identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F362FB0FF6C45C70B37002DBB5C72348.text	F362FB0FF6C45C70B37002DBB5C72348.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nagiella inferior (Hampson 1899)	<div><p>Nagiella inferior (Hampson, 1899) Figures 2C, D, 4A-C, 7A-C</p><p>Botys quadrimaculalis Motschulsky, 1861: 1: 37 (preoccupied).</p><p>Pleuroptya quadrimaculalis: Bae, 2001: 122-124, pl. 5 fig. 172; Park et al. 2007: 177, 239 fig. 94; Bae et al. 2008: 148-149, fig. 167.</p><p>Sylepta inferior Hampson, 1899: 724; Shibuya 1928: 228; Shibuya 1929: 189.</p><p>Nagia inferior: Mutuura, 1957: 122, pl. 21 fig. 636.</p><p>Nagiella inferior: Munroe, 1976: 878; Lu and Du 2020: 149, fig. 5, figs 5, 10, 13.</p><p>Pleuroptya inferior: Inoue, 1982: 1: 343; 2: 234, 454, pl. 40, fig. 16; Li et al. 2012: 625-626, pl. 18, fig. 416; Sasaki and Yamanaka 2013: 81, 451.</p><p>Material examined.</p><p>Japan: 1♂, Tohro, Hokkaido, 3 Jul. 1962, T. Ebato leg., (NSMT-I-L-75498) ; 1♂, Riv. Rusagawa, Shiretoko, Hokkaido, 26 Jul. 1962, K. Tsuchiya leg. (NSMT-I-L-75523) ; 1♂, Sapporo, Hokkaido, 6 Jul. 1933, collector unknown (NSMT-I-L-75529) ; 1♂, Shumarinai, Hokkaido, 20 Jul. 1998, Y. Kishida leg. (NSMT-I-L-75569) ; 1♂, Shibecha, Hokkaido, 8 Jul. 1958, K. Jinbo leg. (NSMT-I-L-75574) ; 2♂, Akan, Hokkaido, 13 Jul. 1958, K. Jinbo leg. (NSMT-I-L-75575, 75576) ; 1♂, Toubai, Nemuro City, Hokkaido, 5 Aug. 2013, U. Jinbo leg. (NSMT-I-L-37577) ; 1♂, Sannai-Ishizawa, Honjoh City, Akita Pref., 29 Jun. 1975, A. Sasaki leg. (NSMT-I-L-75548) ; 1♂, ditto, 29 Jun. 1978, A. Sasaki leg. (NSMT-I-L-75545); 2♂, ditto, 15 Jun. 1979, A. Sasaki leg. (NSMT-I-L-75546, 75547); 1♀, Shinzan Park, Honjoh City, Akita Pref., 11 Jul. 1977, A. Sasaki leg. (NSMT-I-L-75549) ; 1♂, Uwanodai, Kawabe Town, Akita Pref., 5 Jul. 1977, A. Sasaki leg. (NSMT-I-L-75555) ; 1♀, ditto, 12 Jul. 1977, A. Sasaki leg. (NSMT-I-L-75551); 1♀, ditto, 26 Jun. 1978, A. Sasaki leg. (NSMT-I-L-75552); 1♀, ditto, 19 Jul. 1979, A. Sasaki leg.; 1♂, ditto, 19 Jul. 1979, A. Sasaki leg. (NSMT-I-L-75556); 1♂, ditto, 11 Jun. 1980, A. Sasaki leg. (NSMT-I-L-75550); 1♂, ditto, Akita Pref., 2 Jul. 1980, A. Sasaki leg. (NSMT-I-L-75553); 1♀, Kamibiguchi, Gojohme Town, Akita Pref., 26 Jul. 1979, A. Sasaki leg. (NSMT-I-L-75557) ; 1♂, Tazawako Height, Tazawako Town, Akita Pref., 20 Aug. 1979, A. Sasaki leg. (NSMT-I-L-75558) ; 1♂, Chikogi-zaki, Hachimori Town, Akita Pref., 13 Jul. 1979, A. Sasaki leg. (NSMT-I-L-75559) ; 1♀, Niida, Akita City, Akita Pref., 16 Jun. 1978, A. Sasaki leg. (NSMT-I-L-75560) ; 1♂, Asahimata, Akita City, Akita Pref., 16 Jul. 1980, A. Sasaki leg. (NSMT-I-L-75561) ; 1♂, Nibetsu, Akita City, Akita Pref., 3 Jul. 1980, A. Sasaki leg. (NSMT-I-L-75562) ; 1♂, ditto, 23 Jun. 1987, A. Sasaki leg. (NSMT-I-L-75563); 1♂, Mt Takao, Yuma Town, Akita Pref., 8 Jul. 1985, A. Sasaki leg. (NSMT-I-L-75564) ; 1♂, Tōshi, Nikaho Town, Akita Pref., 18 Jul. 1995, A. Sasaki leg. (NSMT-I-L-75565) ; 1♂, ditto, 19 Aug. 1984, A. Sasaki leg. (NSMT-I-L-75566); 1♂, Matsusaka, Ōno-dai, Moriyoshi Town, Akita Pref., 13 Jul. 1996, A. Sasaki leg. (NSMT-I-L-75567) ; 1♀, Aburato, Tsuruoka City, Yamagata Pref., 11 Jul. 1990, A. Sasaki leg. (NSMT-I-L-75568) ; 1♂, Futamata-Spa, Fukushima Pref., 6 Aug. 1967, T. Ebato leg. (NSMT-I-L-75497) ; 1♂, Shiozawa-Spa, Fukushima Pref., 5 Aug. 1967, T. Ebato leg. (NSMT-I-L-75501) ; 1♂, ditto, 6 Aug. 1967, T. Ebato leg. (NSMT-I-L-75519); 1♂, ditto, 29 Jun. 1968, T. Ebato leg. (NSMT-I-L-75518); 2♂, Hanashiki-Spa, Gunma Pref., 22 Jun. 1963, T. Ebato leg. (NSMT-I-L-75459, 75460) ; 1♀, same data as for preceding, (NSMT-I-L-75487); 1♂, ditto, 8 Jul. 1961, T. Ebato leg. (NSMT-I-L-75492); 3♂, Uenohara, Gunma Pref., 8 Jul. 1961, T. Ebato leg. (NSMT-I-L-75488 to 75490) ; 1♂, Kawaburu-Spa, Gunma Pref., 1 Jul. 1967, T. Ebato leg. (NSMT-I-L-75491) ; 1♂, Kitakaruizawa, Gunma Pref., 12 Jul. 1970, T. Okada leg. (NSMT-I-L-75528) ; 1♂, Minakami, Gunma Pref., 22 Jul. 1931, collector unknown (NSMT-I-L-75530) ; 1♀, same data as for preceding (NSMT-I-L-75531); 1♂, Mt Mitsumine, Saitama Pref., 15 Jul. 1961, T. Ebato leg. (NSMT-I-L-75461) ; 1♀, Shikanoyu, Titibu, Saitama Pref., 26 Jul. 1933, collector unknown (NSMT-I-L-75532) ; 1♂, Bushi, Iruma City, Saitama Pref., 10 Sep. 1979, H. Inoue leg. (NSMT-I-L-75635) ; 1♂, Kameyama, Chiba Pref., 12 Aug. 1963, T. Ebato leg. (NSMT-I-L-75517) ; 1♂, Kiyose, Tokyo To, 10 Jun. 1956, T. Ebato leg. (NSMT-I-L-75502) ; 2♀, ditto, 18 Aug. 1958, T. Ebato leg. (NSMT-I-L-75503, 75504); 1♀, ditto, 15 Aug. 1959, T. Ebato leg. (NSMT-I-L-75505); 1♀, ditto, 2 Jul. 1959, T. Ebato leg. (NSMT-I-L-75506); 1♂, ditto, 12 Jun. 1958, T. Ebato leg. (NSMT-I-L-75507); 1♀, ditto, 31 Aug. 1958, T. Ebato leg. (NSMT-I-L-75508); 1♀, ditto, 18 Aug. 1959, T. Ebato leg. (NSMT-I-L-75509); 1♀, ditto, 2 Jul. 1957, T. Ebato leg. (NSMT-I-L-75511); 1♂, ditto, 4 Jun. 1958, T. Ebato leg. (NSMT-I-L-75512); 1♀, Ohizumi, Tokyo To, 18 Aug. 1966, T. Ebato leg. (NSMT-I-L-75513) ; 2♂, Mt Takao, Tokyo To, 27 Jun. 1959, T. Ebato leg. (NSMT-I-L-75514, 75515) ; 1♂, ditto, 7 Jun. 1961, T. Maenami leg. (NSMT-I-L-75527); 1♂, ditto, 19 Jun. 1996, U. Jinbo leg. (NSMT-I-L-33621); 1♂, Mt Mihara, Tokyo To, 31 May 1962, R. Aoki leg. (NSMT-I-L-75521) ; 1♀, Mt Mitake, Tokyo To, 16 Jul. 1960, T. Maenami leg. (NSMT-I-L-75526) ; 1♂, ditto, 20 Aug. 1959, T. Ebato leg. (NSMT-I-L-75510); 1♀, ditto, 25 Jul. 1998, U. Jinbo leg. (NSMT-I-L-36058); 1♂, Institute of Nature Study, Minato-ku, Tokyo To, 6 Jun. 2017, U. Jinbo leg. (NSMT-I-L-55639) ; 1♂, Hodokubo, Hino City, Tokyo To, 3 Jun. 1990, U. Jinbo leg. (NSMT-I-L-75626) ; 1♀, ditto, 16 Jun. 1990, U. Jinbo leg. (NSMT-I-L-75627); 1♂, ditto, 3 Aug. 1991, U. Jinbo leg. (NSMT-I-L-75628); 1♀, ditto, 7 Sep. 1991, U. Jinbo leg. (NSMT-I-L-75629); 1♀, ditto, 23 Aug. 1992, U. Jinbo leg. (NSMT-I-L-75630); 1♂, Yokozawairi, Itsukaichi Town, Tokyo To, 18 Jun. 1994, U. Jinbo leg. (NSMT-I-L-75631) ; 1♂, ditto, Tokyo To, 27 Aug. 1994, U. Jinbo leg. (NSMT-I-L-75632) ; 1♀, Imperial Palace, Chiyoda-ku, Tokyo To, 26 May 2009, Y. Arita et al. leg. (NSMT-I-L-22523) ; 1♂, ditto, 7 Sep. 2010, [Malaise trap] (NSMT-I-L-22525); 1♂, ditto, 9 Aug. 2011, [Malaise trap] (NSMT-I-L-28051); 1♀, ditto, 4 Sep. 2012, Y. Arita, H. Nakajima, M. Owada, Y. Kishida and U. Jinbo leg. (NSMT-I-L-31522); 1♂, Noborito, Tokyo To, 27 May 1932, collector unknown (NSMT-I-L-75533) ; 1♂, Nishitanzawa, Kanagawa Pref., 18 Jun. 1966, Y. Kishida leg. (NSMT-I-L-75525) ; 1♀, Mt Myôjô-san, Itoigawa City, Niigata Pref., 7 Aug. 1999, A. Sasaki leg. (NSMT-I-L-75570) ; 1♂, Teradomari, Niigata Pref., 9 Aug. 2005, R. Sato leg. (NSMT-I-L-75571) ; 1♂, ditto, 7 Jun. 2005, T. Naito leg. (NSMT-I-L-75572); 1♂, Yuzurihara, Yamanashi Pref., 23 Jun. 1945, T. Ebato leg. (NSMT-I-L-75499) ; 1♂, Kiyosato, Yamanashi Pref., 22 Jul. 1967, T. Ebato leg. (NSMT-I-L-75500) ; 1♂, Karuizawa, Nagano Pref., 21 Jul. 1958, T. Ebato leg. (NSMT-I-L-75493) ; 1♂, Miyota, Nagano Pref., 17 Aug. 1965, T. Ebato leg. (NSMT-I-L-75494) ; 1♂, Tobira-Spa, Nagano Pref., 14 Jul. 1957, T. Ebato leg. (NSMT-I-L-75495) ; 1♀, Kumanotaira, Nagano Pref., 7 Jul. 1962, T. Ebato leg. (NSMT-I-L-75496) ; 1♂, ditto, 25 Jun. 1944, H. Inoue leg. (NSMT-I-L-75520); 1♂, Nashimoto, Shizuoka Pref., 10 Jun. 1961, T. Ebato leg. (NSMT-I-L-75516) ; 2♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.59&amp;materialsCitation.latitude=35.42" title="Search Plazi for locations around (long 138.59/lat 35.42)">Asagiri Plateau</a>, Shizuoka Pref., 35.42°N, 138.59°E, 910 m, 27 Aug. 2019, Y. Matsui leg. ; 1♂, Gujo-Rokunori, Gifu Pref., 5 Jul. 1966, S. Sawatani leg. (NSMT-I-L-75573) ; 1♂, Mt Hyôno-sen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.49&amp;materialsCitation.latitude=35.35" title="Search Plazi for locations around (long 134.49/lat 35.35)">Wakasa Town</a>, Tottori Pref., 35.35°N, 134.49°E, 860 m, 6 Nov. 2017 (F1 emerged), Y. Matsui leg. ; 1♂, ditto, 21 Jun. 2020, Y. Matsui leg.; 1♂, Hirodomeno, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.45&amp;materialsCitation.latitude=35.41" title="Search Plazi for locations around (long 134.45/lat 35.41)">Wakasa Town</a>, Tottori Pref., 35.41°N, 134.45°E, 800 m, 4 Sep. 2015, Y. Matsui leg. ; 4♀ 6♂, Tokumaru, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.34&amp;materialsCitation.latitude=35.37" title="Search Plazi for locations around (long 134.34/lat 35.37)">Yazu Town</a>, Tottori Pref., 35.37°N, 134.34°E, 145 m, 9-11 Aug. 2013, 10-20 Oct. 2014 (F1 emerged), H. Naka leg. ; 2♀, Wakabadai-kita, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.26&amp;materialsCitation.latitude=35.45" title="Search Plazi for locations around (long 134.26/lat 35.45)">City</a>, Tottori Pref., 35.45°N, 134.26°E, 40 m, 6 Jun. 2012, 22 Aug. 2014, Y. Matsui leg. ; 1♂, Kôchi, Shikano Town, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.0&amp;materialsCitation.latitude=35.4" title="Search Plazi for locations around (long 134.0/lat 35.4)">City</a>, Tottori Pref., 35.40°N, 134.00°E, 495 m, 12 Jun. 2020, Y. Matsui leg. ; 3♂, Mt Daisen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.55&amp;materialsCitation.latitude=35.35" title="Search Plazi for locations around (long 133.55/lat 35.35)">Kofu Town</a>, Tottori Pref., 35.35°N, 133.55°E, 910 m, 30 Sep.-2 Oct. 2017 (F1 emerged), Y. Matsui leg. ; 2♂, Mt Senjô-san, Kotoura Town, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.6&amp;materialsCitation.latitude=35.43" title="Search Plazi for locations around (long 133.6/lat 35.43)">City</a>, 35.43°N, 133.60°E, 385 m, 15 Jul. 2019, Y. Matsui leg. ; 1♂, Ichibata, Izumo City, Shimane Pref., 27 May 1967, T. Maenami leg. (NSMT-I-L-75522) ; 1♀, same data as for preceding (NSMT-I-L-75524); 1♀ 1♂, Kusandao, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.83&amp;materialsCitation.latitude=35.06" title="Search Plazi for locations around (long 132.83/lat 35.06)">Inan Town</a>, Shimane Pref., 35.06°N, 132.83°E, 915 m, Oct. 2017 (F1 emerged), Y. Matsui leg. ; 1♀, Sugano, Tokuyama City, Yamaguchi Pref., 3 Jun. 1994, T. Ikenoue leg. (NSMT-I-L-75577) ; 1♀, Yunoki, Tokuji Town, Yamaguchi Pref., 15 Jul. 1995, T. Ikenoue leg. (NSMT-I-L-75578) ; 2♀, Mt Tokusagamine, Yamaguchi Pref., 3 Aug. 1996, T. Ikenoue leg. (NSMT-I-L-75579, 75580) ; 1♂, Jakuchikyô, Yamaguchi Pref., 27 Jul. 1995, T. Ikenoue leg. (NSMT-I-L-75581) ; 1♂, Akiyoshi-dai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.31&amp;materialsCitation.latitude=34.24" title="Search Plazi for locations around (long 131.31/lat 34.24)">Mine City</a>, Yamaguchi, 34.24°N, 131.31°E, 240 m, 16 Sep. 2018, Y. Matsui leg. ; 1♀, Kamitsuru, Izumi City, Kagoshima Pref., 14 Jul. 1992, Y. Yanagita leg. (NSMT-I-L-75582) ; 1♀, Shin-Wase-Tunnel, I. Amamiohshima, Kagoshima Pref., 27 Mar. 2009, M. Owada and M. Kimura leg. (NSMT-I-L-75617); 1♂, Yona, I. Okinawajima, Okinawa Pref., 1 Apr. 1964, T. Nagano leg. (NSMT-I-L-75618) ; 1♂, Seifuautaki, Chinen-son, I. Okinawajima, Okinawa Pref., 8 Aug. 1980, R. Sato leg. (NSMT-I-L-75619) ; 2♂, Mt Terukubi-yama, Kunigami-son, I. Okinawajima, Okinawa Pref., 10 Aug. 1980, R. Sato leg. (NSMT-I-L-75620, 75621) ; 1♂, Haneji, Nago City, I. Okinawajima, Okinawa Pref., 17 Aug. 2001, M. Kimura leg. (NSMT-I-L-75622) ; 1♂, ditto, 9 Aug. 2002, M. Kimura leg. (NSMT-I-L-75623); 1♂, ditto, 12 Aug. 2002, M. Kimura leg. (NSMT-I-L-75624); 1♂, Takeda-Rindo, I. Ishigakijima, Okinawa Pref., 5 Jun. 2007, M. Kimura leg. (NSMT-I-L-75625).</p><p>Diagnosis.</p><p>Adult (Fig. 2C, D). Forewing length 9.0- 12.5 mm. This species is similar to N. tristalis and N. quadrimaculalis, but can be distinguished by the following characters: forewing shorter; vertex scales lighter than in N. tristalis and N. quadrimaculalis; subdiscal white spot of forewing usually quadrilateral and distinct; base of discal cell of hindwing broadly white; gnathos of male genitalia nearly triangular, short and small, apex rounded (Fig. 4B); valva of male genitalia dorsally straight margined subapically (Fig. 4C); anterior apophysis of female genitalia slightly curved to dorsally, expansion of the base is sharply triangular (Fig. 7B); signum of female genitalia circular, smaller than in N. tristalis (diameter 0.05-0.06 mm) (Fig. 7C). This species is also similar to N. occultalis, but N. occultalis has the following differences: forewing length 15-16 mm; subdiscal white spot of forewing narrowed or elongated; base of discal cell of hindwing concolorous with ground color; tuba analis of male genitalia sclerotized; gnathos of male genitalia elongated and narrow at the base. Lu and Du (2020) also mentioned genital differences between these species, and as far as we can see from the specimen image by Lu and Du (2020), N. bispina is externally distinguishable from this species by base of discal cell of the hindwing being concolorous with the ground color.</p><p>Distribution.</p><p>Japan, mainland China, Taiwan, Korea, Russia (southeast), India.</p><p>Host plant.</p><p>Rubus buergeri Miq., R. trifidus Thunb. (laboratory reared).</p><p>Remarks.</p><p>Our identification of this species is based on characters of external morphology (Motschulsky 1861; Inoue 1982; Li et al. 2012; Sasaki and Yamanaka 2013; Lu and Du 2020) and male genitalia (Li et al. 2012; Lu and Du 2020). The species status was confirmed by DNA barcodes.</p></div>	https://treatment.plazi.org/id/F362FB0FF6C45C70B37002DBB5C72348	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsui, Yuki;Naka, Hideshi;Jinbo, Utsugi	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
F6509A54AAB65E7DA9AF673568C41C33.text	F6509A54AAB65E7DA9AF673568C41C33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nagiella quadrimaculalis (Kollar & Redtenbacher 1844)	<div><p>Nagiella quadrimaculalis (Kollar &amp; Redtenbacher, 1844) Figures 2E, F, 5A-C, 8A-C</p><p>Scopula quadrimaculalis Kollar and Redtenbacher 1844: IV: 492.</p><p>Coptobasis quadrimaculalis: Lederer, 1863: 429-430; pl. 16 fig. 12.</p><p>Nagia desmialis Walker, 1866: 1320.</p><p>Sylepta quadrimaculalis: Shibuya, 1928: 229; pl. 8 fig. 14; Shibuya 1929: 189.</p><p>Nagia quadrimaculalis: Mutuura, 1957: 122, pl. 21 fig. 635.</p><p>Pleuroptya quadrimaculalis: Inoue, 1982: 1: 343; 2: 234, 454, pl. 40 fig. 17; Li et al. 2012: 624-625, pl. 18 fig. 415; Yamanaka 1995: 187, pl. 125 fig. 21; Sasaki and Yamanaka 2013: 81, 451.</p><p>Nagiella quadrimaculalis: Munroe, 1976: 878; Ullah et al. 2017: 70-72, figs. 2B, 4B, table 3; Lu and Du 2020: 149, fig. 6, figs. 11, 14.</p><p>Material examined.</p><p>Japan: 1♂, Marumori, Onikôbe, Narugo, Miyagi Pref., 30 Jul. 1997, M. Tanaka leg. (NSMT-I-L-75588) ; 1♂, Kirei-pass, Sumison, Miyazaki Pref., 9 Jul. 1992, Y. Yanagita leg. (NSMT-I-L-75592) ; 1♀, Hiromorigawa, Akita Pref., 14 Aug. 1988, A. Sasaki leg. (NSMT-I-L-75583) ; 1♀, Garo-Kyo, Fujisato Town, Akita Pref., 28 Jul. 2002, A. Sasaki leg. (NSMT-I-L-75584) ; 1♀, Yoroibata-Dam, Tazawako Town, Akita Pref., 26 Aug. 1989, A. Sasaki leg. (NSMT-I-L-75585) ; 2♂, Tose, Tamagawa, Tazawako Town, Akita Pref., 21 Aug. 1993, A. Sasaki leg. (NSMT-I-L-75586, 75587) ; 1♂, Futamata-Spa, Fukushima Pref., 6 Aug. 1967, T. Ebato leg. (NSMT-I-L-75462) ; 1♀, Houshi, Gunma Pref., 19 Jul. 1957, T. Ebato leg. (NSMT-I-L-75466); (NSMT-I-L-75472) , 1♂, Kawaburu-Spa, Gunma Pref., 1 Jul. 1967, T. Ebato leg. ; 1♂, Mt Takao, Tokyo To, 23 Jun. 1951, T. Haruta leg. (NSMT-I-L-75479) ; 1♀, ditto, 10 Jul. 1960, T. Ebato leg. (NSMT-I-L-75463); 1♂, ditto, 26 Jun. 1959, T. Ebato leg. (NSMT-I-L-75464); 1♀, ditto, 27 Jun. 1959, T. Ebato leg. (NSMT-I-L-75474); 1♂, Nippara, Tokyo To, 8 Aug. 1961, T. Ebato leg. (NSMT-I-L-75465) ; 1♀, same data as for preceding (NSMT-I-L-75477); 2♂, ditto, 2 Sep. 1961, T. Ebato leg. (NSMT-I-L-75475, 75476); 1♀, Mt Mitake, Tokyo To, 27 Aug. 1960, T. Maenami leg. (NSMT-I-L-75486) ; 1♂, ditto, 20 Jun. 1996, U. Jinbo leg. (NSMT-I-L-36059); 1♀, Ohnita, Ohme City, Tokyo To, 18 Aug. 1996, U. Jinbo leg. (NSMT-I-L-75634) ; 1♂, Yuzurihara, Yamanashi Pref., 1 Sep. 1945, T. Ebato leg. (NSMT-I-L-75471) ; 1♂, ditto, 2 Sep. 1945, T. Ebato leg. (NSMT-I-L-75469); 1♂, ditto, 5 Sep. 1945, T. Ebato leg. (NSMT-I-L-75470); 1♂, ditto, 23 Sep. 1954, T. Ebato leg. (NSMT-I-L-75468); 1♂, Sagashio-Spa, Yamanashi Pref., 9 Aug. 1969, T. Ebato leg. (NSMT-I-L-75473) ; 2♂, Ashiyasu, Yamanashi Pref., 16 Jul. 1977, T. Ebato leg. (NSMT-I-L-75480, 75482) ; 1♂, ditto, 6 Aug. 1977, T. Ebato leg. (NSMT-I-L-75483); 1♀, ditto, 19 Jul. 1980, T. Ebato leg. (NSMT-I-L-75478); 1♂, Nishiyama-Spa, Yamanashi Pref., 17 Aug. 1981, T. Ebato leg. (NSMT-I-L-75481) ; 1♀, Hirayu, Gifu Pref., 7 Aug. 1953, T. Haruta leg. (NSMT-I-L-75467) ; 1♂, Gujo-Rokunori, Gifu Pref., 1 Jul. 1966, S. Sawatani leg. (NSMT-I-L-75589) ; 1♂, Osugi-dani, Wakayama Pref., 4 Aug. 1976, S. Nakatani leg. (NSMT-I-L-75484) ; 1♂, Shimakawa-Osugi-dani, Wakayama Pref., 5 Jul. 1975, S. Nakatani leg. (NSMT-I-L-75485); 1♂, Tatsumi-tôge, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.01&amp;materialsCitation.latitude=35.32" title="Search Plazi for locations around (long 134.01/lat 35.32)">City</a>, Tottori Pref., 35.32°N, 134.01°E, 670 m, 2 Jul. 2019, Y. Matsui leg. ; 2♀ 3♂, Mt Daisen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.55&amp;materialsCitation.latitude=35.35" title="Search Plazi for locations around (long 133.55/lat 35.35)">Kofu Town</a>, Tottori Pref., 35.35°N, 133.55°E, 910 m, 30 Sep.-29 Oct. 2017, and 18 Apr. 2018 (F1 emerged), Y. Matsui leg. ; 1♂, Suemochi, Shikano Town, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.09&amp;materialsCitation.latitude=35.45" title="Search Plazi for locations around (long 134.09/lat 35.45)">City</a>, Tottori Pref., 35.45°N, 134.09°E, 220 m, 1 Jun. 2020, Y. Matsui leg. ; 1♂, Kôchi, Shikano Town, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.0&amp;materialsCitation.latitude=35.4" title="Search Plazi for locations around (long 134.0/lat 35.4)">City</a>, Tottori Pref., 35.40°N, 134.00°E, 495 m, 12 Jun. 2020, Y. Matsui leg. ; 2♀ 1♂, ditto, 12-16 Aug. 2020 (F1 emerged), Y. Matsui leg.; 1♂, Sourokubara, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.11&amp;materialsCitation.latitude=35.46" title="Search Plazi for locations around (long 134.11/lat 35.46)">City</a>, Tottori Pref., 35.46°N, 134.11°E, 110 m, 19 Jul. 2020 (larvae: collected from Rubus buergeri), 20 Aug. 2020 (emerged), Y. Matsui leg. ; 1♀ 1♂, Toyofusa, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.55&amp;materialsCitation.latitude=35.41" title="Search Plazi for locations around (long 133.55/lat 35.41)">Daisen Town</a>, Tottori Pref., 35.41°N, 133.55°E, 680 m, 2 Sep. 2020 (larvae: collected from R. palmatus), 19-20 Oct. 2020 (emerged) Y. Matsui leg. ; 1♂, Yakawa, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.13&amp;materialsCitation.latitude=35.1" title="Search Plazi for locations around (long 133.13/lat 35.1)">Okuizumo Town</a>, Shimane Pref., 35.10°N, 133.13°E, 680 m, 6 Sep. 2013, Y. Matsui leg. ; 2♂, Omogokei, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.1&amp;materialsCitation.latitude=33.72" title="Search Plazi for locations around (long 133.1/lat 33.72)">Kumakogen Town</a>, Ehime Pref., 33.72°N, 133.10°E, 700 m, 8 Jun. 2019, Y. Matsui leg. ; 1♂, Shimomyo, Aira-Cho, Kagoshima Pref., 28 May 1992, Y. Yanagita leg. (NSMT-I-L-75590) ; 1♂, Tobi, Miyanojo-cho, Kagoshima Pref., 26 May 1992, Y. Yanagita leg. (NSMT-I-L-75591) ; 1♀, Mt Ishizukadake, I. Yakushima, Kagoshima Pref., 5-6 Aug. 1958, B. T. leg. (NSMT-I-L-75599) ; 1♂, same data as for preceding (NSMT-I-L-75600); 4♂, ditto, 17 Jul. 1970, K. Tobi leg. (NSMT-I-L-75601 to 75604); 1♀, same data as for preceding (NSMT-I-L-75605) .</p><p>Diagnosis.</p><p>Adult (Fig. 2E, F). Forewing length 15.5-18 mm. This species is similar to N. tristalis and N. inferior, but can be distinguished by the following characters: forewing longer; vertex scales brown, darker than in N. inferior; cilia cream white at Cu2 to A1+2 for forewing, Cu2 to CuP for hindwing; subdiscal white spot of forewing quadrilateral or rounded, rather distinct; base of discal cell of hindwing partially white; gnathos of male genitalia slender and elongated (Fig. 5B); valva of male genitalia broader than in N. tristalis and N. inferior, with straight dorsal margin subapically (Fig. 5C); anterior apophysis of female genitalia slightly curved, expanded near bases, but not triangular (Fig. 8B); signum of female corpus brusae larger than that of N. tristalis and N. inferior (diameter 0.16-0.19 mm), with sharp projections at both edges of posterior margin (Fig. 8C). Ullah et al. (2017) provided diagnostic characters to distinguish this species from N. occultalis . Lu and Du (2020) mentioned genital differences between these species, as far as we can see, the specimen image of N. bispina by Lu and Du (2020) is externally distinguishable from this species by the cilia of each wing being concolorous with the ground color.</p><p>Distribution.</p><p>Japan, mainland China, Taiwan, Korea, Russia (southeast), Southeast Asia, Nepal, India.</p><p>Host plant.</p><p>Rubus buergeri Miq., R. palmatus Thunb. (in the field), R. buergeri, R. trifidus Thunb. (laboratory reared).</p><p>Remarks.</p><p>Nagia incomitata Swinhoe, 1894 has long been considered a synonym of N. quadrimaculalis, but based on the investigation of the type specimen, Lu and Du (2020) considered it likely to belong to Nosophora Lederer, 1863. We also follow this taxonomic treatment.</p><p>Our identification of this species in this study was based on external morphology (Kollar and Redtenbacher 1844; Inoue 1982; Li et al. 2012; Sasaki and Yamanaka 2013; Ullah et al. 2017; Lu and Du 2020) and male genitalia (Li et al. 2012; Ullah et al. 2017; Lu and Du 2020). The species status was confirmed by DNA barcodes.</p></div>	https://treatment.plazi.org/id/F6509A54AAB65E7DA9AF673568C41C33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsui, Yuki;Naka, Hideshi;Jinbo, Utsugi	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
C9D9CE9CFFB3503B98FD90FDE60D40F4.text	C9D9CE9CFFB3503B98FD90FDE60D40F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nagiella tristalis Matsui & Naka 2021	<div><p>Nagiella tristalis Matsui &amp; Naka sp. nov. Figures 2A, B, 3A-C, 6A-C</p><p>Type material.</p><p>Holotype. ♂, Japan: Sourokubara, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.11&amp;materialsCitation.latitude=35.46" title="Search Plazi for locations around (long 134.11/lat 35.46)">City</a>, Tottori Pref., 35.46°N, 134.11°E, 110 m, 7 Nov. 2019 (F1 emerged), Y. Matsui leg. , preserved in National Museum of Nature and Science, NSMT-I-L-75637. Paratypes. 2♀ 3♂, Same locality as holotype, 5 Mar. 2018, 6 Apr. 2018, 4 May 2018, 13 Sep. 2018 (emerged), H. Naka, and Y. Matsui leg. ; 1♀ 3♂, Setagura, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.12&amp;materialsCitation.latitude=35.47" title="Search Plazi for locations around (long 134.12/lat 35.47)">City</a>, Tottori Pref., 35.47°N, 134.12°E, 45 m, 7-22 Mar. 2019 (emerged), 23 Sep. 2019 (F1 emerged), H. Naka leg. ; 2♀, Uemachi, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.24&amp;materialsCitation.latitude=35.5" title="Search Plazi for locations around (long 134.24/lat 35.5)">City</a>, Tottori Pref., 35.50°N, 134.24°E, 40 m, 5 and 10 Feb. 2019 (emerged), Y. Matsui leg. ; 1♀, Mt Honjin-yama, Tottori <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.26&amp;materialsCitation.latitude=35.51" title="Search Plazi for locations around (long 134.26/lat 35.51)">City</a>, Tottori Pref., 35.51°N, 134.26°E, 110 m, 24 Jun. 2012, Y. Matsui leg. ; 1♂, Tokumaru, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.34&amp;materialsCitation.latitude=35.37" title="Search Plazi for locations around (long 134.34/lat 35.37)">Yazu Town</a>, Tottori Pref., 35.37°N, 134.34°E, 145 m, 20 Aug. 2014, H. Naka leg. Other specimens. 1♀, Mt Takao, Tokyo To, 19 Jul. 1960, T. Ebato leg. (NSMT-I-L-75536) ; 1♂, Nashimoto, Shizuoka Pref., 23 May 1953, T. Ebato leg. (NSMT-I-L-75538) ; 1♂, ditto, 5 Jun. 1959, T. Ebato leg. (NSMT-I-L-75537); 1♂, ditto, 10 Jun. 1961, T. Ebato leg. (NSMT-I-L-75539); 2♂, ditto, 24 Aug. 1966, T. Ebato leg. (NSMT-I-L-75534, 75535); 1♀, Kuragari-Valley, Nukata Town, Aichi Pref., 26 Jun. 1993, A. Sasaki leg. (NSMT-I-L-75593) ; 1♂, Sugano, Tokuyama City, Yamaguchi Pref., 27 Jun. 1993, T. Ikenoue leg. (NSMT-I-L-75594) ; 1♀, Shimomyo, Aira Town, Kagoshima Pref., 28 May 1992, Y. Yanagita leg. (NSMT-I-L-75596) ; 1♀, Kamitsuru, Izumi City, Kagoshima Pref., 14 Jul. 1992, Y. Yanagita leg. (NSMT-I-L-75595) ; 1♂, Mt Ishizukadake, I. Yakushima, Kagoshima Pref., 5 Aug. 1958, B. T. leg. (NSMT-I-L-75607) ; 1♂, Nagata, I. Yakushima, Kagoshima Pref., 3 Oct. 2006, M. Owada and T. Fukuda leg. (NSMT-I-L-75606) ; 1♀, Chuo-rindo, Uken, I. Amamiohshima, Kagoshima Pref., 13 Oct. 1988, M. Owada leg. (NSMT-I-L-75541) ; 3♂, ditto, 22 Apr. 2009, M. Owada leg. (NSMT-I-L-75609 to 75611); 1♀, Kinsakubaru, Naze, I. Amamiohshima, Kagoshima Pref., 11 Oct. 1988, M. Owada leg. (NSMT-I-L-75542) ; 1♂, Mt Yuwan-dake, I. Amamiohshima, Kagoshima Pref., 12 Oct. 1988, M. Owada leg. (NSMT-I-L-75543) ; 1♀, Naze, I. Amamiohshima, Kagoshima Pref., 25 Jun. 1968, Y. Kishida leg. (NSMT-I-L-75540) ; 1♂, Shinokawa, Setouchi, I. Amamiohshima, Kagoshima Pref., 21 Apr. 2009, M. Owada leg. (NSMT-I-L-75608) ; 1♂, Mikyo, I. Tokunoshima, Kagoshima Pref., 31 Oct. 1992, M. Owada leg. (NSMT-I-L-75544) ; 1♂, Gogayama, I. Okinawajima, Okinawa Pref., 30 Mar. 1974, T. Naito leg. (NSMT-I-L-75612) ; 1♀, Seifuautaki, Chinen-son, I. Okinawajima, Okinawa Pref., 16 Aug. 1980, R. Sato leg. (NSMT-I-L-75613) ; 1♂, same data as for preceding (NSMT-I-L-75614); 1♀, Ôkuni-bashi, Kunigami-son, I. Okinawajima, Okinawa Pref., 21 Apr. 2001, A. Sasaki leg. (NSMT-I-L-75615).</p><p>Etymology.</p><p>The specific epithet refers to the darker wing color in comparison to that of N. inferior, and the habitat of this species is a shaded place.</p><p>Diagnosis.</p><p>This new species is similar to N. inferior and N. quadrimaculalis, also distributed in Japan, but it can be distinguished by the following characters: length of forewing 12.0-13.0 mm; vertex with erect, dull-orange scales; subdiscal white spot of forewing rounded, small, and blurry; base of discal cell of hindwing identical to ground color; dorsal margin of valva of male genitalia slightly incurved subapically; anterior apophysis of female genitalia straight and narrow; signum of female genitalia nearly elliptical, larger than in N. inferior (diameter 0.09-0.14 mm). This species is also similar to N. occultalis and N. bispina distributed in China, but N. occultalis has the following differences: subdiscal white spot of forewing narrowed or elongated, tuba analis of male genitalia sclerotized, gnathos of male genitalia elongated and narrow at the base; N. bispina exhibits the following differences: gnathos of male genitalia absent, phallus of male genitalia with a hook-shaped cornutus, corpus bursae of female genitalia with two thorn-like signa. From N. hortulatoides, the new species can be easily distinguished by the wing maculation.</p><p>Description</p><p>(Fig. 2A, B). Head: frons brownish grey, smooth. Vertex with erect, dull-orange scales. Labial palpus upturned, dorso-laterally dark brown, ventro-mesally pale white. Antenna dark brown; flagellum filiform with golden cilia ca 1/4 the diameter of flagellum in male.</p><p>Thorax and abdomen: dorsally brownish grey; patagium and tegula with ochreous brown. Ventrally milky white.</p><p>Wings: length of forewing 12.0-13.0 mm. Ground color of both wings brownish grey, with a large comma-shaped white spot at end of discal cell (the bases of R 5 to M3), that of the hindwing somewhat small; cilia concolorous with ground color; postmedial line obscure. Subdiscal white spot of forewing rounded, small and blurry. Base of discal cell of hindwing concolorous with ground color.</p><p>Male genitalia (Fig. 3A-C): uncus short, subtrapezoid, blunt on posterior margin. Gnathos nearly triangular, short and small, apex blunt. Tuba analis elongate, not sclerotized, length ca 0.6 times that of valva. Transtilla subtriangular, completely connected medially. Saccus short, anterior margin rounded. Valva somewhat narrow, length ca 3.3 times that of width, dorsal margin slightly incurved subapically; costa more or less inflated with several setae at apex; clasper large, down curved, the apex blunt. Phallus cylindrical, without cornutus.</p><p>Female genitalia (Fig. 6A-C): ovipositor lobe oblong, setose. Anterior apophysis ca 2 times length of posterior apophysis, narrow, almost straight, with triangular expansion near base. Antrum trapezoid, sclerotized at collar. Ductus bursae almost equal in length to corpus bursae, membranous with wrinkles, slightly narrowed at posterior end. Corpus bursae pear-shaped, membranous; signum nearly elliptical, sparsely covered with denticles.</p><p>Biology.</p><p>In Honshu, Japan, adults are found in May to September, and they are considered bivoltine. They appear to be hardly attracted to light. Larvae feed on Rubus buergeri and the middle instar larvae overwinter in its leaves.</p><p>Distribution.</p><p>Japan: Honshu (Tokyo, Shizuoka, Aichi, Tottori, Yamaguchi), Kyushu (Kagoshima), Ryukyu Islands (Yakushima, Amamioshima, Tokunoshima, Okinawajima) .</p><p>Remarks.</p><p>The shapes of the uncus and gnathos show intraspecific variations, i.e., in several specimens, the posterior margin of the uncus is slightly notched medially, and the projection of the gnathos is smaller than that shown on Figure 3B. Also, in N. inferior, the shape of the gnathos varies similarly as in N. tristalis . Therefore, these characters are unsuitable for diagnosis of N. tristalis and N. inferior . Similarly, Ullah et al. (2017) proposed the shape of the claspers as a diagnostic character for distinguishing between N. occultalis and N. quadrimaculalis, but we could not find any clear difference among N. tristalis, N. inferior, and N. quadrimaculalis, because of the overlap in intraspecific variations.</p></div>	https://treatment.plazi.org/id/C9D9CE9CFFB3503B98FD90FDE60D40F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsui, Yuki;Naka, Hideshi;Jinbo, Utsugi	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
