toward distal end, with lamellar projection and barely visible basal ramus; nodulus extending in same direction as cornu, conical; accessory gland present; sperm duct long; infundibulum absent; apex of bursa copulatrix not modified (Fig. 16). Coxites four times longer than wide, triangular; styli well developed (Fig. 17).</p><p>Variation. Length 3.6–4.1 mm; width 3.0– 3.2 mm.</p><p>Etymology. From the Latin “limitrophus”, indicating their location near on the border between Brazil, Colombia and Peru.</p><p>Type material. Holotype. “ Brasil, Amazonas, / Tabatinga, 50 m / 01-IX-2006, leg. R. / Westerduijn, understory / of open altered forest” “ Male 964” “ Zenoria sp . nov. 2 / Santos P.B. 2013” [DZUP].</p><p>Paratypes (3): (1) “ Brasil, Amazonas, / Tabatinga, 26-VIII al / 6-IX-2006, Leg. R. / Westerduijn. Understory / mature altered forest” “ Female 357” “ Zenoria / sp. 8 / Det.: G. González 2011” “ Zenoria sp . nov. 2 / Santos P.B. 2013” [DZUP]. (2) “ Brasil, Amazonas, / Tabatinga, 50 m / 01-IX-2006, leg. R. / Westerduijn, understory / of open altered forest” “ Male ” “ Zenoria / sp. 10 / Det.: G. González 2011” “ Zenoria sp . nov. 2 / Santos P.B. 2013” [DZUP].</p><p>Distribution. Brazil, Amazonas.</p></div>	https://treatment.plazi.org/id/0A799443FFE05945FF53F8E0B956FC59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Santos, Paula B.;González, Guillermo	Santos, Paula B., González, Guillermo (2016): New species of Zenoria Mulsant (Coleoptera: Coccinellidae) from Brazil and Peru. Zootaxa 4205 (4): 386-394, DOI: 10.11646/zootaxa.4205.4.7
0A799443FFE25942FF53FBD3BA9AF940.text	0A799443FFE25942FF53FBD3BA9AF940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zenoria westerduijni Gonzalez & Santos	<div><p>Zenoria westerduijni González &amp; Santos, sp. nov.</p><p>Figs. 18–30</p><p>Diagnosis. Z. westerduijni sp. nov. is distinguishable from all other species of the genus by the almost entirely black body. It also differs from other species by the slightly angulate postcoxal line and the extremely narrow penis, about 50 times longer than wide, typically 35–40 times in other species of the genus.</p><p>Description of holotype, male. Length 4.0 mm; width 3.4 mm. Body oval, rounded, convex (Figs. 18–19). Head and pronotum black, latter with narrow yellow stripe on anterior border; antennae and mouthparts brown, apex sometimes darker; pubescence grayish white, short, irregularly distributed (Fig. 20). Prothorax black; pubescence light brown, short. Scutellar shield black. Elytra entirely black, shiny; pubescence light brown except for central area, with inconspicuous dark-brown pubescence; this pubescence lacking in some areas, probably due to rubbing (Figs. 18–21). Epipleuron black, inner carina extending more than half distance to outer margin. Meso- and metaventrite brown. Legs brown. Abdominal postcoxal line curved posterolaterally, reaching posterior margin of ventrite for short distance, then redirected anteriorly toward base of ventrite in lateral third (Fig. 22). Posterior margin of ventrite V with smooth notch. Posterior margin of ventrite VI with deep triangular emargination (Fig. 23). Phallobase short, half length of tegminal strut, two times longer than wide (Fig. 24). Penis guide triangular in dorsal view, two times longer than wide, with rounded apex (Figs. 25–27); in lateral view, triangular in basal 3/5, parallel-sided and narrow in distal 2/5 (Figs. 28–29). Penis guide projecting beyond parameres (Fig. 25), parallelsided in ventral view, slightly projected apically (Fig. 27); regularly curved in lateral view and tapering from base to apex, the latter with small truncated widening (Figs. 28–29). Sparse, very short setae at end of paramere (Fig. 29). Penis narrow, more than 50 times longer than wide, semicircular in basal third, gently curved in middle third, and straight in apical third (Fig. 30), ending in pointed apex, membranous areas in distal 1/20 of apex, on lower side (Fig. 31); inner branch of penis capsule slightly curved inward and truncated, outer branch subtriangular, tapered with hook, inconspicuous membranous projection on outer margin (Fig. 30).</p><p>Female. Unknown.</p><p>Variation. Not observed.</p><p>Etymology. This species is named after the biologist and entomologist Rob Westerduijn, who collected this specimen and several others described in this contribution.</p><p>Type material. Holotype. “ Perú, Loreto / Picuroyacu, Pongo / 150 m, 18-XI-2008 Leg. / R. Westerduijn, / Understory of altered / mature forest” “ Male 1612” [UNMSM].</p><p>Distribution. Peru, Loreto.</p></div>	https://treatment.plazi.org/id/0A799443FFE25942FF53FBD3BA9AF940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Santos, Paula B.;González, Guillermo	Santos, Paula B., González, Guillermo (2016): New species of Zenoria Mulsant (Coleoptera: Coccinellidae) from Brazil and Peru. Zootaxa 4205 (4): 386-394, DOI: 10.11646/zootaxa.4205.4.7
0A799443FFE55940FF53F8A5B937FBA5.text	0A799443FFE55940FF53F8A5B937FBA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zenoria carbo Gonzalez & Santos	<div><p>Zenoria carbo González &amp; Santos, sp. nov.</p><p>Figs. 32–45</p><p>Diagnosis. Z. carbo sp. nov. resembles a group of species with a long subtriangular penis guide, about 3 times longer than wide, and parameres with setae only at the apex. Other species that have these features are Z.</p><p>circumcincta Gordon, 1971; Z. emarginata Gordon, 1971; Z. linteolata Mulsant, 1850 and Z. purpurea Gordon, 1972 . The new species differs from Z. circumcincta in the more elongated phallobase (3 times longer than wide) and the parameres parallel sided with a straight outer margin in ventral view. It is distinguished from the other species by having a penis guide with a sinuous inner margin in lateral view. Apart from this, all other species are predominantly lighter in color (with or without dark spots on the elytral disc), except Z. linteolata and Z. purpurea, which have a dark metallic-green and dark metallic-purple elytra respectively.</p><p>Description of holotype, male. Length 3.2 mm; width 2.6 mm. Body oval, convex (Figs. 32–33). Head and prothorax pale yellow; antennae and mouthparts brown; pubescence yellowish white (Fig. 34). Pronotum with basal black spot occupying central ¾ of base and extending to middle; pubescence yellowish white, about half scutellar shield length (Figs. 32–34). Elytra black, shiny with yellowish-white pubescence, except central medial area with inconspicuous dark-brown pubescence (Figs. 32–35). Scutellar shield black (Fig. 32). Epipleuron black, inner carina extending half distance to outer margin. Meso- and metaventrite black. Legs pale yellow. Abdominal postcoxal line curved posterolaterally, closely paralleling posterior margin of ventrite for short distance, then abruptly redirected anteriorly toward base of ventrite in lateral third (Fig. 36). Posterior margin of ventrite V with smooth emargination. Posterior margin of ventrite VI with wide emargination, accentuated medially, deep (Fig. 37). Phallobase long, ½ length of tegminal strut, three times longer than wide (Fig. 37). Penis guide triangular in dorsal view, nearly three times longer than wide, sides almost parallel, slightly narrower in apical fourth; apex rounded (Figs. 39–40); in lateral view triangular, inner margin slightly sinuous (Figs. 42–43). Parameres projecting slightly beyond penis guide (Fig. 39), in ventral view subtriangular with straight lateral margins, directed slightly inward in distal 1/5 (Fig. 41), regularly curved in lateral view and parallel sided. Sparse short setae at end of paramere (Figs. 42–43). Penis normal, wide, no more than 40 times longer than wide, semicircular in basal 3/5, straight in apical 2/5, except apex recurved upward relative to penis capsule, with membranous areas on outer side, fewer on inner side (Figs. 44–45); penis capsule with inner branch slightly longer than outer branch, curved inward, narrow at base and widening to 2/3 of length with membranous areas, outer branch with almost parallel sides, truncated, with outer membranous area (Fig. 44).</p><p>Female. Unknown.</p><p>Variation. Length 3.1–3.2 mm; width 2.5–2.6 mm. The pale yellow coloration of the pronotal anterior margins may be reduced to ½ or ¼ of the size.</p><p>Etymology. From Latin " carbo " meaning coal, referring to the intense black color of the elytra; proposed as a noun in apposition.</p><p>Type material. Holotype. “ Perú, Madre de Dios / Tahuamanu, Iberia / 292 mnsm, 10-I-2010 / Leg. J. Miró, Cítricos ” “ Male 1617” [MEUT].</p><p>Paratypes (2): (1) “ Perú, Madre de Dios / Tahuamanu, Iberia / 292 mnsm, 10-I-2010 / Leg. J. Miró, Cítricos ” “ Male 612” [MEUT] . (1) “ Perú Madre de Dios dept. / CICRA <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.10962&amp;materialsCitation.latitude=-12.55207" title="Search Plazi for locations around (long -70.10962/lat -12.55207)">FldStn</a>, trail 6 research / plot, 12.55207°S 70.10962°W / 295m, 11-13.VI.2011, Chaboo / team, canopy Malaise, bottom / sample, PER-11-CMB-002” “[bar code] / SEM C0986969 / KUNHM-ENT” “ Male 1571” [UNMSM] .</p><p>Distribution. Peru, Madre de Dios.</p></div>	https://treatment.plazi.org/id/0A799443FFE55940FF53F8A5B937FBA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Santos, Paula B.;González, Guillermo	Santos, Paula B., González, Guillermo (2016): New species of Zenoria Mulsant (Coleoptera: Coccinellidae) from Brazil and Peru. Zootaxa 4205 (4): 386-394, DOI: 10.11646/zootaxa.4205.4.7
0A799443FFE75941FF53FB88B937FE40.text	0A799443FFE75941FF53FB88B937FE40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zenoria miroi Gonzalez & Santos	<div><p>Zenoria miroi González &amp; Santos, sp. nov.</p><p>Figs. 46–59</p><p>Diagnosis. Zenoria miroi sp. nov. is easily distinguishable from other species with the same color pattern, Z. crotchi Gordon, 1971; Z. delicatula Weise, 1910; Z. discoidalis (Kirsch, 1876); Z. lativerpa González &amp; Honour, 2012; Z. luciae González &amp; Honour, 2012; and Z. sylvatica González &amp; Honour, 2012 by the subtriangular black basal spot on the pronotum. The male genitalia with short, inflated parameres, curved in lateral view with sensory setae limited to the apex are distinctly different from other described species. Although the male genitalia resemble Z. annularis Gordon, 1971 this species has broad triangular parameres, angulate at the base and straight on the outer margin in ventral view.</p><p>Description of holotype, male. Length 3.2 mm; width 2.5 mm. Body oval, sligthly convex (Figs. 46–47). Head pale yellow with yellowish-white pubescence, longer and more abundant toward clypeus; antennae and mouthparts brown (Fig. 48). Pronotum dark yellow with subrectangular basal black spot; pubescence yellowish white (Figs. 46–48). Elytra pale yellow with large black common spot on disc, covering part of humeral callus; pubescence yellowish-white, except elytral disc with dark-brown pubescence (Figs. 46–49). Scutellar shield black (Fig. 46). Epipleuron yellow, inner carina extending less than half distance to outer margin. Meso- and metaventrite black. Legs pale yellow. Abdominal postcoxal line curved posterolaterally, closely paralleling posterior margin of ventrite for short distance, then abruptly redirected anteriorly toward base of ventrite in lateral third (Fig. 50). Posterior margin of ventrite V arcuate. Posterior margin of ventrite VI with broad and deep semicircular emargination (Fig. 51). Phallobase short, less than half length of tegminal strut, two times longer than wide (Fig. 52). Penis guide mamilliform, basal third broad with convex sides, apex rounded (Figs. 53–55); in lateral view broad at base, outer margin straight, inner margin curved and not sinuous, with tubercles (Figs. 56–57). Parameres projecting beyond penis guide in ventral view; outer margin wide and curved, inner margin parallel-sided (Figs. 53–54); curved in lateral view, apex truncate; short sensory setae at apex (Figs. 56–57). Penis normal, wide, no more than 40 times longer than wide, basal third semicircular, middle third curved, straight in apical third except apex curved downward and tapering upward relative to penis capsule in short, very narrow projection (Fig. 58); apical membranous projections on both sides (Fig. 59); penis capsule with inner branch curved inward, ending in enlargement, and triangular outer branch (Fig. 58).</p><p>Female. Unknown.</p><p>Variation. Not observed.</p><p>Etymology. This species is named after the agronomist Jimmy Miro, collector of this species and many other species of Coccinellidae in Tumbes and Madre de Dios departments, Peru.</p><p>Holotype. “ Perú, Madre de Dios / Tambopata, Las / Piedras, Alegría, 5-VIII / 2012, leg. Jimmy Miró ” “N: 8660120 E: 486620 / Cítricos: Naranjo ” “ Male 1100” [MEUT].</p><p>Distribution. Peru, Madre de Dios.</p></div>	https://treatment.plazi.org/id/0A799443FFE75941FF53FB88B937FE40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Santos, Paula B.;González, Guillermo	Santos, Paula B., González, Guillermo (2016): New species of Zenoria Mulsant (Coleoptera: Coccinellidae) from Brazil and Peru. Zootaxa 4205 (4): 386-394, DOI: 10.11646/zootaxa.4205.4.7
