taxonID	type	description	language	source
090C879DFFE28E1C899AFE3CA97FFA93.taxon	vernacular_names	[Japanese name: Hime-kinokotsuyakeshi-hanekakushi-zoku]	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E1C899AFE3CA97FFA93.taxon	type_taxon	type species: Pseudatheta elegans Cameron, 1920).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E1C899AFE3CA97FFA93.taxon	discussion	Remarks. Species of this genus often have intraspecific variation in body coloration, the ratio of the length and width of each antennomere, and the processes on the posterior margin of the tergite VIII of males, which do not necessarily correspond to those described or illustrated in this paper. The genus Pseudatheta can be distinguished from the genus Phymatura by following character states: mesoventrite without longitudinal carina; number of setose lobes on flabellum of hind wing varies from zero to one.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E1C899AFE3CA97FFA93.taxon	discussion	Remarks. Pseudatheta hilaris (Sharp), comb. nov., is excluded from this key because only female specimens were available.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E19899AFA40AC27FEC6.taxon	description	(Figs. 1 A, 2 C, 5 A – C)	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E19899AFA40AC27FEC6.taxon	vernacular_names	[Japanese name: Miiro-kinokotsuyakeshi-hanekakushi]	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E19899AFA40AC27FEC6.taxon	distribution	type locality: “ Bukenji ” [Yokohama-shi, Kanagawa-ken, central Honshu, Japan]); Sawada, 1977: 217 (as a synonym of Ph. oligotinula).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E19899AFA40AC27FEC6.taxon	discussion	(transferred to the genus Phymatura from the genus Atheta); Shibata et al., 2013: 115 (catalogue); Schülke & Smetana, 2015: 633 (catalogue).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E19899AFA40AC27FEC6.taxon	discussion	(transferred to genus Pseudatheta from Phymatura; probably based on Sawada (1977 )).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E19899AFA40AC27FEC6.taxon	distribution	type locality: “ China: Sichuan, Ganzi Prefecture, Daxue Shan, Gongga Shan, Hailougou glacier park, 102 ° 04 ’ E 29 ° 36 ’ N, river valley, ca. 1 km above camp I, 2100 m ”); Schülke & Smetana, 2015: 634 (catalogue); Pace, 2016: 301 (Yunnan). Syn. nov.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E19899AFA40AC27FEC6.taxon	materials_examined	Material examined. Type material. Atheta (Datomicra) crenulicauda: Lectotype: here designated, male “ Hans Sauter 4000 / Bukenji 10 XI 05 / Unter Schilfhaufen [handwritten] // Bukenji / Japan. Sauter [handwritten] // crenulicauda / Brnh. Typ. [handwritten] // Chicago NHMus / M. Bernhauer / Collection ” (abdominal segments VIII – X and aedeagus were dissected and mounted in Euparal by MM) (FMNH). Paralectotype: 1 female, “ crenulicauda / Bernh. Cotypus [handwritten] // Unzen. 2600 ’ / Japan. Sauter [handwritten] // Chicago NHMus / M. Bernhauer / Collection ” (abdominal segments VIII – X and spermatheca were dissected and mounted in Euparal by MM) (FMNH). Additional material examined. JAPAN: [Aomori-ken]: 1 male, Yasumiya, Towada-shi, 4 – 5 VIII 1966, M. & M. T. Ch ̊ jô leg. (KUM); [Miyagi-ken]: 2 males, 1 female, Yoshida, Taiwa-chô, 20 VIII 2009, J. Aoki leg. (KUM); [Fukushima-ken]: 1 male, Shitokigawa-keikoku, Iwaki-shi, 24 V 1992, K. Haga leg. (KUM); [Tochigi-ken]: 1 male, 1 female, Watarase, 6 V 1979, M. Tao leg. (KUM); [Gunma-ken]: 1 male, Naganohara, Naganohara-machi, 9 VIII 1997, T. Kishimoto leg. (TKc); [Saitama-ken]: 1 male, 1 female, Yoshimi-machi, 16 VII 2000, K. Toyoda leg. (KUM); 3 males, 2 females, Mitsumine, Chichibu-shi, 28 V 1997, M. Maruyama leg. (KUM); [Chiba-ken]: 1 male, Noda-shi, 17 V 1991, T. Kishimoto leg. (TKc); [Tokushima-ken]: 1 male, Shikibidani, Naka-chô, 17 VII 2010, J. Aoki leg. (KUM); [Ehime-ken]: 1 male, Sara-ga-mine, 29 IV 1962, S. Hisamatsu leg. (EUM); Komenono, 18 VII 1975, A. Yonetsu leg. (EUM); 1 male, Sara-ga-mine, 27 VI 1959, M. Satô leg. (EUM); [Kochi-ken]: 1 male, 1 female, Yusuhara-chô, 5 V 2010, T. & T. Miyata leg. (KUM); 2 males, 1 female, Kamioriwatashi, Yusuhara-chô, 24 V 1997, M. Sakai leg. (EUM); [Nagasaki-ken]: Tsushima Is.: 1 male, Kamiagatamachi Sasuna, Tsushima-shi, 18 VI 2022, T. Hashizume leg. (KUM).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E19899AFA40AC27FEC6.taxon	description	Redescription. Measurements (n = 5): BL ≈ 1.86 – 2.39; FBL, 1.00 – 1.22; HL, 0.34 – 0.38; HW, 0.38 – 0.44; PL, 0.35 – 0.42; PW, 0.50 – 0.59; EL, 0.38 – 0.48; EW, 0.60 – 0.73. (Lectotype of Atheta crenulicauda: BL ≈ 2.0; PL, 0.40; PW, 0.55; HTL, 0.41). Relative length of antennomeres I – XI (n = 1): 25: 24: 21: 11: 11: 11: 11: 10: 11: 12: 37. Ratio of length / width of antennomeres I – XI (n = 1): 1.84: 2.07: 2.16: 0.88: 0.71: 0.68: 0.64: 0.55: 0.57: 0.63: 1.83. Body (Figs. 1 A, 2 C) reddish brown; head darker; posterolateral areas of elytra darker; abdominal segments V – VII darker. Head almost as long as wide, HW / HL: 1.11 – 1.26; surface densely covered with setae. Antenna with antennomeres I – III and XI longer than wide, antennomeres IV – X wider than long; antennomeres I and II with rounded distal end, antennomeres III – X strongly angulated near distal end, antennomere XI oval. Pronotum transverse, PW / PL: 1.36 – 1.50, PW / HW: 1.32 – 1.41; surface densely covered with setae, finely punctured, without microsculpture; posterior margin slightly bisinuate. Elytra wider than long, EW / EL: 1.60 – 1.70, EL / PL: 1.00 – 1.10, EW / PW: 1.20 – 1.24; surface densely covered with setae and finely punctured; posterior margin sinuate near posterolateral corners. Hind wings well developed; flabellum with one seta. Mesoventrite without longitudinal carina; mesoventral process extended to slightly beyond middle of mesocoxal cavities, with pointed apex; metaventral process shorter than mesoventral process, with rounded apex; isthmus present. Abdomen slightly narrowed posteriad; surface densely covered with setae. Male. Elytra with a pair of small tubercles at about posterior 1 / 3 on sutural margin. Sternite VI with small medial lobe on posterior margin. Tergite VII with a tubercle on posteromedian area. Tergite VIII (Fig. 5 A) with a tubercle on median area; three to five processes on each side of posterior margin, outer ones slightly curved, longer and sharper than the others. Aedeagus as in Fig. 5 B; median lobe with large basal bulb; apical process broad, dilated in lateral view, apical end thin and elongated, apex pointed; flagellum moderately long. Female. Spermatheca (Fig. 5 C) curved twice, with a transverse band-like structure at base of distal portion; distal portion elongated, U-shaped; median portion elongated, slightly curved; proximal portion longer than wide.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E19899AFA40AC27FEC6.taxon	distribution	Distribution. Japan (Honshu, Shikoku, Tsushima Is., Kyushu?); China (Sichuan, Yunnan).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE28E19899AFA40AC27FEC6.taxon	discussion	Remarks. Sawada (1977) redescribed this species in detail as Ph. oligotinula. Based on its characteristics (e. g., small body size, absence of the longitudinal carina on the mesoventrite, and the shape of the median lobe of the aedeagus and the spermatheca), this species belongs to Pseudatheta rather than to Phymatura. The illustration of the median lobe of the aedeagus of Ps. similis, shown by Pace (2010), fully agrees with that of Ps. crenulicauda (as Ps. similis syn. nov.). Pace (2016) recorded this species from Yunnan as Ps. similis. Kim & Ahn (2014) listed Ph. crenulicauda as a Korean species, based on the record of “ Ph. japonica ” from North Korea in Paśnik (2001), but later found that the specimens examined in Paśnik (2001) were not Ph. crenuicauda (Kim & Ahn, 2016). While the true species identity of these specimens is unknown, we tentatively consider this record as a record of Ph. japonica from North Korea. This species can be distinguished from its congeners by the apically dilated apical process and the absence of a ventral process of the median lobe of the aedeagus. There is currently no practical key to distinguish the female of this species from the female of its allies [i. e., Ps. hilaris, Ps. taiwanensis, Ps. thailandensis, and Ph. bigranipennis (Bernhauer, 1915)]. Thus, despite our detailed morphological observations, these species could not be distinguished based on female specimens. The paralectotype of this species was collected from Unzen, Nagasaki, Kyushu, and is the only record of this species from Kyushu; however, because it was a female, it could not be positively identified. Further study is needed to allow morphologically based species identification in Pseudatheta females, supported perhaps by DNA information (see also the remarks on Ph. hilaris).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE78E17899AFCDBACE9FEC7.taxon	description	(Figs. 1 B, 4, 5 D – F)	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE78E17899AFCDBACE9FEC7.taxon	vernacular_names	[Japanese name: Taiwan-miiro-kinokotsuyakeshi-hanekakushi]	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE78E17899AFCDBACE9FEC7.taxon	distribution	type locality: “ Taiwan, Kaohsiung Hs., for. abv. Tona For. Sta. 1100 m ”).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE78E17899AFCDBACE9FEC7.taxon	materials_examined	Materials examined. JAPAN: [Hokkaido]: 2 males, Midorimachi, Assabu-chô, 8 IX 2022, T. Hashizume leg. (KUM); 2 males, Ôsawaguchi, Nopporo-shinrin-kôen, Ebetsu-shi, 16 – 19 VI 2001, S. Hori leg. (KUM); 1 male, 1 female, Kannonzawa-rindô, Minami-ku, Sapporo-shi, 8 VIII 2021, T. Nozaki leg. (KUM); [Saitama-ken]: 1 male, 2 females, Futago-yama, 28 VIII 1984, H. Oda leg. (KUM); [Tokyo-to]: 1 male, Nippara, Okutama-machi, 15 VI 1997, K. Haga leg. (KUM); 1 male, 1 female, same data, but 5 V 1998 (KUM); [Tottori-ken]: 1 male, Hyô-no-sen, Wakasa-chô, 6 – 9 VIII 1968, I. H. B. C. (EUM); [Fukuoka-ken]: 2 males, 1 female, Hiko-san, Soeda-machi, 23 V 2020, S. Inoue leg. (KUM); 1 male, 2 female, Hiko-san, 7 V 1971, K. Takeno leg. (KUM); 1 male, Hiko-san, 15 X 1968, M. - T. Ch ̊ jô leg. (KUM); [Kumamoto-ken]: 1 male, Shiratori-yama, Izumimachi Momigi, Yatsushiro-shi, 14 V 2021, S. Inoue leg. (KUM); [Oita-ken]: 1 male, Kyûsuikei, Kokonoe-machi, 31 V – 10 VI 2011, S. Imasaka leg. (KUM).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE78E17899AFCDBACE9FEC7.taxon	description	Redescription. Measurements (n = 5): BL ≈ 1.90 – 2.13; FBL: 1.09 – 1.25; HL: 0.34 – 0.38; HW: 0.39 – 0.44; PL: 0.38 – 0.40; PW: 0.53 – 0.59; EL: 0.36 – 0.43; EW: 0.62 – 0.72. Relative length of antennomeres I – XI (n = 1): 20: 22: 18: 10: 12: 13: 13: 11: 12: 12: 33. Ratio of length / width of antennomeres I – XI (n = 1): 1.90: 2.27: 1.86: 0.80: 0.73: 0.78: 0.77: 0.63: 0.73: 0.69: 1.79. Body (Fig. 1 B) reddish brown; head darker; posterolateral areas of elytra darker; abdominal segments V – VII darker. Head almost as long as wide, HW / HL: 1.07 – 1.23; surface densely covered with setae. Antenna with antennomeres I – III and XI longer than wide, antennomeres IV – X wider than long; antennomeres I and II with rounded distal end, antennomeres III – X strongly angulated near distal end, antennomere XI oval. Pronotum transverse, PW / PL: 1.35 – 1.45, PW / HW: 1.31 – 1.41; surface densely covered with setae, finely punctured, without microsculpture; posterior margin slightly bisinuate. Elytra wider than long, EW / EL: 1.60 – 1.79, EL / PL: 0.97 – 1.06, EW / PW: 1.14 – 1.22; surface densely covered with setae and finely punctured; posterior margin sinuate near posterolateral corners. Hind wings well developed; flabellum with one seta. Mesoventrite without longitudinal carina; mesoventral process extended to slightly beyond middle of mesocoxal cavities, with rounded apex; metaventral process shorter than mesoventral process, with rounded apex; isthmus present. Abdomen slightly narrowed posteriad; surface densely covered with setae. Male. Elytra with a pair of small tubercles at about posterior 1 / 3 on sutural margin. Sternite VI with small medial lobe on posterior margin. Tergite VII with a tubercle on posteromedian area. Tergite VIII (Fig. 5 D) with a tubercle on median area; three to five processes on each side of posterior margin, outer ones slightly curved, longer and sharper than the others. Aedeagus as in Fig. 5 E; median lobe with large basal bulb; apical process broad, parallel to near apex, not dilated in lateral view, apical end thin and elongated, apex pointed; flagellum moderately long. Female. Spermatheca (Fig. 5 F) curved twice, with a transverse band-like structure at base of distal portion; distal portion elongated, U-shaped; median portion elongated, slightly curved; proximal portion longer than wide.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE78E17899AFCDBACE9FEC7.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, Kyushu) — new record; Taiwan.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE78E17899AFCDBACE9FEC7.taxon	discussion	Remarks. This species can be distinguished from Ps. crenulicauda by the non-dilated, parallel-sided apical process of its aedeagus. Most of the aedeagi of specimens from Japan have an abruptly narrowed and protruding apical part on their apical process, which is not seen in Pace’s illustrations of the specimen from Taiwan (2008). Similar aedeagi are seen in a few Japanese specimens, but it is likely that they were damaged. While we cannot rule out the possibility that the original description of Ps. taiwanensis was based on a specimen with a damaged apical process of aedeagus, we nonetheless identified the Japanese specimen as Ps. taiwanensis. The female morphology is not informative for distinguishing this species from Ps. crenulicauda and other relatives.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE98E14899AF8FFABFFFBB5.taxon	description	(Figs. 1 C, 6 A)	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE98E14899AF8FFABFFFBB5.taxon	distribution	type locality: “ Nikko ” [Nikkô-shi, Tochigi-ken, central Honshu]); Schülke & Smetana, 2015: 711 (catalogue).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE98E14899AF8FFABFFFBB5.taxon	materials_examined	Type material. Lectotype: here designated, female, “ Oxypoda hi- / laris Type / D. S. / Nikko. Japan. / 31.10.80. Lewis [handwritten, paper card] // Type [red round label pinned by a curator] // Japan. / G. Lewis. // Sharp Coll. / 1905 - 313. ” (abdominal segments VIII – X and spermatheca were dissected and glued on paper card, and labium and maxilla were mounted in Euparal by MM). (BMNH) Paralectotypes: 2 females, “ Kuro / matsu [handwritten, paper card, underside of label] // Japan. / G. Lewis. / 1910 - 320. // Oxypoda / hilaris [handwritten] ” (BMNH). See remarks.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE98E14899AF8FFABFFFBB5.taxon	description	Redescription. Measurements of lectotype of Oxypoda hilaris: BL ≈ 2.6; PL, 0.39; PW, 0.56; HTL, 0.40. Relative length of antennomeres I – XI (n = 1): 23: 24: 22: 10: 13: 13: 13: 12: 14: 13: 36. Ratio of length / width of antennomeres I – XI (n = 1): 1.95: 1.59: 1.96: 0.65: 0.70: 0.70: 0.67: 0.59: 0.64: 0.63: 1.64. Body (Fig. 1 C) reddish brown; head darker; posterolateral areas of elytra darker; abdominal segments V – VII darker. Head. Surface densely covered with setae. Antenna with antennomeres I – III and XI longer than wide, antennomeres IV – X wider than long; antennomeres I and II with rounded distal end, antennomeres III – X strongly angulated near distal end, antennomere XI oval. Pronotum transverse (PW / PL: 1.44); surface densely covered with setae, finely punctured; posterior margin slightly bisinuate. Elytra wider than long; surface densely covered with setae and finely punctured; posterior margin sinuate near posterolateral corners. Abdomen slightly narrowed posteriad; surface densely covered with setae. Male. Unknown. Female. Spermatheca (Fig. 6 A) curved twice, with a transverse band-like structure at base of distal portion; distal portion elongated, U-shaped; median portion elongated, slightly curved; proximal portion longer than wide.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE98E14899AF8FFABFFFBB5.taxon	distribution	Distribution. Japan (Honshu).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFE98E14899AF8FFABFFFBB5.taxon	discussion	Remarks. This species clearly does not belong to the genus Oxypoda, judging from the shape of the spermatheca, but to the genus Pseudatheta. As the type series of Oxypoda hilaris consists only of females, there is currently no method to distinguish this species from its allies. It is unclear whether this species is identical to Ps. crenulicauda, Ps. taiwanensis, both known from Japan, or neither. There is no reason to believe that the paralectotypes are the same species as the lectotype. Since this species was described in 1888 (hilaris is the oldest name), one or more of the similar species are likely to be junior synonyms of this species.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEA8E15899AFB6CABF8FDC3.taxon	description	(Figs. 1 D, 6 B – D)	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEA8E15899AFB6CABF8FDC3.taxon	distribution	type locality: “ Chiang Mai Prov., Doi Suthep ”); Pace, 2004: 66 (China, Shaanxi); Schülke & Smetana, 2015: 634 (catalogue).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEA8E15899AFB6CABF8FDC3.taxon	materials_examined	Material examined. THAILAND: 2 males, 1 female, Doi Inthanon, Chaing Mai, 1560 m, 20 X 1983, M. Sakai leg. (EUM). Additional description. Measurements (n = 1): BL ≈ 2.07; FBL: 1.22; HL: 0.36; HW: 0.43; PL: 0.52; PW: 0.60; EL: 0.43; EW: 0.72. Ratios (n = 1): HW / HL: 1.18, PW / PL: 1.44, EW / EL: 1.67, PW / HW: 1.39, EL / PL: 1.03, EW / PW: 1.20.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEA8E15899AFB6CABF8FDC3.taxon	description	Relative length of antennomeres I – XI (n = 1): 20: 24: 15: 10: 12: 10: 12: 12: 12: 11: 34. Ratio of length / width of antennomeres I – XI (n = 1): 1.79: 2.36: 1.50: 0.68: 0.76: 0.65: 0.66: 0.63: 0.58: 0.55: 1.61. Male. Elytra with a pair of small tubercles at about posterior 1 / 3 on sutural margin. Sternite VI with small medial lobe on posterior margin. Tergite VII with a tubercle on posteromedian area. Tergite VIII (Fig. 6 B) with a tubercle on median area; four processes on each side of posterior margin, outer ones slightly curved, longer and sharper than the others. Aedeagus as in Fig. 6 C; median lobe with large basal bulb; apical process relatively slender, narrowed apically in lateral view, apex pointed; flagellum moderately long.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEA8E15899AFB6CABF8FDC3.taxon	distribution	Distribution. Thailand, China (Shaanxi).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEA8E15899AFB6CABF8FDC3.taxon	discussion	Remarks. This species was described only from a female and could not be distinguished from its allied species based on the spermatheca. According to the original description by Pace (1992), the median part of the spermatheca is short, but this description may have derived from a deformed specimen. In many aleocharine species, spermathecae are easily deformed immediately after mounting in mounting medium. Later, Pace (2004) recorded this species from China, but it was again a female specimen. In this study, we were able to examine specimens of both sexes of the genus Pseudatheta, which were similar to Ps. thailandensis collected in an area close to the type locality of Ps. thailandensis. Since there are areas where more than one species of this genus is distributed, such as Japan, whether these specimens are Ps. thailandensis is unclear. Nonetheless, since this species is the only known thailandensis - like species from Thailand, we tentatively identified the specimens as Ps. thailandensis and described the aedeagus. This species can be distinguished from its allies (Ps. crenulicauda, Ps. taiwanensis) by the apical process of the aedeagus, which tapers gradually toward the apex. However, Ph. bigranipennis (Bernhauer, 1915), redescribed by Benick (1980) and Kim & Ahn (2016), cannot be distinguished from Ps. thailandensis judging from the illustrations and description, and specimens described by the authors are considered identical. Since Benick (1980) and Kim & Ahn (2016) did not state that they examined the type specimen, we cannot rule out the possibility of a misidentification. Thus, we refrain from taxonomic treatment of Ph. bigranipennis at this time. The relationship between Ps. thailandensis and Ps. hilaris could not be determined.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEB8E12899AFDB0AB3FFE0B.taxon	description	(Figs. 1 E, 7 A – D)	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEB8E12899AFDB0AB3FFE0B.taxon	vernacular_names	[Japanese name: Minami-hime-kinokotsuyakeshi-hanekakushi]	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEB8E12899AFDB0AB3FFE0B.taxon	distribution	type locality: “ China, Jiangsu Prov., Nanjing Zijinshan ”): Schülke & Smetana, 2015: 634 (catalogue); Pace, 2016: 301 (Yunnan).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEB8E12899AFDB0AB3FFE0B.taxon	materials_examined	Material examined. JAPAN: [Kagoshima-ken]: Yaku-shima Is.: 1 male, Aiko-dake, Yakushima-chô, 25 IX – 23 X 2006, T. Yamauchi et al. leg, (KUM); 1 male, 2 females, Kurio, Yakushima-chô, 21 VII 2021, T. Hashizume leg. (KUM); [Okinawa-ken]: Okinawa-jima Is.: 2 males, Takazato, Ôgimi-son, 3 XI 1993, M. Kimura leg. (KUM); 2 males, Yona, 25 – 27 V 1974, M. Sato leg. (KUM); Iriomote-jima Is.: 2 males, 2 females, Kampiree, 27 III 1984, S. Nomura leg. (KUM). TAIWAN: 3 males, Wulai, Taipei Hsien, 17 V 1972, M, Sakai leg. (EUM).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEB8E12899AFDB0AB3FFE0B.taxon	description	Redescription. Measurements (n = 5): BL ≈ 1.65 – 1.84; FBL, 0.91 – 1.00; HL, 0.28 – 0.34; HW, 0.35 – 0.39; PL, 0.31 – 0.35; PW, 0.46 – 0.50; EL, 0.30 – 0.35; EW, 0.55 – 0.60. Relative length of antennomeres I – XI (n = 1): 25: 27: 19: 10: 10: 10: 11: 11: 13: 13: 35. Ratio of length / width of antennomeres I – XI (n = 1): 1.94: 2.21: 1.80: 0.87: 0.66: 0.64: 0.59: 0.58: 0.63: 0.61: 1.62. Body (Fig. 1 E) reddish brown; head darker; elytra dark brown except for the anterior 1 / 4 – 1 / 3; abdominal segments V – VI darker. Head slightly transverse, HW / HL: 1.12 – 1.36; surface densely covered with setae. Antenna with antennomeres I – III and XI longer than wide, antennomeres IV – X wider than long; antennomeres I and II with rounded distal end, antennomeres III – X strongly angulated near distal end, antennomere XI oval. Pronotum transverse, PW / PL: 1.43 – 1.50, PW / HW: 1.29 – 1.33; surface densely covered with setae, finely punctured, without microsculpture; posterior margin arcuate. Elytra wider than long, EW / EL: 1.71 – 1.84, EL / PL: 0.92 – 1.00, EW / PW: 1.13 – 1.24; surface densely covered with setae and finely punctured; posterior margin sinuate near posterolateral corners. Mesoventrite without longitudinal carina; mesoventral process extended to slightly beyond middle of mesocoxal cavities, with pointed apex; metaventral process shorter than mesoventral process, with rounded apex; isthmus absent. Abdomen slightly narrowed posteriad; surface densely covered with setae. Male. Elytra with a pair of small tubercles at about posterior 1 / 3 on sutural margin. Tergite VII with a tubercle on anteromedian area. Tergite VIII (Fig. 7 A) with a tubercle on median area; six processes on posterior margin. Aedeagus as in Fig. 7 C; median lobe with large basal bulb; apical process elongated, simply curved ventrally, slightly widened at basal part; ventral process large, apex pointed, with a distinct angle on dorsa side; flagellum very long. Female. Spermatheca (Fig. 7 D) curved twice; distal portion large, nearly conical shape, slightly curved; median portion elongate, curved; proximal portion very small.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEB8E12899AFDB0AB3FFE0B.taxon	distribution	Distribution. Japan (Yaku-shima Is., Okinawa-jima Is., Iriomote-jima Is.), Taiwan — new record; China (Jiangsu, Yunnan).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEB8E12899AFDB0AB3FFE0B.taxon	discussion	Remarks. We report the first discovery of males of Ps. cooteri and describe them here. Variations in the shape of the ventral process of the median lobe of the aedeagus were observed (Fig. 7 B). The aedeagus and spermatheca of Ps. spinosa, a species from Taiwan described by Pace (2008), are similar to those of Ps. cooteri. However, judging from the illustrations in that report, Ps. cooteri can be distinguished from Ps. spinosa by the longer apical process of the median lobe of its aedeagus and by the longer, weakly curved distal portion of the spermatheca. We were able to examine specimens of Pseudatheta from Taiwan and found that the aedeagi were identical to those of Ps. cooteri from Japan. We consider the former specimens to be Ps. cooteri from Taiwan, although we were unable to compare the spermathecae because females were not collected with the male specimens. While it is possible that the two species are distributed in Taiwan, the slight differences may have instead been due to intraspecific variation, deformation of the specimens, or misinterpretations in sketching. Therefore, we cannot rule out the possibility that these two species are synonyms.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFED8E10899AFF4DA98FFB1F.taxon	description	(Figs. 1 F, 3, 7 E – G)	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFED8E10899AFF4DA98FFB1F.taxon	vernacular_names	[Japanese name: Ko-hime-kinokotsuyakeshi-hanekakushi]	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFED8E10899AFF4DA98FFB1F.taxon	distribution	type locality: “ Bukit Timah, woodlands, Mandai ” [Singapore]): Cameron, 1939: 226 (Northern India); Schülke & Smetana, 2015: 634 (catalogue).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFED8E10899AFF4DA98FFB1F.taxon	materials_examined	Material examined. Type materials compared. Syntypes: 1 male, “ Fungus // Mandai, / Singapore. / Dr. Cameron. // SYN- / TYPE [blue round label pinned by a curator] // Pseudatheta / elegans Cam. / M. E. Bacchus det 1978 / SYNTYPE // Pseudatheta / elegans CAM. / det. R. Pace 1984 / LECTOTYPUS // NHMUK 015025222 ” (BMNH); 1 male, “ Fungus / (rotten) // Mandai, / Singapore. / Dr. Cameron. // SYN- / TYPE [blue round label pinned by a curator] // Pseudatheta / elegans Cam / M. E. Bacchus det 1978 / SYNTYPE // Pseudatheta / elegans CAM. / det. R. Pace 1984 / PARALECTOTYPUS // NHMUK 015020370 ” (BMNH); 1 male, “ SYN- / TYPE [blue round label pinned by a curator] // Mandai, / Singapore. / Dr. Cameron // TYPE / Pseudatheta / elegans. / Dr. CAMERON. // Pseudatheta / elegans Cam. / M. E. Bacchus det 1978 / SYNTYPE // Pseudatheta / elegans CAM. / det. R. Pace 1984 / PARALECTOTYPUS / Dissection By? // NHMUK 015020337 ” (BMNH): 1 male, “ Fungus / (rotten) // Woodlands, / Singapore. / Dr. Cameron. // SYN- / TYPE [blue round label pinned by a curator] // Pseudatheta / elegans CAM. / det. R. Pace 1984 / PARALECTOTYPUS // Pseudatheta / elegans Cam. / M. E. Bacchus det 1978 / SYNTYPE / NHMUK 015025223 ” (BMNH); 1 female, “ Wood / (rotten) // Bukit Timah, / Singapore. / Dr. Cameron. / SYN- / TYPE [blue round label pinned by a curator] // Pseudatheta / elegans Cam. / M. E. Bacchus det 1978 / SYNTYPE // Pseudatheta / elegans CAM. / det. R. Pace 1984 / PARALECTOTYPUS // NHMUK 015025224 ” (BMNH) Additional materials studied. JAPAN: [Miyagi-ken]: 1 male, Yoshida, Taiwa-chô, 20 VIII 2009, J. Aoki leg. (KUM); [Fukuoka-ken]: 1 male, Tonoue-yama, Moji-ku, Kitakyûshû-shi, 15 V 2022, J. - H. Park leg. (KUM); 1 male, 1 female, Hiko-san, Soeda-machi, 24 VII 2020, T. Nozaki leg. (KUM); 1 male, Ryôgenji, Munakata-shi, 15 V 2022, T. Hashizume leg. (KUM); [Saga-ken]: 1 male, 2 females, Uki-dake, Nanayama, Karatsu-shi, 6 X 2022, N. Tsuji leg. (KUM); [Nagasaki-ken]: Tsushima Is.: 1 male, 1 female, Kamiagatamachi Sagohigashisato, Tsushima-shi, 18 VI 2022, T. Hashizume leg. (KUM); 1 male, Kamiagatamachi Sasuna, 19 VI 2022, T. Hashizume leg. (KUM); Fukue-jima Is.: 1 female, Kishikumachi Nakadake, Gotô-shi, 11 X 2022, T. Hashizume leg. (KUM); [Oita-ken]: 1 male, Takatoriya-yama, Minamitabaru, Ume, Saeki-shi, 14 XI 2020, S. Inoue leg. (KUM); [Kagoshima-ken]: 1 female, Sata-misaki, 5 V 1985, Y. Takai leg. (KUM); Yaku-shima Is.: 2 males, 1 female, Koseda, Yakushima-chô, 22 VII 2021, T. Hashizume leg. (KUM); Amami-Ôshima Is.: 1 male, Sumiyôchô Nishinakama, Amami-shi, 27 VI 2021, T. Hashizume leg. (KUM); 1 male, Shinmura, 11 IV 1971, M. Sakai leg. (EUM); Tokuno-shima Is.: 1 female, Mikyo, 12 IV 1968, M. Tomokuni leg. (EUM); [Okinawa-ken] Ishigaki-jima Is.: 1 male, Omoto-dake, Hirae, Ishigaki-shi, 24 III 2022, S. Inoue leg. (KUM).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFED8E10899AFF4DA98FFB1F.taxon	description	Redescription. Measurements (n = 5): BL ≈ 1.49 – 1.94; FBL, 0.86 – 0.96; HL, 0.29 – 0.30; HW, 0.34 – 0.36; PL, 0.30 – 0.34; PW, 0.44 – 0.48; EL, 0.30 – 0.35; EW, 0.51 – 0.56. Relative length of antennomeres I – XI (n = 1): 23: 25: 18: 11: 10: 11: 10: 10: 11: 11: 31. Ratio of length / width of antennomeres I – XI (n = 1): 1.88: 2.35: 2.10: 0.96: 0.77: 0.75: 0.96: 0.58: 0.63: 0.57: 1.46. Body (Fig. 1 F) reddish brown; head darker; elytra dark brown except for the anterior 1 / 4 – 1 / 3; abdominal segments V – VI darker. Head almost as long as wide, HW / HL: 1.17 – 1.21; surface densely covered with setae. Antenna with antennomeres I – III and XI longer than wide, antennomeres IV – X wider than long; antennomeres I and II with rounded distal end, antennomeres III – X strongly angulated near distal end, antennomere XI oval. Pronotum transverse, PW / PL: 1.30 – 1.46, PW / HW: 1.30 – 1.31; surface densely covered with setae, finely punctured, without microsculpture; posterior margin arcuate. Elytra wider than long, EW / EL: 1.54 – 1.71, EL / PL: 1.00 – 1.04, EW / PW: 1.17 – 1.29; surface densely covered with setae and finely punctured; posterior margin sinuate near posterolateral corners. Hind wings well developed; without flabellum. Mesoventrite without longitudinal carina; mesoventral process extended to slightly beyond middle of mesocoxal cavities, with pointed apex; metaventral process shorter than mesoventral process, with rounded apex; isthmus absent. Abdomen slightly narrowed posteriad; surface densely covered with setae. Male. Elytra with a pair of small tubercles at about posterior 1 / 3 on sutural margin. Tergite VII with a tubercle on anteromedian area. Tergite VIII (Fig. 7 E) with a tubercle on median area; eight to ten processes on posterior margin. Aedeagus as in Fig. 7 F; median lobe with large basal bulb; apical process elongated, sinuated, apical half slightly widened; ventral process narrow, sinuated; flagellum moderately long. Female. Spermatheca (Fig. 7 G) curved twice; distal portion nearly reniform; median portion elongate, curved; proximal portion round, wider than long; constriction between median and proximal portion distinct.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFED8E10899AFF4DA98FFB1F.taxon	distribution	Distribution. Japan (Honshu, Kyushu, Tsushima Is., Gotô-rettô Isls. (Fukue-jima Is.), Yaku-shima Is., Amami-Ôshima Is., Tokuno-shima Is., Ishigaki-jima Is.) — new record; Singapore, India.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFED8E10899AFF4DA98FFB1F.taxon	discussion	Remarks. We compared photographs of the habitus and the aedeagi of the type series with those of the specimens collected from Japan, and determined them to be the same species based on the similarity of dorsal habitus, the slightly sinuate apical process of the median lobe of aedeagus, and moderately long flagellum of the aedeagus. This species is similar in the shapes of the aedeagus and the spermatheca to Ps. tronqueti, described from Sri Lanka by Pace (1988), Ps. mendica, described from Nepal by Pace (1989), Ps. rougemonti, described from Sabah in East Malaysia by Pace (2014), Ps. borneensis, described from Sabah by Pace (2007), Ps. kinabaluensis, described from Sabah by Pace (2007), and Ps. pahangensis, described from Peninsular Malaysia by Pace (2012). These similar species, except for Ps. mendica, can be distinguished from Ps. elegans as follows: in Ps. rougemonti, the ventral process of the median lobe of aedeagus is almost straight;	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFED8E10899AFF4DA98FFB1F.taxon	description	in Ps. borneensis the flagellum of the median lobe of aedeagus is longer, and the proximal portion of the spermatheca is slightly smaller; in Ps. kinabaluensis, the ventral process of the median lobe of aedeagus is simply slightly curved dorsally, and the shape of the proximal portion of the spermatheca is different; in Ps. tronqueti, the apical process of the median lobe of aedeagus is almost straight, and the proximal portion of the spermatheca is slightly smaller; in Ps. pahangensis, the distal portion of the spermatheca is slightly broader and shorter. It is desirable to find males of Ps. pahangensis because the spermatheca is very similar to that of Ps. elegans. Unfortunately, Ps. mendica cannot be distinguished from the Japanese Ps. elegans based on the information in the original description based on the female holotype (Pace, 1989). Some of species compared here are difficult to distinguish and we cannot refute the possibility that they are geographic or individual variation of this species. In fact, the aedeagus of the Pseudatheta sp. from Taiwan we examined is almost identical to those of specimens from Japan, but the distal portion of the spermatheca is slightly shorter. In this study, we were not able to examine the spermatheca of the type series of Ps. elegans and it is desirable to do so in the future.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEE8E10899AFA8DAD17F82E.taxon	vernacular_names	[Japanese name: Hime-tobimushi-hanekakushi]	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEE8E10899AFA8DAD17F82E.taxon	discussion	(transferred to the genus Platyola from the genus Phymatura).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEE8E10899AFA8DAD17F82E.taxon	discussion	(transferred to the genus Pseudatheta from the genus Phymatura; probably based on Sawada (1977 )); Schülke & Smetana, 2015: 634 (catalogue). nec. Phymatura oligotinula sensu Sawada, 1977: 217 (transferred to the genus Phymatura from the genus Homalota).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEE8E10899AFA8DAD17F82E.taxon	materials_examined	Material examined. Type material. Lectotype: here designated, female, “ Homalota / oligotinula / Type D. S. / Kumamoto / 26. IV. 81. Lewis [paper card, handwritten] // SYN- / TYPE [blue round label pinned by a curator] // Type [red round label pinned by a curator] // Japan. / G. Lewis. // Kumamoto / 23. IV. - 26. IV. 81. // Sharp Coll. / 1905 - 313 // Platyola / oligotinula (SH.) // det. R. Pace, 1982 ” (spermatheca is mounted in glycerin by previous researcher) (BMNH).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEE8E10899AFA8DAD17F82E.taxon	discussion	Remarks. This species was redescribed based on misidentified specimens of Ps. crenulicauda by Sawada (1977) and transferred to the genus Phymatura. However, as Pace (1984) stated that this species belongs to the genus Platyola of the tribe Athetini, not Homalotini. Later, Pace (1992) probably forgot this arrangement (Pace, 1984) and moved this species from the genus Phymatura to the genus Pseudatheta, and this treatment was followed in the later catalogs (e. g., Schülke & Smetana 2015; Newton 2022). We here place this species again in the genus Platyola.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E11899AFF4DADE7FD4A.taxon	vernacular_names	[Japanese name: Kinokotsuyakeshi-hanekakushi-zoku]	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E11899AFF4DADE7FD4A.taxon	type_taxon	type species: Bolitochara brevicollis Kraatz, 1856, fixed by subsequent designation by Casey, 1906: 264).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E11899AFF4DADE7FD4A.taxon	synonymic_list	type species: Venusa picta Casey, 1906, fixed by subsequent designation by Fenyes, 1918: 26); Ashe, 1992: 371 (as a synonym of Phymatura).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E11899AFF4DADE7FD4A.taxon	discussion	Remarks. The genus Phymatura can be distinguished from the genus Pseudatheta by the following character states: mesoventrite with a longitudinal carina; flabellum of hind wing with at least three setose lobes.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E0F899AFD39AB80FC37.taxon	description	(Figs. 2 A, E, 8 A – C)	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E0F899AFD39AB80FC37.taxon	vernacular_names	[Japanese name: Nihon-kinokotsuyakeshi-hanekakushi]	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E0F899AFD39AB80FC37.taxon	distribution	type locality; “ Chuzenji ” [Ch ̊ zenji, Nikkô-shi, Tochigi-ken, central Honshu]); Sawada, 1977: 217 (as a synonym of Ps. oligotinula); Schülke & Smetana, 2015: 633 (catalogue, as a synonym of Ph. crenulicauda).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E0F899AFD39AB80FC37.taxon	distribution	type locality: “ Bai He [Baihe], Jilin Prov., China ”); Schülke & Smetana, 2015: 633 (catalogue); Assing, 2021: 330 (Japan: Amami-Ôshima Is.; Russia). Syn. nov.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E0F899AFD39AB80FC37.taxon	materials_examined	Material examined. Type material. Phymatura japonica: Holotype: male, “ Holo- / type [red round label pinned by a curator] // JAPAN / CHUZENJI // AT FT J. E. A. Lewis // Cameron / Bequest. / B. M. 1955 - 147. // Phymatura / japonica / TYPE Cam. [handwritten] // Phymatura / japonica Cam. / P. M. Hammond / det. 1973 / HOLOTYPE ” (abdominal segments VIII – X and aedeagus are dissected and mounted in Euparal by MM) (BMNH). Additional materials studied. JAPAN: [Hokkaido]: 1 ex., Akan-chô, 2 VIII 1988, K. Haga leg. (KUM); 1 ex., Akan N. Park, 6 VII 1958, M. Miyatake leg. (EUM); 1 male, Taiki-chô, 2 IX 1990, K. Haga leg. (KUM); 7 exs., Sounkyô, 17 – 18 VII 1970, S. Kinoshita leg. (EUM); [Fukushima-ken]: 1 male, 3 females, Shindenpara, Minamiaizu-machi, 13 VII 1996, K. Haga leg. (KUM); [Saitama-ken]: 1 male, 3 females, Mitsumine, Chichibu-shi, 28 V 1997, M. Maruyama leg. (KUM); 10 males, 10 females, Taki-gawa, Chichibu-shi, 30 V 1997, M. Maruyama leg. (KUM); [Tokyo-to]: 1 male, 1 female, Nippara, Okutama-machi, 8 VI 1991, K. Haga leg. (KUM): 1 female, same data, but 9 VI 1991 (KUM); 1 male, 1 female, Nippara, Okutama-machi, 7 V 1997, T. Shimada leg. (KUM); [Kanagawa-ken]: 1 male, 2 females, Hakone, 1 V 1974, Y. Hirano leg. (KUM); 1 female, Nishi Tanzawa, 8 IX 1984, Y. Hirano leg. (KUM); [Niigata-ken]: 1 female, Yuzawa-machi, 1 X 1995, K. Haga leg. (KUM); [Yamanashiken]: 1 male, Sentouboshi-yama, Minaniarupusu-shi, 2 VIII 1992, K. Haga leg. (KUM); [Shizuoka-ken]: 1 female, Sessokyo, Kawanehon-chô, 9 – 10 V 1992, T. Kishimoto leg. (TKc); 1 femle, Amagi-tôge, Kawazu-chô, 14 – 16 III 1996, S. Naomi leg. (KUM); [Nara-ken]: 1 female, Ôdaigahara, 29 V 1985, S. Nomura leg. (KUM); [Hiroshimaken]: 1 male, Kure-shi, 2 XII 1995, I. Okamoto leg. (KUM); [Tokushima-ken]: 1 female, Tsurugi-san, 27 VII 1961, M. T. Chûjô leg. (KUM); 5 males, 2 females, same data, but 28 VII 1961 (KUM); 1 ex., Tsurugi-san, 11 VII 1993, M. Sakai leg. (EUM); 1 male, Iyadani, Tsurugi-san, 19 VIII 1989, K. Matsumoto leg. (KUM); 1 female, Shikibidani, Naka-chô, 17 VII 2010, J. Aoki leg. (KUM); [Ehime-ken]: 1 ex., Omogo-kei, 2 V 1976, A. Oda leg. (EUM); 1 ex., Omogo-kei, 4 VI 1983, K. Ishida leg. (EUM); 1 ex., Banshodani, Omogo, 11 VI 1988, M. Sakai leg. (EUM); 1 ex., Ôhira, Tôon-shi, 27 IV 2007, T. Kitano leg. (EUM); [Fukuoka-ken]: 1 female, Tachibana-yama, Fukuoka-shi, 27 XI 1983, S. Naomi leg. (KUM); 1 male, 1 female, Hiko-san, 24 IX 1938, Hori & Fujino leg. (KUM); 2 males, Hiko-san, 7 V 1971, K. Takano leg. (KUM); 1 female, Hiko-san, Soeda-machi, 23 V 2020, S. Inoue leg. (KUM); [Saga-ken]: 1 female, Sefuri-san, 3 X 1976, H. Oishi leg. (KUM); [Kumamoto-ken]: 1 male, 1 female, Izumimachi Momigi, 14 V 2021, S. Inoue leg. (KUM); 1 male, Gokanoshô, 17 X 1988, S. Naomi leg. (KUM); 4 males, 8 females, Asoshi, 3 X 1995, Y. Tomishima leg. (KUM); 1 male, Ichifusa-yama, Mizukami-mura, 13 X 1995, Y. Tomishima leg. (KUM); 2 exs., Ichifusa-yama, 11 VI 1972, S. Hisamatsu leg. (EUM); 1 male, Kirihagi, Yamato-chô, 15 X 1993, Y. Tomishima leg. (KUM); [Oita-ken]: 1 male, Shônaichô Asono, Yufu-shi, 20 V 2021, T. Hashizume leg. (KUM); [Kagoshima-ken]: Amami-Ôshima Is.: 12 exs., Hatsuno, 2 V 1977, A. Oda leg. (EUM); 3 exs., same data, but M. Sakai & A. Oda leg. (EUM); 1 ex., same data, but M. Sakai leg. (EUM); 3 exs., Yuwan, 3 IV 1968, M. Tomokuni leg. (EUM); Tokuno-shima Is.: 6 exs., Mikyo, 12 IV 1968, M. Tomokuni leg. (EUM).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E0F899AFD39AB80FC37.taxon	description	Redescription. Measurements (n = 5): BL ≈ 2.69 – 3.05; FBL, 1.35 – 1.65; HL, 0.38 – 0.46; HW, 0.45 – 0.51; PL, 0.43 – 0.51; PW, 0.58 – 0.69; EL, 0.49 – 0.61; EW, 0.75 – 0.86. (Holotype: BL ≈ 3.2; PL, 0.46; PW, 0.63; HTL, 0.51) Relative length of antennomeres I – XI (n = 1): 21: 21: 19: 10: 13: 12: 13: 13: 12: 14: 31. Ratio of length / width of antennomeres I – XI (n = 1): 1.88: 2.22: 1.94: 0.82: 0.85: 0.71: 0.76: 0.77: 0.68: 0.73: 1.58. Body (Fig. 2 A, D) relatively large. Head dark brown; pronotum brown; posterolateral areas of elytra dark brown, anterior 1 / 4 – 1 / 3, sutural area, and narrow area of posterior margin yellowish brown; abdomen reddish brown, around midline of tergite III – V dark brown, segments VI – VII dark brown. Head almost as long as wide, HW / HL: 1.11 – 1.20; surface densely covered with setae. Antenna with antennomeres I – III and XI longer than wide, antennomeres IV – X wider than long; antennomeres I and II with rounded distal end, antennomeres III – X strongly angulated near distal end, antennomere XI oval. Pronotum transverse, PW / PL: 1.32 – 1.35, PW / HW: 1.28 – 1.34; surface densely covered with setae, finely punctured, without microsculpture; posterior margin slightly bisinuate. Elytra wider than long, EW / EL: 1.34 – 1.54, EL / PL: 1.12 – 1.21, EW / PW: 1.18 – 1.30; surface densely covered with setae and finely punctured; posterior margin sinuate near posterolateral corners. Hind wings well developed; flabellum with several setae. Mesoventrite with short longitudinal carina from anterior margin; mesoventral process extended to slightly beyond middle of mesocoxal cavities, with pointed apex; metaventral process shorter than mesoventral process, with rounded apex; listhmus absent. Abdomen slightly narrowed posteriad; surface densely covered with setae. Male. Tergite VII with a tubercle on posteromedian area. Tergite VIII (Fig. 8 A) broadly emarginate; one tubercle on median area; eight to ten small processes on posterior margin. Aedeagus as in Fig. 8 B; median lobe with relatively narrow basal bulb; apex of apical process pointed; flagellum well sclerotized, very long. Female. Spermatheca as in Fig. 8 C; distal portion large; wall of distal end protrudes inward; median portion and proximal portion small.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E0F899AFD39AB80FC37.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Amami-Ôshima Is., Tokuno-shima Is.); China (Jilin, Heilongjiang), North Korea, Russia.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFEF8E0F899AFD39AB80FC37.taxon	discussion	Remarks. There is only the male holotype of Ph. japonica in BMNH. The original description of Phymatura cooteri agrees with the feature of the type specimen of Ph. japonica in the shapes of the tergite VIII, sternite VIII and the aedeagus. Therefore, Ph. cooteri is here synonymized with Ph. japonica. Assing (2021) recorded this species from Southwestern Japan (Amami-Ôshima) and Russia. Paśnik (2001) recorded this species from North Korea (see Assing (2005) for more detailed description of Ph. japonica (as Ph. cooteri )).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFF18E0F899AFBE4ACA2F937.taxon	description	(Figs. 2 B, 8 D – F)	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFF18E0F899AFBE4ACA2F937.taxon	vernacular_names	[Japanese name: Aka-kinokotsuyakeshi-hanekakushi]	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFF18E0F899AFBE4ACA2F937.taxon	distribution	type locality: “ CHINA: S-Shaanxi (Qinling Shan), pass on rd. Zhouzhi — Foping, 105 km SW Xi′an, N-slope, 1990 m, 33 ° 44 ′ N, 107 ° 59 ′ E ”).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFF18E0F899AFBE4ACA2F937.taxon	materials_examined	Material examined. JAPAN: [Hokkaido]: 1 female, Nopporo-shinrin-kôen, Ebetsu-shi, 16 – 19 VI 2001, S. Hori leg. (KUM); 1 ex., Kamibisei, Memuro-chô, 25 VIII 1995, M. Sakai leg. (EUM); [Ehime-ken]: 2 males, Komenomachi, Matsuyama-shi, 8 V 1999, T. Kan leg. (EUM).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFF18E0F899AFBE4ACA2F937.taxon	description	Measurements. (n = 3): BL ≈ 1.98 – 2.31; FBL: 1.08 – 1.19; HL: 0.31 – 0.36; HW: 0.35 – 0.36; PL: 0.38 – 0.40; PW: 0.48; EL: 0.39 – 0.41; EW: 0.59 – 0.62. Ratios (n = 3): HW / HL: 1.04 – 1.13, PW / PL: 1.20 – 1.28, EW / EL: 1.50, PW / HW: 1.32 – 1.37, EL / PL: 1.00 – 1.03, EW / PW: 1.22 – 1.29. Relative length of antennomeres I – XI (n = 1): 26: 25: 15: 10: 12: 11: 12: 12: 13: 13: 30. Ratio of length / width of antennomeres I – XI (n = 1): 2.09: 2.08: 1.21: 0.71: 0.75: 0.62: 0.60: 0.55: 0.61: 0.55: 1.46.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFF18E0F899AFBE4ACA2F937.taxon	distribution	Distribution. Japan (Hokkaido, Shikoku) — new record; China (Shaanxi).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
090C879DFFF18E0F899AFBE4ACA2F937.taxon	discussion	Remarks. Phymatura russa is recorded from Japan for the first time. This species can be distinguished from the other Phymatura species recorded from Japan by its nearly uniformly reddish coloration, the smaller eyes, and the shapes of the aedeagus and the spermatheca. This species has been collected from Hokkaido and Shikoku, and may be widely distributed in mainland Japan (see Assing (2021) for detailed description of Ph. russa).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 5319 (1): 27-47, DOI: 10.11646/zootaxa.5319.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.2
