taxonID	type	description	language	source
F362FB0FF6C45C70B37002DBB5C72348.taxon	materials_examined	Material examined. Japan: 1 ♂, Tohro, Hokkaido, 3 Jul. 1962, T. Ebato leg., (NSMT-I-L- 75498); 1 ♂, Riv. Rusagawa, Shiretoko, Hokkaido, 26 Jul. 1962, K. Tsuchiya leg. (NSMT-I-L- 75523); 1 ♂, Sapporo, Hokkaido, 6 Jul. 1933, collector unknown (NSMT-I-L- 75529); 1 ♂, Shumarinai, Hokkaido, 20 Jul. 1998, Y. Kishida leg. (NSMT-I-L- 75569); 1 ♂, Shibecha, Hokkaido, 8 Jul. 1958, K. Jinbo leg. (NSMT-I-L- 75574); 2 ♂, Akan, Hokkaido, 13 Jul. 1958, K. Jinbo leg. (NSMT-I-L- 75575, 75576); 1 ♂, Toubai, Nemuro City, Hokkaido, 5 Aug. 2013, U. Jinbo leg. (NSMT-I-L- 37577); 1 ♂, Sannai-Ishizawa, Honjoh City, Akita Pref., 29 Jun. 1975, A. Sasaki leg. (NSMT-I-L- 75548); 1 ♂, ditto, 29 Jun. 1978, A. Sasaki leg. (NSMT-I-L- 75545); 2 ♂, ditto, 15 Jun. 1979, A. Sasaki leg. (NSMT-I-L- 75546, 75547); 1 ♀, Shinzan Park, Honjoh City, Akita Pref., 11 Jul. 1977, A. Sasaki leg. (NSMT-I-L- 75549); 1 ♂, Uwanodai, Kawabe Town, Akita Pref., 5 Jul. 1977, A. Sasaki leg. (NSMT-I-L- 75555); 1 ♀, ditto, 12 Jul. 1977, A. Sasaki leg. (NSMT-I-L- 75551); 1 ♀, ditto, 26 Jun. 1978, A. Sasaki leg. (NSMT-I-L- 75552); 1 ♀, ditto, 19 Jul. 1979, A. Sasaki leg.; 1 ♂, ditto, 19 Jul. 1979, A. Sasaki leg. (NSMT-I-L- 75556); 1 ♂, ditto, 11 Jun. 1980, A. Sasaki leg. (NSMT-I-L- 75550); 1 ♂, ditto, Akita Pref., 2 Jul. 1980, A. Sasaki leg. (NSMT-I-L- 75553); 1 ♀, Kamibiguchi, Gojohme Town, Akita Pref., 26 Jul. 1979, A. Sasaki leg. (NSMT-I-L- 75557); 1 ♂, Tazawako Height, Tazawako Town, Akita Pref., 20 Aug. 1979, A. Sasaki leg. (NSMT-I-L- 75558); 1 ♂, Chikogi-zaki, Hachimori Town, Akita Pref., 13 Jul. 1979, A. Sasaki leg. (NSMT-I-L- 75559); 1 ♀, Niida, Akita City, Akita Pref., 16 Jun. 1978, A. Sasaki leg. (NSMT-I-L- 75560); 1 ♂, Asahimata, Akita City, Akita Pref., 16 Jul. 1980, A. Sasaki leg. (NSMT-I-L- 75561); 1 ♂, Nibetsu, Akita City, Akita Pref., 3 Jul. 1980, A. Sasaki leg. (NSMT-I-L- 75562); 1 ♂, ditto, 23 Jun. 1987, A. Sasaki leg. (NSMT-I-L- 75563); 1 ♂, Mt Takao, Yuma Town, Akita Pref., 8 Jul. 1985, A. Sasaki leg. (NSMT-I-L- 75564); 1 ♂, Tōshi, Nikaho Town, Akita Pref., 18 Jul. 1995, A. Sasaki leg. (NSMT-I-L- 75565); 1 ♂, ditto, 19 Aug. 1984, A. Sasaki leg. (NSMT-I-L- 75566); 1 ♂, Matsusaka, Ōno-dai, Moriyoshi Town, Akita Pref., 13 Jul. 1996, A. Sasaki leg. (NSMT-I-L- 75567); 1 ♀, Aburato, Tsuruoka City, Yamagata Pref., 11 Jul. 1990, A. Sasaki leg. (NSMT-I-L- 75568); 1 ♂, Futamata-Spa, Fukushima Pref., 6 Aug. 1967, T. Ebato leg. (NSMT-I-L- 75497); 1 ♂, Shiozawa-Spa, Fukushima Pref., 5 Aug. 1967, T. Ebato leg. (NSMT-I-L- 75501); 1 ♂, ditto, 6 Aug. 1967, T. Ebato leg. (NSMT-I-L- 75519); 1 ♂, ditto, 29 Jun. 1968, T. Ebato leg. (NSMT-I-L- 75518); 2 ♂, Hanashiki-Spa, Gunma Pref., 22 Jun. 1963, T. Ebato leg. (NSMT-I-L- 75459, 75460); 1 ♀, same data as for preceding, (NSMT-I-L- 75487); 1 ♂, ditto, 8 Jul. 1961, T. Ebato leg. (NSMT-I-L- 75492); 3 ♂, Uenohara, Gunma Pref., 8 Jul. 1961, T. Ebato leg. (NSMT-I-L- 75488 to 75490); 1 ♂, Kawaburu-Spa, Gunma Pref., 1 Jul. 1967, T. Ebato leg. (NSMT-I-L- 75491); 1 ♂, Kitakaruizawa, Gunma Pref., 12 Jul. 1970, T. Okada leg. (NSMT-I-L- 75528); 1 ♂, Minakami, Gunma Pref., 22 Jul. 1931, collector unknown (NSMT-I-L- 75530); 1 ♀, same data as for preceding (NSMT-I-L- 75531); 1 ♂, Mt Mitsumine, Saitama Pref., 15 Jul. 1961, T. Ebato leg. (NSMT-I-L- 75461); 1 ♀, Shikanoyu, Titibu, Saitama Pref., 26 Jul. 1933, collector unknown (NSMT-I-L- 75532); 1 ♂, Bushi, Iruma City, Saitama Pref., 10 Sep. 1979, H. Inoue leg. (NSMT-I-L- 75635); 1 ♂, Kameyama, Chiba Pref., 12 Aug. 1963, T. Ebato leg. (NSMT-I-L- 75517); 1 ♂, Kiyose, Tokyo To, 10 Jun. 1956, T. Ebato leg. (NSMT-I-L- 75502); 2 ♀, ditto, 18 Aug. 1958, T. Ebato leg. (NSMT-I-L- 75503, 75504); 1 ♀, ditto, 15 Aug. 1959, T. Ebato leg. (NSMT-I-L- 75505); 1 ♀, ditto, 2 Jul. 1959, T. Ebato leg. (NSMT-I-L- 75506); 1 ♂, ditto, 12 Jun. 1958, T. Ebato leg. (NSMT-I-L- 75507); 1 ♀, ditto, 31 Aug. 1958, T. Ebato leg. (NSMT-I-L- 75508); 1 ♀, ditto, 18 Aug. 1959, T. Ebato leg. (NSMT-I-L- 75509); 1 ♀, ditto, 2 Jul. 1957, T. Ebato leg. (NSMT-I-L- 75511); 1 ♂, ditto, 4 Jun. 1958, T. Ebato leg. (NSMT-I-L- 75512); 1 ♀, Ohizumi, Tokyo To, 18 Aug. 1966, T. Ebato leg. (NSMT-I-L- 75513); 2 ♂, Mt Takao, Tokyo To, 27 Jun. 1959, T. Ebato leg. (NSMT-I-L- 75514, 75515); 1 ♂, ditto, 7 Jun. 1961, T. Maenami leg. (NSMT-I-L- 75527); 1 ♂, ditto, 19 Jun. 1996, U. Jinbo leg. (NSMT-I-L- 33621); 1 ♂, Mt Mihara, Tokyo To, 31 May 1962, R. Aoki leg. (NSMT-I-L- 75521); 1 ♀, Mt Mitake, Tokyo To, 16 Jul. 1960, T. Maenami leg. (NSMT-I-L- 75526); 1 ♂, ditto, 20 Aug. 1959, T. Ebato leg. (NSMT-I-L- 75510); 1 ♀, ditto, 25 Jul. 1998, U. Jinbo leg. (NSMT-I-L- 36058); 1 ♂, Institute of Nature Study, Minato-ku, Tokyo To, 6 Jun. 2017, U. Jinbo leg. (NSMT-I-L- 55639); 1 ♂, Hodokubo, Hino City, Tokyo To, 3 Jun. 1990, U. Jinbo leg. (NSMT-I-L- 75626); 1 ♀, ditto, 16 Jun. 1990, U. Jinbo leg. (NSMT-I-L- 75627); 1 ♂, ditto, 3 Aug. 1991, U. Jinbo leg. (NSMT-I-L- 75628); 1 ♀, ditto, 7 Sep. 1991, U. Jinbo leg. (NSMT-I-L- 75629); 1 ♀, ditto, 23 Aug. 1992, U. Jinbo leg. (NSMT-I-L- 75630); 1 ♂, Yokozawairi, Itsukaichi Town, Tokyo To, 18 Jun. 1994, U. Jinbo leg. (NSMT-I-L- 75631); 1 ♂, ditto, Tokyo To, 27 Aug. 1994, U. Jinbo leg. (NSMT-I-L- 75632); 1 ♀, Imperial Palace, Chiyoda-ku, Tokyo To, 26 May 2009, Y. Arita et al. leg. (NSMT-I-L- 22523); 1 ♂, ditto, 7 Sep. 2010, [Malaise trap] (NSMT-I-L- 22525); 1 ♂, ditto, 9 Aug. 2011, [Malaise trap] (NSMT-I-L- 28051); 1 ♀, ditto, 4 Sep. 2012, Y. Arita, H. Nakajima, M. Owada, Y. Kishida and U. Jinbo leg. (NSMT-I-L- 31522); 1 ♂, Noborito, Tokyo To, 27 May 1932, collector unknown (NSMT-I-L- 75533); 1 ♂, Nishitanzawa, Kanagawa Pref., 18 Jun. 1966, Y. Kishida leg. (NSMT-I-L- 75525); 1 ♀, Mt Myojo-san, Itoigawa City, Niigata Pref., 7 Aug. 1999, A. Sasaki leg. (NSMT-I-L- 75570); 1 ♂, Teradomari, Niigata Pref., 9 Aug. 2005, R. Sato leg. (NSMT-I-L- 75571); 1 ♂, ditto, 7 Jun. 2005, T. Naito leg. (NSMT-I-L- 75572); 1 ♂, Yuzurihara, Yamanashi Pref., 23 Jun. 1945, T. Ebato leg. (NSMT-I-L- 75499); 1 ♂, Kiyosato, Yamanashi Pref., 22 Jul. 1967, T. Ebato leg. (NSMT-I-L- 75500); 1 ♂, Karuizawa, Nagano Pref., 21 Jul. 1958, T. Ebato leg. (NSMT-I-L- 75493); 1 ♂, Miyota, Nagano Pref., 17 Aug. 1965, T. Ebato leg. (NSMT-I-L- 75494); 1 ♂, Tobira-Spa, Nagano Pref., 14 Jul. 1957, T. Ebato leg. (NSMT-I-L- 75495); 1 ♀, Kumanotaira, Nagano Pref., 7 Jul. 1962, T. Ebato leg. (NSMT-I-L- 75496); 1 ♂, ditto, 25 Jun. 1944, H. Inoue leg. (NSMT-I-L- 75520); 1 ♂, Nashimoto, Shizuoka Pref., 10 Jun. 1961, T. Ebato leg. (NSMT-I-L- 75516); 2 ♂, Asagiri Plateau, Shizuoka Pref., 35.42 ° N, 138.59 ° E, 910 m, 27 Aug. 2019, Y. Matsui leg.; 1 ♂, Gujo-Rokunori, Gifu Pref., 5 Jul. 1966, S. Sawatani leg. (NSMT-I-L- 75573); 1 ♂, Mt Hyono-sen, Wakasa Town, Tottori Pref., 35.35 ° N, 134.49 ° E, 860 m, 6 Nov. 2017 (F 1 emerged), Y. Matsui leg.; 1 ♂, ditto, 21 Jun. 2020, Y. Matsui leg.; 1 ♂, Hirodomeno, Wakasa Town, Tottori Pref., 35.41 ° N, 134.45 ° E, 800 m, 4 Sep. 2015, Y. Matsui leg.; 4 ♀ 6 ♂, Tokumaru, Yazu Town, Tottori Pref., 35.37 ° N, 134.34 ° E, 145 m, 9 - 11 Aug. 2013, 10 - 20 Oct. 2014 (F 1 emerged), H. Naka leg.; 2 ♀, Wakabadai-kita, Tottori City, Tottori Pref., 35.45 ° N, 134.26 ° E, 40 m, 6 Jun. 2012, 22 Aug. 2014, Y. Matsui leg.; 1 ♂, Kochi, Shikano Town, Tottori City, Tottori Pref., 35.40 ° N, 134.00 ° E, 495 m, 12 Jun. 2020, Y. Matsui leg.; 3 ♂, Mt Daisen, Kofu Town, Tottori Pref., 35.35 ° N, 133.55 ° E, 910 m, 30 Sep. - 2 Oct. 2017 (F 1 emerged), Y. Matsui leg.; 2 ♂, Mt Senjo-san, Kotoura Town, Tottori City, 35.43 ° N, 133.60 ° E, 385 m, 15 Jul. 2019, Y. Matsui leg.; 1 ♂, Ichibata, Izumo City, Shimane Pref., 27 May 1967, T. Maenami leg. (NSMT-I-L- 75522); 1 ♀, same data as for preceding (NSMT-I-L- 75524); 1 ♀ 1 ♂, Kusandao, Inan Town, Shimane Pref., 35.06 ° N, 132.83 ° E, 915 m, Oct. 2017 (F 1 emerged), Y. Matsui leg.; 1 ♀, Sugano, Tokuyama City, Yamaguchi Pref., 3 Jun. 1994, T. Ikenoue leg. (NSMT-I-L- 75577); 1 ♀, Yunoki, Tokuji Town, Yamaguchi Pref., 15 Jul. 1995, T. Ikenoue leg. (NSMT-I-L- 75578); 2 ♀, Mt Tokusagamine, Yamaguchi Pref., 3 Aug. 1996, T. Ikenoue leg. (NSMT-I-L- 75579, 75580); 1 ♂, Jakuchikyo, Yamaguchi Pref., 27 Jul. 1995, T. Ikenoue leg. (NSMT-I-L- 75581); 1 ♂, Akiyoshi-dai, Mine City, Yamaguchi, 34.24 ° N, 131.31 ° E, 240 m, 16 Sep. 2018, Y. Matsui leg.; 1 ♀, Kamitsuru, Izumi City, Kagoshima Pref., 14 Jul. 1992, Y. Yanagita leg. (NSMT-I-L- 75582); 1 ♀, Shin-Wase-Tunnel, I. Amamiohshima, Kagoshima Pref., 27 Mar. 2009, M. Owada and M. Kimura leg. (NSMT-I-L- 75617); 1 ♂, Yona, I. Okinawajima, Okinawa Pref., 1 Apr. 1964, T. Nagano leg. (NSMT-I-L- 75618); 1 ♂, Seifuautaki, Chinen-son, I. Okinawajima, Okinawa Pref., 8 Aug. 1980, R. Sato leg. (NSMT-I-L- 75619); 2 ♂, Mt Terukubi-yama, Kunigami-son, I. Okinawajima, Okinawa Pref., 10 Aug. 1980, R. Sato leg. (NSMT-I-L- 75620, 75621); 1 ♂, Haneji, Nago City, I. Okinawajima, Okinawa Pref., 17 Aug. 2001, M. Kimura leg. (NSMT-I-L- 75622); 1 ♂, ditto, 9 Aug. 2002, M. Kimura leg. (NSMT-I-L- 75623); 1 ♂, ditto, 12 Aug. 2002, M. Kimura leg. (NSMT-I-L- 75624); 1 ♂, Takeda-Rindo, I. Ishigakijima, Okinawa Pref., 5 Jun. 2007, M. Kimura leg. (NSMT-I-L- 75625).	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
F362FB0FF6C45C70B37002DBB5C72348.taxon	diagnosis	Diagnosis. Adult (Fig. 2 C, D). Forewing length 9.0 - 12.5 mm. This species is similar to N. tristalis and N. quadrimaculalis, but can be distinguished by the following characters: forewing shorter; vertex scales lighter than in N. tristalis and N. quadrimaculalis; subdiscal white spot of forewing usually quadrilateral and distinct; base of discal cell of hindwing broadly white; gnathos of male genitalia nearly triangular, short and small, apex rounded (Fig. 4 B); valva of male genitalia dorsally straight margined subapically (Fig. 4 C); anterior apophysis of female genitalia slightly curved to dorsally, expansion of the base is sharply triangular (Fig. 7 B); signum of female genitalia circular, smaller than in N. tristalis (diameter 0.05 - 0.06 mm) (Fig. 7 C). This species is also similar to N. occultalis, but N. occultalis has the following differences: forewing length 15 - 16 mm; subdiscal white spot of forewing narrowed or elongated; base of discal cell of hindwing concolorous with ground color; tuba analis of male genitalia sclerotized; gnathos of male genitalia elongated and narrow at the base. Lu and Du (2020) also mentioned genital differences between these species, and as far as we can see from the specimen image by Lu and Du (2020), N. bispina is externally distinguishable from this species by base of discal cell of the hindwing being concolorous with the ground color.	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
F362FB0FF6C45C70B37002DBB5C72348.taxon	distribution	Distribution. Japan, mainland China, Taiwan, Korea, Russia (southeast), India.	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
F6509A54AAB65E7DA9AF673568C41C33.taxon	materials_examined	Material examined. Japan: 1 ♂, Marumori, Onikobe, Narugo, Miyagi Pref., 30 Jul. 1997, M. Tanaka leg. (NSMT-I-L- 75588); 1 ♂, Kirei-pass, Sumison, Miyazaki Pref., 9 Jul. 1992, Y. Yanagita leg. (NSMT-I-L- 75592); 1 ♀, Hiromorigawa, Akita Pref., 14 Aug. 1988, A. Sasaki leg. (NSMT-I-L- 75583); 1 ♀, Garo-Kyo, Fujisato Town, Akita Pref., 28 Jul. 2002, A. Sasaki leg. (NSMT-I-L- 75584); 1 ♀, Yoroibata-Dam, Tazawako Town, Akita Pref., 26 Aug. 1989, A. Sasaki leg. (NSMT-I-L- 75585); 2 ♂, Tose, Tamagawa, Tazawako Town, Akita Pref., 21 Aug. 1993, A. Sasaki leg. (NSMT-I-L- 75586, 75587); 1 ♂, Futamata-Spa, Fukushima Pref., 6 Aug. 1967, T. Ebato leg. (NSMT-I-L- 75462); 1 ♀, Houshi, Gunma Pref., 19 Jul. 1957, T. Ebato leg. (NSMT-I-L- 75466); (NSMT-I-L- 75472), 1 ♂, Kawaburu-Spa, Gunma Pref., 1 Jul. 1967, T. Ebato leg.; 1 ♂, Mt Takao, Tokyo To, 23 Jun. 1951, T. Haruta leg. (NSMT-I-L- 75479); 1 ♀, ditto, 10 Jul. 1960, T. Ebato leg. (NSMT-I-L- 75463); 1 ♂, ditto, 26 Jun. 1959, T. Ebato leg. (NSMT-I-L- 75464); 1 ♀, ditto, 27 Jun. 1959, T. Ebato leg. (NSMT-I-L- 75474); 1 ♂, Nippara, Tokyo To, 8 Aug. 1961, T. Ebato leg. (NSMT-I-L- 75465); 1 ♀, same data as for preceding (NSMT-I-L- 75477); 2 ♂, ditto, 2 Sep. 1961, T. Ebato leg. (NSMT-I-L- 75475, 75476); 1 ♀, Mt Mitake, Tokyo To, 27 Aug. 1960, T. Maenami leg. (NSMT-I-L- 75486); 1 ♂, ditto, 20 Jun. 1996, U. Jinbo leg. (NSMT-I-L- 36059); 1 ♀, Ohnita, Ohme City, Tokyo To, 18 Aug. 1996, U. Jinbo leg. (NSMT-I-L- 75634); 1 ♂, Yuzurihara, Yamanashi Pref., 1 Sep. 1945, T. Ebato leg. (NSMT-I-L- 75471); 1 ♂, ditto, 2 Sep. 1945, T. Ebato leg. (NSMT-I-L- 75469); 1 ♂, ditto, 5 Sep. 1945, T. Ebato leg. (NSMT-I-L- 75470); 1 ♂, ditto, 23 Sep. 1954, T. Ebato leg. (NSMT-I-L- 75468); 1 ♂, Sagashio-Spa, Yamanashi Pref., 9 Aug. 1969, T. Ebato leg. (NSMT-I-L- 75473); 2 ♂, Ashiyasu, Yamanashi Pref., 16 Jul. 1977, T. Ebato leg. (NSMT-I-L- 75480, 75482); 1 ♂, ditto, 6 Aug. 1977, T. Ebato leg. (NSMT-I-L- 75483); 1 ♀, ditto, 19 Jul. 1980, T. Ebato leg. (NSMT-I-L- 75478); 1 ♂, Nishiyama-Spa, Yamanashi Pref., 17 Aug. 1981, T. Ebato leg. (NSMT-I-L- 75481); 1 ♀, Hirayu, Gifu Pref., 7 Aug. 1953, T. Haruta leg. (NSMT-I-L- 75467); 1 ♂, Gujo-Rokunori, Gifu Pref., 1 Jul. 1966, S. Sawatani leg. (NSMT-I-L- 75589); 1 ♂, Osugi-dani, Wakayama Pref., 4 Aug. 1976, S. Nakatani leg. (NSMT-I-L- 75484); 1 ♂, Shimakawa-Osugi-dani, Wakayama Pref., 5 Jul. 1975, S. Nakatani leg. (NSMT-I-L- 75485); 1 ♂, Tatsumi-toge, Tottori City, Tottori Pref., 35.32 ° N, 134.01 ° E, 670 m, 2 Jul. 2019, Y. Matsui leg.; 2 ♀ 3 ♂, Mt Daisen, Kofu Town, Tottori Pref., 35.35 ° N, 133.55 ° E, 910 m, 30 Sep. - 29 Oct. 2017, and 18 Apr. 2018 (F 1 emerged), Y. Matsui leg.; 1 ♂, Suemochi, Shikano Town, Tottori City, Tottori Pref., 35.45 ° N, 134.09 ° E, 220 m, 1 Jun. 2020, Y. Matsui leg.; 1 ♂, Kochi, Shikano Town, Tottori City, Tottori Pref., 35.40 ° N, 134.00 ° E, 495 m, 12 Jun. 2020, Y. Matsui leg.; 2 ♀ 1 ♂, ditto, 12 - 16 Aug. 2020 (F 1 emerged), Y. Matsui leg.; 1 ♂, Sourokubara, Tottori City, Tottori Pref., 35.46 ° N, 134.11 ° E, 110 m, 19 Jul. 2020 (larvae: collected from Rubus buergeri), 20 Aug. 2020 (emerged), Y. Matsui leg.; 1 ♀ 1 ♂, Toyofusa, Daisen Town, Tottori Pref., 35.41 ° N, 133.55 ° E, 680 m, 2 Sep. 2020 (larvae: collected from R. palmatus), 19 - 20 Oct. 2020 (emerged) Y. Matsui leg.; 1 ♂, Yakawa, Okuizumo Town, Shimane Pref., 35.10 ° N, 133.13 ° E, 680 m, 6 Sep. 2013, Y. Matsui leg.; 2 ♂, Omogokei, Kumakogen Town, Ehime Pref., 33.72 ° N, 133.10 ° E, 700 m, 8 Jun. 2019, Y. Matsui leg.; 1 ♂, Shimomyo, Aira-Cho, Kagoshima Pref., 28 May 1992, Y. Yanagita leg. (NSMT-I-L- 75590); 1 ♂, Tobi, Miyanojo-cho, Kagoshima Pref., 26 May 1992, Y. Yanagita leg. (NSMT-I-L- 75591); 1 ♀, Mt Ishizukadake, I. Yakushima, Kagoshima Pref., 5 - 6 Aug. 1958, B. T. leg. (NSMT-I-L- 75599); 1 ♂, same data as for preceding (NSMT-I-L- 75600); 4 ♂, ditto, 17 Jul. 1970, K. Tobi leg. (NSMT-I-L- 75601 to 75604); 1 ♀, same data as for preceding (NSMT-I-L- 75605).	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
F6509A54AAB65E7DA9AF673568C41C33.taxon	diagnosis	Diagnosis. Adult (Fig. 2 E, F). Forewing length 15.5 - 18 mm. This species is similar to N. tristalis and N. inferior, but can be distinguished by the following characters: forewing longer; vertex scales brown, darker than in N. inferior; cilia cream white at Cu 2 to A 1 + 2 for forewing, Cu 2 to CuP for hindwing; subdiscal white spot of forewing quadrilateral or rounded, rather distinct; base of discal cell of hindwing partially white; gnathos of male genitalia slender and elongated (Fig. 5 B); valva of male genitalia broader than in N. tristalis and N. inferior, with straight dorsal margin subapically (Fig. 5 C); anterior apophysis of female genitalia slightly curved, expanded near bases, but not triangular (Fig. 8 B); signum of female corpus brusae larger than that of N. tristalis and N. inferior (diameter 0.16 - 0.19 mm), with sharp projections at both edges of posterior margin (Fig. 8 C). Ullah et al. (2017) provided diagnostic characters to distinguish this species from N. occultalis. Lu and Du (2020) mentioned genital differences between these species, as far as we can see, the specimen image of N. bispina by Lu and Du (2020) is externally distinguishable from this species by the cilia of each wing being concolorous with the ground color.	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
F6509A54AAB65E7DA9AF673568C41C33.taxon	distribution	Distribution. Japan, mainland China, Taiwan, Korea, Russia (southeast), Southeast Asia, Nepal, India.	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
C9D9CE9CFFB3503B98FD90FDE60D40F4.taxon	etymology	Etymology. The specific epithet refers to the darker wing color in comparison to that of N. inferior, and the habitat of this species is a shaded place.	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
C9D9CE9CFFB3503B98FD90FDE60D40F4.taxon	diagnosis	Diagnosis. This new species is similar to N. inferior and N. quadrimaculalis, also distributed in Japan, but it can be distinguished by the following characters: length of forewing 12.0 - 13.0 mm; vertex with erect, dull-orange scales; subdiscal white spot of forewing rounded, small, and blurry; base of discal cell of hindwing identical to ground color; dorsal margin of valva of male genitalia slightly incurved subapically; anterior apophysis of female genitalia straight and narrow; signum of female genitalia nearly elliptical, larger than in N. inferior (diameter 0.09 - 0.14 mm). This species is also similar to N. occultalis and N. bispina distributed in China, but N. occultalis has the following differences: subdiscal white spot of forewing narrowed or elongated, tuba analis of male genitalia sclerotized, gnathos of male genitalia elongated and narrow at the base; N. bispina exhibits the following differences: gnathos of male genitalia absent, phallus of male genitalia with a hook-shaped cornutus, corpus bursae of female genitalia with two thorn-like signa. From N. hortulatoides, the new species can be easily distinguished by the wing maculation.	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
C9D9CE9CFFB3503B98FD90FDE60D40F4.taxon	description	Description (Fig. 2 A, B). Head: frons brownish grey, smooth. Vertex with erect, dull-orange scales. Labial palpus upturned, dorso-laterally dark brown, ventro-mesally pale white. Antenna dark brown; flagellum filiform with golden cilia ca 1 / 4 the diameter of flagellum in male. Thorax and abdomen: dorsally brownish grey; patagium and tegula with ochreous brown. Ventrally milky white. Wings: length of forewing 12.0 - 13.0 mm. Ground color of both wings brownish grey, with a large comma-shaped white spot at end of discal cell (the bases of R 5 to M 3), that of the hindwing somewhat small; cilia concolorous with ground color; postmedial line obscure. Subdiscal white spot of forewing rounded, small and blurry. Base of discal cell of hindwing concolorous with ground color. Male genitalia (Fig. 3 A-C): uncus short, subtrapezoid, blunt on posterior margin. Gnathos nearly triangular, short and small, apex blunt. Tuba analis elongate, not sclerotized, length ca 0.6 times that of valva. Transtilla subtriangular, completely connected medially. Saccus short, anterior margin rounded. Valva somewhat narrow, length ca 3.3 times that of width, dorsal margin slightly incurved subapically; costa more or less inflated with several setae at apex; clasper large, down curved, the apex blunt. Phallus cylindrical, without cornutus. Female genitalia (Fig. 6 A-C): ovipositor lobe oblong, setose. Anterior apophysis ca 2 times length of posterior apophysis, narrow, almost straight, with triangular expansion near base. Antrum trapezoid, sclerotized at collar. Ductus bursae almost equal in length to corpus bursae, membranous with wrinkles, slightly narrowed at posterior end. Corpus bursae pear-shaped, membranous; signum nearly elliptical, sparsely covered with denticles.	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
C9D9CE9CFFB3503B98FD90FDE60D40F4.taxon	biology_ecology	Biology. In Honshu, Japan, adults are found in May to September, and they are considered bivoltine. They appear to be hardly attracted to light. Larvae feed on Rubus buergeri and the middle instar larvae overwinter in its leaves.	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
C9D9CE9CFFB3503B98FD90FDE60D40F4.taxon	distribution	Distribution. Japan: Honshu (Tokyo, Shizuoka, Aichi, Tottori, Yamaguchi), Kyushu (Kagoshima), Ryukyu Islands (Yakushima, Amamioshima, Tokunoshima, Okinawajima).	en	Matsui, Yuki, Naka, Hideshi, Jinbo, Utsugi (2021): DNA barcoding and morphology reveal a new cryptic species of Nagiella (Lepidoptera, Crambidae, Spilomelinae) from Japan. ZooKeys 1023: 171-192, DOI: http://dx.doi.org/10.3897/zookeys.1023.60934, URL: http://dx.doi.org/10.3897/zookeys.1023.60934
