taxonID	type	description	language	source
0F7C87E7EF46FE57FF708B1CFB7FF9FF.taxon	diagnosis	Diagnosis. Apical margin of frontoclypeal region feebly trilobed and more than twice as wide as the base; antennal grooves absent; male protarsomere I simple with basally curved sex combs; tarsal claws simple; tarsomere IV excavate-emarginate; metacoxal plates medially 3.0 – 6.0 times wider than laterally; last visible ventrite strongly produced; lateral surfaces of meso- and metatibiae with setae and transverse rows of spine combs; metathoracic episterna parallel-sided; male aedeagus dorsoventrally compressed, with laterally attached secondary lateral lobes; median lobe simple, with moderately and narrowly bifurcate apices; lateral lobes simple, entire; flagellum simple (Muona 1993, 2011; Otto 2016). These diagnostic characteristics, especially the presence of excavate-emarginate tarsomere IV and parallelsided metathoracic episterna will distinguish the group from any genera within the tribe Nematodini. The new identification key will largely feature the overall length of the femoral ridge at the base of the hypomera where the femur rests when legs are reposed as a means to diagnose each species present in this region.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF40FE53FF708FA3FD1AFBAB.taxon	description	Fig. 6 – 8	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF40FE53FF708FA3FD1AFBAB.taxon	diagnosis	Diagnosis. Elongate and strongly-developed incomplete femoral ridge extending about 1 / 2 the length of the hypomeral base will readily distinguish the species from other Nematodes species present in the Nearctic region. Specimens examined. Sixty-three specimens were available for study: FLORIDA: “ FLORIDA: Gainesville, 3517 NW 10 TH Ave., 17 - V- 1993, R. E. Woodruff, blacklight trap ” (1, FSCA); “ FL: Alachua Co.:, Gainesville, NW 51 st Terrace, 6 May 2009, coll. O. J. Blanchard ” (‘ st’ superscript on label) / “ black, light ” (1, FSCA); “ FL: Alachua Co.:, Gainesville, NW 51 st Terrace, 13 May 2009, coll. O. J. Blanchard ” (‘ st’ superscript on label) / “ black, light ” (1, FSCA); “ Florida: Alachua Co., Paynes Prairie Preserve, State Park campground, May 3, 2012 at light, Kyle E. Schnepp ” (1, KESC); “ Florida: Alachua Co., San Felasco Preserve, April 5 – May 6, 2014, Lindgren funnel trap, Kyle E. Schnepp ” (1, KESC); “ Florida: Alachua Co., San Felasco Hammock, Preserve State Park, April 15, 2014, Kyle E. Schnepp ” / “ beaten from fallen, chinkapin oak branch, Quercus muehlenbergii ” (1, KESC); “ Florida: Alachua Co., 3.4 miles WNW Archer, March 28, 2016 beaten, from dead oak branch, Kyle E. Schnepp ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2017 ” (‘ 17 ’ handwritten on label) (1, KESC); “ Florida: Suwanee Co., ~ 7 miles W White Springs, May 16 – June 20, 2017, Lindgren funnel trap, Kyle E. Schnepp ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2017 ” (1, KESC); “ Florida: Liberty Co., Torreya S. P. at light, May 11, 2018, Kyle E. Schnepp ” (1, KESC); “ Florida: Alachua Co., San Felasco H. P. S. P., April 24 – May 5 2021, Lindgren funnel trap, Kyle E. Schnepp ” (2, KESC). GEORGIA: “ GEORGIA: Chatham Co., 1.4 km WNW Port Wentworth, 32.15091, - 81.17832, 13 m, 22 Mar – 6 Apr 2016, LFT, R. Morgan, BSF # 66614 ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2017 ” (1, CMNH); “ GEORGIA: Chatham Co., 1.4 km WNW Port Wentworth, 32.15091, - 81.17832, 13 m, 5 – 19 Apr 2016, LFT, R. Morgan, BSF # 66615 ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2017 ” (1, CMNH); “ GEORGIA: Chatham Co., 1.4 km WNW Port Wentworth, 32.15091, - 81.17832, 13 m, 5 – 19 Apr 2016, Lindgren FT, R. Morgan, BSF # 66618 ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2017 ” (2, CMNH); “ GEORGIA: Chatham Co., 1.4 km WNW Port Wentworth, 32.15091, - 81.17832, 13 m, 16 Apr – 2 May 2016, LFT, R. Morgan, BSF # 66583 ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2017 ” (1, CMNH); “ GEORGIA: Chatham Co., 1.4 km WNW Port Wentworth, 32.15091, - 81.17832, 13 m, 19 Apr – 3 May 2016, LFT, R. Morgan, BSF # 66613 ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2017 ” (3, CMNH); “ GEORGIA: Chatham Co., 1.9 km N Garden City, 32.13098, - 81.15114, 3 m, 19 Apr – 3 May 2018, LFT, S. Davis, BSF # 78777 ” (1, CMNH); “ GEORGIA: Chatham Co., 0.9 km ENE Port Wentworth, 32.15005, - 81.15388, 3 m, 19 Apr – 3 May 2018, LFT, S. Davis, BSF # 78796 ” (2, CMNH); “ GEORGIA: Chatham Co., 2.3 km NW Garden City, 32.1235, - 81.17599, 5 m, 17 – 31 May 2018, LFT, S. Davis, BSF # 79491 ” (1, CMNH); “ GEOR- GIA: Chatham Co., 0.8 km ENE Port Wentworth, 32.15031, - 81.15459, 3 m, 17 – 31 May 2018, LFT, S. Davis, BSF # 79559 ” (1, CMNH). ILLINOIS: “ ILLINOIS: White County, 1.2 km SSE Grayville, 38.248574, - 87.98698, 125 m, 4 – 8 Jun 2018, T. Vorce, LFT, BSF # 82162 ” (1, CMNH). INDIANA: “ Indiana: Morgan Co., June 12 – July 10, 2009, purple sticky trap, Kyle E. Schnepp ” (1, KESC); “ Indiana: Morgan Co., July 10 – 20, 2009, purple sticky trap, Kyle E. Schnepp ” (1, KESC); “ Indiana: Morgan Co., June 14 – 29, 2010, black panel trap EtOH, Kyle E. Schnepp ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2014 ” (1, KESC); “ Indiana: Morgan Co., June 29 – July 11, 2010, black panel trap EtOH, Kyle E. Schnepp ” (1, KESC). KANSAS: “ USA: Kansas: Labelle Co., Big Horn Resevoir, NW of dam, 37 ° 16.55 ’ N 95 ° 28.43 ’ W, 2 – 13 - VI- 2005 G. A. Salsbury, ex. canopy trap KAN 1 S 05 100 ” (1, GERP); “ USA: Kansas: Johnson Co., Overland Park Arboretum, 38 ° 48.094 ’ N 94 ° 41.310 ’ W, 12 – 20 - VI- 2006, A. Cruz ex., canopy trap KAN 1 S 06 067 ” (1, GERP); “ KANSAS: Wyandotte Co., 3.5 km WSW Kansas City, 39.10374, - 94.66579, 268 m, 17 May – 7 June 2018, LFT, S. Roberts, BSF # 83457 ” (2, CMNH); “ KANSAS: Jefferson Co., 2.5 km N Grantville, 293 m, 39.105735, - 95.557166, 10 Jun – 1 Jul 2019, LFT, S. Pellato, BSF # 91322 ” (1, CMNH); “ KANSAS: Jefferson Co., 2.5 km N of Grantville, 39.105735, - 95.557166, 293 m, 1 – 8 Jul 2019, LFT, S. Pellato, BSF # 91323 ” (1, CMNH). LOUISIANA: “ LA: Baton Rouge, Place DuPlantier, Apts. V. 26.85, Coll. E. G. Riley ” (date handwritten) / “ window trap ” (3, GERP). MISSOURI: “ MO: Jackson Co., Jackson, June 1998 ” (2, GERP); “ MISSOURI: St. Louis Co., 2.4 km E Maryland Heights, 38.71261, - 90.40263, 193 m, 14 – 27 June 2016, LFT, L. Trevathan, BSF # 70244 ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2017 ” (1, CMNH). OHIO: “ OHIO, Scioto Co., Shawnee St. For., IX- 1980 ” (“ IX- ” and “ 80 ” were handwritten) / “ collectors:, B & BValentine ” / “ Valentine, Coll’n, Rec. 2016 ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2019 ” (10, FSCA); “ USA: OH Cuyahoga Co., 09 - VI- 2000, on dead Hickory, colld by RJBuss ” / “ Nematodes, collaris? ” (handwritten in pencil) (1, GERP); “ OH: Preble Co., Camden, 29 June 2001, Michael Ulyschen, Blacklight ” (1, GERP); “ OHIO: Allen County, 3 km SW Lima, 263 m, 40.7273, - 84.1348, 19 Jun – 3 Jul 2018, LFT, C. Poe, BSF # 82558 ” (1, CMNH). SOUTH CAROLINA: “ Dovehaven, ” 7 mi., N. E. of Pickens, S. C., 11. VII. 1979, H. L. Dozier ” (“ 11. VII. ” and “ 79 ” were handwritten) (1, FSCA); “ South Carolina: Pickens Co., ~ 5 miles north Clemson, June 20, 2018, under bark of fallen tree, Kyle E. Schnepp ” / “ Nematodes, atropos, (Say), Det. R. L. Otto, 2019 ” (2, KESC). TEXAS: “ TEXAS: Sabine Co., 9 mi., E. Hemphill “ Beech, Bottom ”, V- 8 – 21 - 1989, R. Anderson & E. Morris, Malaise trap, beech-magnolia forest ” (2, GERP). VIRGINIA: “ VA: Appomattox Co., Appomattox, 29 – 30 June 2000, Robert Vigneault ” (4, GERP).	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF40FE53FF708FA3FD1AFBAB.taxon	description	Redescription. Length, 6.0 – 10.0 mm. Width, 1.5 – 2.0 mm. Body subcylindrical, elongate; uniformly dark brown; antennae dark brown; legs including tarsi dark brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (Fig. 6). Head: Subspherical; integument closely punctate, somewhat dullish; frons convex, with median, round, shallow fovea above frontoclypeal region; apical margin of frontoclypeal region weakly trilobed, about 2 times wider than base; mandibles stout, bidentate, densely punctate. Antenna: Filiform to weakly serriform from antennomeres III – XI, attaining nearly 1 / 3 the length of the body; antennomere III longer than IV; antennomeres IV – V each sub-equal, longer than wide, each slightly shorter than VI; antennomeres VI – X each sub-equal, longer than wide; antennomere XI stout, slightly longer than X, less than 2.0 times longer than wide. Pronotum: Integument dullish, densely punctate to rugose; longer than wide, with moderate, sharp hind angles; lateral sides basally constricted above pronotal hind angles, parallel-sided, cranially arcuate; disc convex with shallow median groove and pair of shallow, circular fovea; base sinuous. Scutellar shield: Elongate, sub-triangular, setose, shallowly punctate and distally rounded. Elytra: Distinctly striate; interstices elevated; integument somewhat shiny, transversely rugose at basal 1 / 2, closely punctate at apical 1 / 2. Legs: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I – III simple; metatarsomere IV excavate-emarginate; metatarsomere V short with simple claws. Venter (Fig. 7): Closely punctate, with short, recumbent yellowish setae; hypomeron with basally obliterated lateral antennal grooves; strongly-developed femoral ridge incomplete, elongate, extending about 1 / 2 the length of the hypomeral base; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially more than 3.0 – 6.0 times wider than laterally; last abdominal ventrite apically produced. Aedeagus (Fig. 8): Basal piece longer than wide, laterally parallel-sided, dorsally open, apically rounded; remaining parts elongate, constricted laterally just above the basal piece, laterally sinuous; parameres elongate, apically rounded, triangular-shaped lateral tooth present near apices; median lobe elongate and narrow, apically pointed, deeply and narrowly bifid, longer than parameres.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF40FE53FF708FA3FD1AFBAB.taxon	distribution	Distribution. This widespread, somewhat common eucnemid species has been taken from Alabama, Arkansas, Connecticut, Florida, Georgia, Illinois, Indiana, Kansas, Kentucky, Louisiana, Maryland, Massachusetts, Mississippi, Missouri, New Jersey, New York, North Carolina, Ohio, Oklahoma, Pennsylvania, South Carolina, Tennessee, Texas, Virginia, and West Virginia (Otto and Karns 2017).	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF40FE53FF708FA3FD1AFBAB.taxon	biology_ecology	Biology. Specimens were reared from dead Carya sp. Nutt. (Juglandaceae) limbs as well as captured in Lindgren funnel traps and other types of flight intercept traps. Five specimens were taken from lights or blacklights in Florida. Two other specimens were beaten from a dead oak limbs. Immature stages were briefly noted by Van Horn (1909), but never formally described.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF42FE5CFF708B09FB26FB63.taxon	description	Fig. 9 – 11	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF42FE5CFF708B09FB26FB63.taxon	diagnosis	Diagnosis. Very short, poorly-developed, incomplete femoral ridge at the base of the hypomera will readily distinguish N. collaris from all other Nematodes species present in the region. Specimens examined. Ten specimens were available for study: FLORIDA: “ FLORIDA: Alachua Co., NW Gainesville, 1 - IV- 1974, J. B. Heppner ” / “ Nematodes, collaris, Bonvouloir, Det. R. L. Otto, 2019 ” (1, FSCA). ILLI- NOIS: “ ILLINOIS: St. Clair County, 1.1 km NNW East St. Louis, 36.63384, - 90.15347, 128 m, 2 – 14 Jun 2016, Lindgren FT, S. Mills, BSF # 71866 ” (1, GERP); “ ILLINOIS St. Clair Co., 1.1 km NNW East St. Louis, 36.63384, - 90.15347, 128 m, 14 – 30 Jun 2016, LFT, S. Mills, BSF # 71543 ” (1, GERP). INDIANA: “ Indiana: Morgan Co., June 12 – July 10, 2009, purple sticky trap, Kyle E. Schnepp ” / “ Nematodes, collaris, Bonvouloir, Det. R. L. Otto, 2014 ” (1, KESC); “ Indiana: Montgomery Co., Shades State Park, June 13 – July 1, 2013, Lindgren funnel trap, Kyle E. Schnepp ” (1, KESC). OHIO: “ OHIO, Scioto Co., Shawnee St. For., IX- 1980 ” (“ IX- ” and “ 80 ” were handwritten) / “ collectors:, B & BValentine ” / “ Valentine, Coll’n, Rec. 2016 ” / “ Nematodes, collaris, Bonvouloir, Det. R. L. Otto, 2019 ” (2, FSCA); “ OHIO: Sandusky Co., 1.4 km NNW Clyde, 41.3153, - 82.9843, 200 m, 14 – 30 June 2016, LFT, C. Poe, BSF # 68617 ” / “ Nematodes, collaris, Bonvouloir, Det. R. L. Otto, 2017 ” (1, CMNH); “ OHIO: Lucas County, 4.8 km SSW Sylvania, 41.6792, - 83.7361, 250 m, 22 Jun – 5 Jul 2016, LFT, C. Poe, BSF # 68323 ” (1, GERP); “ OHIO: Huron County, 0.8 km WSW Willard, 41.0514, - 82.7373, 285 m, 26 Jun – 19 Jul 2018 ” / “ cross-vane panel, trap, C. Mitchell, BSF # 82591 ” (1, CMNH).	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF42FE5CFF708B09FB26FB63.taxon	description	Redescription. Length, 6.5 – 9.0 mm. Width, 1.0 – 2.0 mm. Body subcylindrical, elongate; either uniformly dark brown-black or bicolored with dark orange head / pronotum and dark brown-black elytra / venter; antennae dark brown; legs including tarsi dark brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (Fig. 9). Head: Subspherical; integument closely punctate, somewhat dullish; frons convex, with median, round, deep fovea above frontoclypeal region; apical margin of frontoclypeal region weakly trilobed, about 2 times wider than base; mandibles stout, bidentate, densely punctate. Antenna: Filiform to weakly serriform from antennomeres III – XI, attaining nearly 1 / 3 the length of the body; antennomere III longer than IV; antennomeres IV – V each sub-equal, longer than wide, each slightly shorter than VI; antennomeres VI – X each sub-equal, longer than wide; antennomere XI elongate, longer than X, more than 2.0 times longer than wide. Pronotum: Integument dullish, densely punctate to rugose; longer than wide, with moderate, sharp hind angles; lateral sides parallel-sided, cranially arcuate; disc convex with deep median groove and pair of deep, circular fovea; base sinuous. Scutellar shield: Elongate, sub-triangular, setose, shallowly punctate and distally rounded. Elytra: Distinctly striate; interstices elevated; integument somewhat shiny, transversely rugose at basal 1 / 2, closely punctate at apical 1 / 2. Legs: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I – III simple; metatarsomere IV excavate-emarginate; metatarsomere V short with simple claws. Venter (Fig. 10): Closely punctate, with short, recumbent yellowish setae; hypomeron with basally obliterated lateral antennal grooves; poorly-developed femoral ridge, short, incomplete, extending about 1 / 3 the length of the hypomeral base; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially more than 3.0 – 6.0 times wider than laterally; last abdominal ventrite apically produced. Aedeagus (Fig. 11): Basal piece longer than wide, laterally parallel-sided, dorsally open, apically rounded; remaining parts elongate, parallel-sided; parameres elongate, apically rounded, triangular-shaped lateral tooth present near apices; median lobe elongate and narrow, apically pointed, deeply and narrowly bifid, longer than parameres.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF42FE5CFF708B09FB26FB63.taxon	distribution	Distribution. This very rarely seen eucnemid species was previously recorded in Illinois, Indiana, Kansas, Louisiana, Maryland, Ohio, South Carolina (Otto and Karns 2017) and now newly recorded for the first time in Florida. A single specimen from Missouri was misidentified as N. collaris. There are currently no Missouri records for the species.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF42FE5CFF708B09FB26FB63.taxon	biology_ecology	Biology. Adult specimens were taken from Lindgren funnel traps, purple sticky trap, cross-vane panel trap, canopy traps, flight intercept traps, and canopy Malaise traps. Immature stages remain unknown. Note. According to Muona (pers. comm.), the editors had accidentally switched the illustrations of the aedeagus between this species and that of N. humphreyi in the 2000 Nearctic revision of the family.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF4DFE5EFF708BB1FBCEFD5F.taxon	description	Fig. 12 – 14	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF4DFE5EFF708BB1FBCEFD5F.taxon	diagnosis	Diagnosis. Complete femoral ridge extending to the lateral side of the pronotum will distinguish N. humphreyi from all known Nematodes species present in the Nearctic region. Specimens examined. Twenty-one specimens were available for study: FLORIDA: “ FLORIDA: Dade Co., Camp Mahachee, nr., Matheson Hammock, 12 - IV- 1991, J. Gleason, Blacklight trap ” (1, GERP); “ FLORIDA: Miami- Dade Co., Miami, Greynolds Park West, 25.944, - 80.1556, 5 - V- 2011, P. Perez, M. YongCong, J. Garcia, lindgren funnel trap w / phoebe oil ” (1, FSCA); “ FLORIDA: Broward Co., Dania Beach, Secret, Woods Nature Ctr., 19 - IV- 2013, M. DaCosta, J. Garcia Stickyboard trap ” (1, FSCA); Florida: Monroe Co., Upper Key Largo, May 25, 2014 at light, Kyle E. Schnepp ” / “ Nematodes, humphreyi, Muona, Det. R. L. Otto, 2014 ” (1, KESC); “ Florida: Monroe Co., Upper Key Largo, May 15, 2016 stump, of small tree at night, Kyle E. Schnepp ” (1, GERP; 4, KESC); “ Florida: Monroe Co., Upper Key Largo, May, 15, 2016 at light, Kyle E. Schnepp ” (1, KESC); “ Florida: Monroe Co., Upper Key Largo, May 29, 2016 at light, Kyle E. Schnepp ” (1, GERP; 1, KESC); “ Florida: Monroe Co., Upper Key Largo, May 30, 2016 at light, Kyle E. Schnepp ” (1, GERP; 5, KESC); “ Florida: Monroe Co., Upper Key Largo, May 30, 2016, on cut tree at night, Kyle E. Schnepp ” (2, KESC); “ Florida: Monroe Co., Upper Key Largo, May 18, 2018 at light, Kyle E. Schnepp ” (1, KESC).	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF4DFE5EFF708BB1FBCEFD5F.taxon	description	Redescription. Length, 5.0 – 7.5 mm. Width, 1.0 – 2.0 mm. Body subcylindrical, elongate; uniformly dark brown; antennae dark reddish-brown; legs including tarsi dark reddish-brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (Fig. 12). Head: Subspherical; integument closely punctate / rugose to confluently rugose, somewhat dullish; frons convex, with median, round, shallow fovea above frontoclypeal region; apical margin of frontoclypeal region weakly trilobed, about 2 times wider than base; mandibles stout, bidentate, densely punctate. Antenna: Filiform to weakly serriform from antennomeres III – XI, attaining nearly 1 / 3 the length of the body; antennomere III longer than IV; antennomeres IV – V each sub-equal, slightly longer than wide, each shorter than VI; antennomeres VI – X each sub-equal, longer than wide; antennomere XI elongate, slightly longer than X, more than 2.0 times longer than wide. Pronotum: Integument somewhat dullish, densely punctate to rugose; longer than wide, with moderate, sharp hind angles; lateral sides parallel-sided at basal half, arcuate at cranial 1 / 2; disc convex with shallow median groove extending through entire length and pair of shallow, circular fovea; base sinuous. Scutellar shield: Elongate, sub-triangular, setose, shallowly punctate and distally rounded. Elytra: Distinctly striate; interstices elevated; integument somewhat shiny, transversely rugose throughout. Legs: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I – III simple; metatarsomere IV excavate-emarginate; metatarsomere V short with simple claws. Venter (Fig. 13): Closely punctate, with short, recumbent yellowish setae; hypomeron with indicated, medially undefined lateral antennal grooves; well-developed femoral ridge complete, extending to the lateral side of the pronotum; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially more than 3.0 – 6.0 times wider than laterally; last abdominal ventrite apically produced. Aedeagus (Fig. 14): Basal piece longer than wide, laterally parallel-sided, dorsally open, apically rounded; remaining parts elongate, basally bulbous above basal piece, laterally sinuous; parameres elongate, apically rounded, hook-like lateral tooth present base of parameres; median lobe elongate and narrow, apically pointed, deeply and narrowly bifid, slightly longer than parameres.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF4DFE5EFF708BB1FBCEFD5F.taxon	distribution	Distribution. This precinctive Florida eucnemid species is known from a small number of locations in southern Florida.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF4DFE5EFF708BB1FBCEFD5F.taxon	biology_ecology	Biology. One specimen has been taken from a blacklight trap in Dade County. Two specimens were taken from a small cut tree at night. Eleven specimens were taken from lights in the Upper Key Largo. Five specimens were taken from a stump of a small tree at night. Immature stages remain unknown. Note. According to Muona (pers. comm.), the editors had accidentally switched the illustrations of the aedeagus between this species and that of N. collaris in the 2000 Nearctic revision of the family.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF4FFE44FF708DBDFAACF87B.taxon	description	Fig. 15 – 17	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF4FFE44FF708DBDFAACF87B.taxon	diagnosis	Diagnosis. A strongly-developed, elongate, incomplete femoral ridge will distinguish this eucnemid species from all known Nematodes species in the Nearctic region, except N. rugosipennis new species. A sub-equal antennomere V in relation to antennomere IV will further distinguish N. penetrans from N. rugosipennis new species. Specimens examined. Three hundred and seventy-five specimens were available for study: CANADA: QUÉ- BEC: “ CANADA:, Quebec, Oka, 4 July 1999, Robert Vigneault ” (1, GERP). USA: ARKANSAS: “ AR Little Rock, VI / 20 / 03 Hanging, sugar traps, Brian Baldwin ” (1, GERP). ILLINOIS: “ ILLINOIS: Clinton County, 6.2 km SSW Harbor Light, Bay, 38.66203, - 89.32758, 141 m, 25 May – 8 Jun 2018, S. Mills, LFT, BSF # 82133 ” (1, CMNH); “ ILLINOIS: St. Clair Co., 2.4 km WNW Signal Hill, 38.583173, - 90.083133, 124 m, 8 – 22 Jun 2018, S. Mills, LFT, BSF # 82234 ” (1, CMNH); “ ILLINOIS: Moultrie County, 4.6 km ESE Kirksville, Forest W. Wood Access, Area, 39.55210, - 88.62136, 194 m, 13 – 27 Jun 2018 ” / “ Lindgren funnel, trap, C. Wetzel, BSF # 81099 ” (1, CMNH). INDIANA: “ Indiana: Tippecanoe Co., July 3 – 6, 2007 red, Lindgren funnel trap, Kyle E. Schnepp ” (3, KESC); “ Indiana: Tippecanoe Co., July 25 – 27, 2008, white PVC trap, Kyle E. Schnepp ” (1, KESC); “ Indiana: Tippecanoe Co., ex: dead butternut, Juglans cinerea branch, Kyle E. Schnepp ” / “ wood collected fall, 2009 emerged, spring 2010 ” / “ Nematodes, penetrans, (LeConte), Det. R. L. Otto, 2019 ” (1, KESC). KANSAS: “ USA: Kansas: Jefferson Co., The Falin Property, 1.5 km, N jct. 94 th St. & Kingman, Rd. 39 ° 13.38 ’ N 95 ° 24.24 ’ W, 10 – 20 - VI- 2006 Z. H. Falin, ex. canopy FIT, near lower, meadow KAN 1 F 05 118 ” (1, GERP). MAINE: “ MAINE: Penobscot Co., 3.8 km W Bangor, 52 m, 44.79578, - 68.8247, 16 – 30 Jul 2018, LFT, R. Nyce, BSF # 81958 ” (1, CMNH). MISSOURI: “ MIS- SOURI: St. Louis Co., 2.4 km E Maryland Heights, 38.71261, - 90.40263, 193 m, 14 – 27 June 2016, LFT, L. Trevathan, BSF # 70244 ” / “ Nematodes, penetrans, (LeConte), Det. R. L. Otto, 2017 ” (2, CMNH). NEW JERSEY: “ NEW JERSEY: Monmouth, Co., 1.2 km S Farmingdale, 40.18605, - 74.16875, 30 m, 26 Jun – 9 Jul 2018, LFT, D. Armstrong, BSF # 80353 ” (1, CMNH). NEW YORK: “ NEW YORK: Franklin Co., 6.2 km ENE Hogansburg, 44.9875, - 74.5869, 48 m, 2 – 19 Jul 2019, LFT, A. Johnson, BSF # 86283 ” (1, CMNH); “ NEW YORK: Franklin Co., 0.9 km ENE Hogansburg, 44.9728, - 74.6517, 60 m, 3 – 18 Jul 2019, LFT, A. Johnson, BSF # 86310 ” (1, CMNH); “ NEW YORK: Franklin Co., 7.5 km ENE Hogansburg, 44.9484, - 74.5748, 62 m, 14 – 21 Aug 2019, LFT, A. Johnson, BSF # 88005 ” (1, CMNH); “ NEW YORK: Franklin Co., 5.2 km ENE Hogansburg, 44.9574, - 74.6012, 66 m, 14 – 21 Aug 2019, LFT, A. Johnson, BSF # 88019 ” (1, CMNH). OHIO: “ OHIO: Hamilton County, 4.1 km W Forest Park, 39.2884, - 84.551, 258 m, 12 – 28 Jun 2018, LFT, E. Larue, BSF # 82508 ” (5, CMNH). VERMONT: “ Vermont Orleans Co., Newport, PT- 3; Jules Rd; N 44.9853, W 72.12628; EAB Purple Trap, Emerald Ash Borer Pheromone; 5 / 30 / 2014 – 7 / 21 / 2014; Coll., USDA (5535) ” / “ Eucnemidae, Eschscholtz 1829, Det. E. R. Hoebeke, 2015 ” (5, CMNH); “ VERMONT: Chittenden, Co, 2.1 km NNE Burlington, 44.494317, - 73.208966, 33 m, 7 – 29 Jun 2018 ” / “ panel intercept, trap, E. Inoue, BSF # 80820 ” (4, CMNH); “ VERMONT: Windham Co., 2.5 km NW Rockingham, 43.20408, - 72.51019, 150 m, 26 Jun – 10 Jul 2018, LFT, L. Modesto, BSF # 80319 ” (1, CMNH). VIR- GINIA: “ VA: Appomattox Co., Appomattox, 14 – 28 June 1999, Robert Vigneault ” (1, GERP). WEST VIRGINIA: “ WEST VIRGINIA: Raleigh, Co, 7 km ENE Beckley, 37.7937, - 81.1116, 747 m, 6 – 18 Jul 2018, LFT, R. Hays, BSF # 80665 ” (1, CMNH). WISCONSIN: “ Jackson Co., WI, T 21 N R 24 W s 23, 27 VI – 3 VII 1977, Gypsy Moth- M. T. ” (handwritten) (1, WIRC); “ USA: WI: Green Co., Abraham’s Woods SNA, 42 ° 41 ’ 05 ” N / 89 ° 29 ’ 04 ” W, 11 July – 5 Aug. 2001, Jeffrey P. Gruber ” / “ flight intercept trap near, several fallen trees in, southern mesic hardwood, forest ” / “ Nematodes, penetrans, (LeConte), Det. R. L. Otto, 2015 ” (1, WIRC); “ USA: WI: Grant County, Wyalusing State Park, 42 ° 59.647 ’ N, 91 ° 06.648 ’ W, 18 – 26 July 2004, Coll. John J. Dorshorst ” / “ Flight intercept trap, baited with cantharidin, in southern mesic to, southern dry-mesic forest ” (3, WIRC); “ WI: Oconto Co., Rueckert’s private property, T 29 N R 16 E sec 16, 17 July 1996, Robert L. Otto ” / “ collected on, dead Acer ” (1, GERP); “ WI: Oconto Co., Rueckert’s private property, T 29 N R 17 E sec 16, 14 July 1997, Robert L. Otto ” / “ collected beneath, Acer bark ” (3, GERP); “ WI: Richland Co., LWRSWA, Lone Rock Unit, T 8 N / R 3 E / Sec. 3, July 1 – 5 1999, Alistair Ramsdale ” / “ Flight intercept, Trap, Oak, savanna ” (1, GERP); “ WI: Sauk Co., Pine Hollow, T 10 N / R 5 E / Sec. 4, July 7 – 13 1999, Alistair Ramsdale ” / “ Flight intercept, Trap, Mesic, Hardwood forest ” (1, GERP); “ WI: Fond du Lac Co., Long Lake, Kettle Moraine St. For., N 43.66355 °, W 088.1652 °, July 20, 2004, MDD & JMP ” / “ Girdled ash tree, EAB TT # 1 ” (handwritten) (1, WIRC); “ WI: Oconto Co., Rueckert’s private property, T 29 N R 16 E sec 16, 10 April 2007, Robert L. Otto ” / “ reared from larvae, in rotten Acer on, 13 May 2007 ” (1, GERP); “ WI: Oconto Co., Rueckert’s private property, N 44 ° 59 ’ W 88 ° 25 ’, 9 April 2009, Robert L. Otto ” / “ reared from larva, in Acer on, 7 May 2009 ” (10, GERP); “ WI: Oconto Co., Rueckert’s private property, N 44 ° 59 ’ W 88 ° 25 ’, 9 April 2009, Robert L. Otto ” / “ reared from larva, in Acer on, 9 May 2009 ” (2, GERP); “ WI: Oconto Co., Rueckert’s private property, N 44 ° 59 ’ W 88 ° 25 ’, 9 April 2009, Robert L. Otto ” / “ reared from larva, in Acer on, 16 May 2009 ” (1, GERP); “ WI: Forest Co., along County Hwy W, N 45.54697 ° / W 088.87092 °, 14 July 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, GERP; 1, WIRC); “ WI: Forest Co., along Interstate 8, N 45.64411 ° / W 088.66409 °, 21 July 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (2, GERP; 2, WIRC); “ WI: Forest Co., along State Hwy 139, N 45.76769 ° / W 088.69083 °, 4 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, GERP; 1, WIRC); “ WI: Forest Co., along County Hwy W, N 45.54697 ° / W 088 .. 87092 °, 11 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, GERP; 4, WIRC); “ WI: Forest Co., along Range Line Rd., N 45.60146 ° / W 088.93555 °, 11 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, GERP; 1, WIRC); “ WI: Forest Co., along Lundt Rd., N 45.58689 ° / W 088.86453 °, 11 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Potawatomi Trl., N 45.56523 ° / W 088.86207 °, 11 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Otter Creek Ln., N 45.60744 ° / W 088.80075 °, 12 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, GERP; 2, WIRC); “ WI: Forest Co., along Peshtigo Preserve Rd, N 45.56194 ° / W 088.57435 °, 17 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (4, WIRC); “ WI: Forest Co., along Goodman Park Rd., N 45.52111 ° / W 088.564901 °, 17 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Interstate 8, N 45.64411 ° / W 088.66409 °, 18 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (3, GERP; 12, WIRC); “ WI: Forest Co., along Peshtigo River Rd., N 45.60692 ° / W 088.59950 °, 18 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Browns Rd., N 45.65729 ° / W 088.59246 °, 18 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (2, WIRC); “ WI: Forest Co., along Interstate 8, N 45.65477 ° / W 088.59246 °, 18 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (2, WIRC); “ WI: Forest Co., along Spur Ln., N 45.64630 ° / W 088.56099 °, 19 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (4, WIRC); “ WI: Forest Co., along County Hwy G, N 45.65222 ° / W 088.78085 °, 19 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (2, WIRC); “ WI: Forest Co., along S. Town Rd., N 45.65641 ° / W 088.56300 °, 19 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Kalata Rd., N 45.67459 ° / W 088.65600 °, 19 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Double Bend Rd., N 45.74372 ° / W 088.72631 °, 20 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Hanson Flat, N 45.66603 ° / W 088.96741 °, 24 August 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Windsor Dam Rd., N 45.86884 ° / W 088.88438 °, 1 September 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (2, WIRC); “ WI: Forest Co., along Lundt Rd., N 45.58689 ° / W 088.86455 °, 3 September 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (3, WIRC); “ WI: Forest Co., along County Hwy W, N 45.54697 ° / W 088.87097 °, 3 September 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Potawatomi Trl., N 45.56523 ° / W 088.86207 °, 8 September 2009, Andy E. Anderson ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Interstate 8, N 45.64411 ° / W 088.66409 °, 9 September 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Forest Co., along Browns Rd., N 45.65729 ° / W 088.59246 °, 9 September 2009, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Door Co., EAB Trap # T 014697, N 44.98045055 °, W 087.31120038 °, September 21, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (1, GERP; 1, WIRC); “ WI: Door Co., EAB Trap # T 015991, N 45.14915940 °, W 087.15348867 °, September 21, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (3, WIRC); “ WI: Door Co., EAB Trap # T 015651, N 45.11035387 °, W 087.21469416 °, September 21, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Door Co., EAB Trap # T 014541, N 44.95521801 °, W 087.36600042 °, September 21, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (4, WIRC); “ WI: Door Co., EAB Trap # T 014239, N 44.92717752 °, W 087.3908807 °, September 21, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Door Co., EAB Trap # T 017944, N 45.40686160 °, W 086.87317260 °, September 22, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (2, WIRC); “ WI: Door Co., EAB Trap # T 017477, N 45.348739948 °, W 086.87006073 °, September 22, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (2, WIRC); “ WI: Door Co., EAB Trap # T 016989, N 45.28424241 °, W 087.06588454 °, September 22, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (2, WIRC); “ WI: Door Co., EAB Trap # T 016658, N 45.23784128 °, W 086.98765598 °, September 22, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Door Co., EAB Trap # T 015993, N 45.14882656 °, W 087.09470211 °, September 22, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (2, WIRC); “ WI: Door Co., EAB Trap # T 015652, N 45.12242680 °, W 087.19298151 °, September 22, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (8, WIRC); “ WI: Door Co., EAB Trap # T 016159, N 45.17046310 °, W 087.12216558 °, September 22, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Door Co., EAB Trap # T 015656, N 45.11044281 °, W 087.06869499 °, September 22, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Door Co., EAB Trap # T 015483, N 45.08630276 °, W 087.19392263 °, September 22, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (3, WIRC); “ WI: Door Co., EAB Trap # T 015484, N 45.08352122 °, W 087.16285209 °, September 22, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (3, WIRC); “ WI: Door Co., EAB Trap # T 014704, N 44.98264829 °, W 087.22309529 °, September 24, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (2, GERP; 3, WIRC); “ WI: Door Co., EAB Trap # T 015012, N 45.02535123 °, W 087.28214468 °, September 24, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (2, WIRC); “ WI: Door Co., EAB Trap # T 015168, N 45.04545975 °, W 087.25530423 °, September 24, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (1, WIRC); “ WI: Door Co., EAB Trap # T 014241, N 44.91611643 °, W 087.33672235 °, September 24, 2009, Robert L. Otto ” / “ Taken from EAB, prism trap baited, w / Manuka oil ” (2, WIRC); “ WI: Shawano Co., along County Road C, N 44.73930 °. W- 088.33105 °, EABT 012765, 21 June 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, GERP); “ WI: Oconto Co., along County Road G / M, N 45.00425 °. W- 088.24133 °, EABT 014676, 29 June 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, GERP); “ WI: Marinette Co., along County Road C, N 45.40716 °. W- 088.34933 °, EABT 017771, 1 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, GERP); “ WI: Marinette Co., along County Road F, N 45.39204 °, W- 088.35157 °, EABT 017607, 1 July 2010, Robert L. Otto ” / “ Taken from EAB, Prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Lafayette Co., along County Road U, N 42.54630 °. W- 090.23315 °, EABT 108073, 7 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, GERP; 36, WIRC); “ WI: Lafayette Co., along County Road O, N 42.52951 °. W- 090.25043 °, EABT 108146, 8 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, GERP); “ WI: Shawano Co., along County Road M, N 44.76789 °. W- 088.69987 °, EABT 012934, 12 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, GERP; 1, WIRC); “ WI: Shawano Co., along Maple Road, N 44.77976 °. W- 088.67004 °, EABT 013102, 12 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, GERP; 3 WIRC); “ WI: Marinette Co., along Packard Road, N 45.33680138 °, W- 087.69561374 °, EABT 017300, 12 July 2010, Randy LaPlant ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (2, WIRC); “ WI: Marinette Co., along Forest Road 901, N 45.32403046 °, W- 087.83637892 °, EABT 017112, 12 July 2010, Randy LaPlant ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Chapin Road, N 45.33556528 °, W- 087.68496905 °, EABT 017476, 12 July 2010, Randy LaPlant ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Shawano Co., along Cherry Road, N 44.82852 °, W- 088.72285 °, EABT 013439, 13 July 2010, Robert L. Otto ” / Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Shawano Co., along Hickory Road, N 44.80491 °, W- 088.71557 °, EABT 013284, 13 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (2, WIRC); “ WI: Rock Co., along S. Johnson Road, N 42.645037 °, W- 089.13047997 °, EABT 107501, 14 July 2010, Victor Starostka ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Shawano Co., along Woods Road, N 44.84645 °, W- 088.52161 °, EABT 025222, 15 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (2, WIRC); “ WI: Shawano Co., along Old Lakes Road, N 44.81015 °, W- 088.58273 °, EABT 013288, 15 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (2, WIRC); “ WI: Shawano Co., along Industrial Drive, N 44.76722 °, W- 088.57491 °, EABT 012938, 15 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Nejedlo Road, N 45.35224772 °, W- 088.01709023 °, EABT 017291, 15 July 2010, Randy LaPlant ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Shawano Co., along Cheese Factory Road, N 44.84477 °, W- 088.42981 °, EABT 013600, 19 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Pike River Road, N 45.38102841 °, W- 087.89911386 °, EABT 017622, 19 July 2010, Randy LaPlant ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (2, WIRC); “ WI: Green Co., along Mt. Hope Road, N 42.50737103 °, W- 089.39527133 °, EABT 108122, 21 July 2010, Larry Broitzman ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oconto Co., along County Road R, N 44.87707 °, W- 088.45058 °, EABT 013758, 21 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Green Co., along Mt. Hope Road, N 42.50737103 °. W- 089.39527133 °, EABT 108122, 21 July 2010, Larry Broitzman ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, GERP); “ WI: Marinette Co., along Point Lane, N 45.24940663 °, W- 087.891258 °, EABT 016635, 24 July 2010, Randy LaPlant ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oconto Co., along Brush Lane, N 45.00051 °, W- 088.45244 °, EABT 014669, 26 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oconto Co., along Pine Road, N 44.96420 °, W- 088.43286 °, EABT 014360, 26 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oconto Co., along Peshtigo Brook Road, N 45.01746 °, W- 088.36530 °, EABT 014807, 26 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oconto Co., along County Road M, N 45.00508 °, W- 088.29438 °, EABT 014674, 27 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oconto Co., along County Roads G & M, N 45.00425 °, W- 088.24133 °, EABT 014676, 27 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oconto Co., along Rugg Road, N 45.30374 °, W- 088.54252 °, EABT 016959, 27 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oconto Co., along Tillman Road, N 44.98399 °, W- 088.33889 °, EABT 014512, 27 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Green Co., along County Road EE, N 42.7408787 °, W- 089.48130059 °, EABT 106993, 28 July 2010, Larry Broitzman ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Kirby Lake Road, N 45.22294 °, W- 088.11060 °, EABT 016297, 28 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (2, WIRC); “ WI: Marinette Co., along Brandywine Lane, N 45.40898 °, W- 088.31567 °, EABT 017772, 29 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Parkway Road, N 45.30199 °, W- 088.20638 °, EABT 016970, 29 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Bluebird Road, N 45.39176 °, W- 088.32729 °, EABT 017608, 29 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (2, WIRC); “ WI: Marinette Co., along Eagle River Road, N 45.43129 °, W- 088.23354 °, EABT 017910, 29 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Country Road F, N 45.39204 °, W- 088.35157 °, EABT 017607, 29 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Boat Landing no. 13 Road, N 45.34381 °, W- 088.22379 °, EABT 017284, 29 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Ranch Road, N 45.320095 °, W- 088.23276 °, EABT 017104, 29 July 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Vilas Co., July 2010, Marie Eicksen-Pilch ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Rusk Co., July 2010, Marie Eicksen-Pilch ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Grant Co., along County Road VV, N 42.73084 °, W- 091.01829 °, EABT 025487, 2 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (11, WIRC); “ WI: Grant Co., along County Road VV, N 42.74561 °. W- 091.03953 °, EABT 025486, 2 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, GERP); “ WI: Grant Co., along Birchwood Lane, N 42.55105 °, W- 090.61307 °, EABT 108134, 3 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (2, WIRC); “ WI: Grant Co., along Kelly Lane, N 42.65854 °, W- 090.64565 °, EABT 107667, 3 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Grant Co., along Indian Creek Road, N 42.61946 °, W- 090.63913 °, EABT 107901, 3 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Grant Co., Potosi Recreation Area, N 42.66558 °, W- 090.72927 °, EABT 107664, 3 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Grant Co., Grant River Recreation Area, N 42.65148 °, W- 090.70064 °, EABT 107665, 3 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Grant Co., along River Lane, N 42.64566 °, W- 090.69144 °, EABT 000802, 3 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (2, WIRC); “ WI: Grant Co., O’Leary’s Lake Recreation Area, N 42.54082 °, W- 090.63676 °, EABT 000151, 3 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (4, WIRC); “ WI: Grant Co., along Far-Nuff Road, N 42.69210 °, W- 090.90461 °, EABT 107550, 4 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Grant Co., K- 7 Koral Campground, N 42.70224 °, W- 090.95731 °, EABT 10548, 4 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Grant Co., along Closing Dam Road, N 42.76075 °, W- 091.05850 °, EABT 107215, 4 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (3, WIRC); “ WI: Grant Co., along Burton Lane, N 42.71818 °, W- 090.81564 °, EABT 107444, 4 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (2, WIRC); “ WI: Grant Co., along Lynn Hollow Lane, N 42.67606 °, W- 090.81497 °, EABT 107661, 4 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Grant Co., along Dutch Hollow Road, N 42.70730 °, W- 090.78826 °, EABT 107445, 4 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Grant Co., along Airport Road, N 42.77551 °, W- 090.67876 °, EABT 107117, 5 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Grant Co., along County Road O, N 42.68338 °, W- 090.63218 °, EABT 107559, 5 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Shawano Co., along County Road M, N 44.76789 °, W- 088.69987 °, EABT 012934, 9 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oconto Co., along Peshtigo Brook Road, N 45.01746 °, W- 088.36530 °, EABT 014807, 16 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oconto Co., along Brush Lane, N 45.00051 °, W- 088.45244 °, EABT 014669, 16 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along County Road C, N 45.39984 °, W- 088.14449 °, EABT 017614, 24 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along County Road A, N 45.42694 °, W- 088.10370 °, EABT 017914, 24 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Davis Road, N 45.41748 °, W- 088.08846 °, EABT 017779, 24 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Marinette Co., along Smith Creek Road, N 45.26758 °. W- 088.05819 °, EABT 016630, 24 August 2010, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, GERP; 2, WIRC); “ WI: Rusk Co., August 2010, Marie Eicksen-Pilch ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Oneida Co., August 2010, Marie Eicksen-Pilch ” / “ Taken from EAB, prism trap baited, with Manuka oil ” (1, WIRC); “ WI: Sauk Co., along Schiffet Road, N 43.1677610 °, W- 090.078893 °, EABT 104953 C, 11 July 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (2, WIRC); “ WI: Grant Co., along Walnut Road, N 42.9040730 °, W- 091.099555 °, EABT 106539, 14 July 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (3, WIRC); “ WI: Waupaca Co., along County Road T, N 44.42629 °. W- 088.75619 °, EABT 010405 A, 21 July 2011, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil & Z 3 - Hexon- 1 - ol ” (2, GERP); “ WI: Waupaca Co., along Big Eddy Road, N 44.39886 °. W- 088.77316 °, EABT 010267 A, 21 July 2011, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil & Z 3 - Hexon- 1 - ol ” (1, GERP); “ WI: Waupaca Co., along Big Eddy Road, N 44.39883 °. W- 088.77314 °, EABT 010267 B, 21 July 2011, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil & Z 3 - Hexon- 1 - ol ” (3, GERP); “ WI: Richland Co., along State Highway 60, N 43.2070650 °, W- 090.565659 °, EABT 104865, 21 July 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (1, WIRC); “ WI: Richland Co., along Country Lane, N 43.3549120 °, W- 090.413616 °, EABT 104037 C, 28 July 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (3, WIRC); “ WI: Marinette Co., along H & S Lane, N 45.37822 °, W- 088.26212 °, EABT 017450, 29 July 2011, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (1, WIRC); “ WI: Winnebago Co., along Mill Pond Lane, N 44.22178 °. W- 088.47646 °, EABT 100276 C, 15 August 2011, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil & Z 3 - Hexon- 1 - ol ” (1, GERP); “ WI: Grant Co., along Bluff Road, N 42.5388840 °, W- 090.622583 °, EABT 000151, 16 August 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (3, WIRC); “ WI: Grant Co., along Dutch Hollow Road, N 42.7041840 °, W- 090.770282 °, EABT 107446, 18 August 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (3, WIRC); “ WI: Grant Co., along State Highway 133, N 42.7039690 °, W- 090.943685 °, EABT 107549, 18 August 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (1, WIRC); “ WI: Grant Co., along Pigeon River Road, N 42.785030 °, W- 090.809306 °, EABT 107112, 23 August 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (1, WIRC); “ WI: Grant Co., along Mississippi Drive, N 42.8654220 °, W- 091.050628 °, EABT 106656, 24 August 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (5, WIRC); “ WI: Grant Co., along Wightman Road, N 43.1874800 °, W- 090.528244 °, EABT 104978, 6 September 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (4, WIRC); “ WI: Richland Co., along State Highway 60, N 43.2023310 °, W- 090.434931 °, EABT 104869 D, 7 September 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (1, WIRC); “ WI: Iowa Co., along State Highway 14, N 43.1678660 °, W- 090.039619 °, EABT 104994 B, 7 September 2011, Andy Anderson ” / “ Taken from EAB, prism trap baited, with Manuka oil, & Z 3 - Hexon- 1 - ol ” (4, WIRC); “ WI: Florence Co., along Brook Street, N 45.82581 °. W- 088.24797 °, EABT 020739 A, 7 September 2011, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil & Z 3 - Hexon- 1 - ol ” (1, GERP); “ WI: Florence Co., along Brook Street, N 45.82539 °. W- 088.24696 °, EABT 020739 B, 7 September 2011, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil & Z 3 - Hexon- 1 - ol ” (1, GERP); “ WI: Florence Co., along County Road N, N 45.83457 °. W- 088.22651 °, EABT 020740 A, 7 September 2011, Robert L. Otto ” / “ Taken from EAB, prism trap baited, with Manuka oil & Z 3 - Hexon- 1 - ol ” (1, GERP); “ WI: Oconto Co., along Groniger St., N 44.998073 °, W- 088.378475 °, 26 June – 13 July 2012, Robert L. Otto ” / “ Taken from elevated, flight intercept trap, baited with Manuka oil ” (1, GERP); “ USA: WI: Ozaukee Co., UW-Mil. Field Station, 43.38295 ° N / - 88.02534 ° W, WGS 84; 09 – 16 July 2013, Daniel K. Young; Malaise, Trap Fagus - Acer forest ” / “ Nematodes, penetrans, (LeConte), Det. R. L. Otto, 2015 ” (2, WIRC); “ USA: WI: Langlade County, Gardner Dam BSA Camp, 45.13667 ° N / 88.68449 ° W, WGS 84; 13 – 20 July 2013, Kari Gullickson, Duane Hille, Malaise trap; hardwood forest ” (1, WIRC); “ USA: WI: Ozaukee Co., UW-Mil. Field Station, 43.38295 ° N / - 88.02534 ° W, WGS 84; 22 – 30 July 2013, Daniel K. Young; Malaise, Trap Fagus - Acer forest ” / “ Nematodes, penetrans, (LeConte), Det. R. L. Otto, 2015 ” (1, WIRC); “ USA: WI: Ozaukee Co., UW-Mil. Field Station, 43.38295 ° N / - 88.02534 ° W, WGS 84; 13 – 20 August 2013, Daniel K. Young; Malaise, Trap Fagus - Acer forest ” / “ Nematodes, penetrans, (LeConte), Det. R. L. Otto, 2015 ” (1, WIRC); “ USA: WI: Dodge Co., Ledge County Park, ~ 43.47357 ° N 88.57654 ° W, 28 June 2014, Jeffrey P. Gruber ” / “ found on mostly debarked, trunk of dead standing, hardwood tree, mixed, hardwood forest ” / “ Nematodes, penetrans, (LeConte), Det. R. L. Otto, 2016 ” (“ 16 ” handwritten) (1, WIRC); “ USA: WI: Waukesha Co., Muskego Park, Hardwoods SNA, ~ 42.89587 ° N 88.16227 ° W, 5 July 2014, Jeffrey P. Gruber ” / “ crawling on large debarked, hardwood log, southern, dry-mesic forest ” / “ Nematodes, penetrans, (LeConte), Det. R. L. Otto, 2014 ” (7, WIRC); “ WI: Oconto Co., Jeff Otto Property, N 44 ° 59.075 ’, W- 088 ° 28.725 ’, 5 April 2015, Robert L. Otto ” / “ Reared from larvae, in rotten Acer limb, on 8 May 2015 ” (1, GERP); “ WI: Oconto Co., Jeff Otto Property, N 44 ° 59.075 ’, W- 088 ° 28.725 ’, 5 April 2015, Robert L. Otto ” / “ Reared from larvae, in rotten Acer limb, on 18 May 2015 ” (1, GERP); “ WI: La Crosse Co., Coulee Exp. Forest, N 43 ° 51.612 ’, W- 91 ° 01.797 ’, 15 July 2015, Robert L. Otto ” / “ Taken from, downed elm, tree ” (1, GERP); “ USA: WI: Outagamie County, LaSage Bottom SNA, 44.38264 ° N / 88.60455 ° W, 24 June – 09 July 2016, WGS 84; Ann Marsh ” / “ Non-baited Townes Malaise, Trap; second growth flood-, plain forest; shade, between, ephemeral pond and river; Specify Collection, Event Code, 2016 AM 0148 ” (1, WIRC); “ USA: WI: Juneau County, 15 m E of Lemonwear River, 43.91632 ° N / 90.16554 ° W, 27 June – 11 July 2016, WGS 84; Ann Marsh ” / “ Ocotopoda baited Lindgren, Funnel Trap; flood plain, forest; amid ephemeral, ponds and river; mixed, hardwood canopy; shade, Specify Collection, Event Code, 2016 AM 0152 ” (5, WIRC); “ USA: WI: Outagamie County, LaSage Bottom SNA, 44.38264 ° N / 88.60455 ° W, 09 – 25 July 2016, WGS 84; Ann Marsh ” / “ Non-baited Townes Malaise, Trap; second growth flood-, plain forest; shade, between, ephemeral pond and river; Specify Collection, Event Code, 2016 AM 0171 ” (1, WIRC); “ USA: WI: Juneau County, 15 m E of Lemonwear River, 43.91632 ° N / 90.16554 ° W, 11 – 28 July 2016, WGS 84; Ann Marsh ” / “ Ocotopoda baited Lindgren, Funnel Trap; flood plain, forest; amid ephemeral, ponds and river; mixed, hardwood canopy; shade, Specify Collection, Event Code, 2016 AM 0181 ” (1, WIRC).	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF4FFE44FF708DBDFAACF87B.taxon	description	Redescription. Length, 3.0 – 6.0 mm. Width, 1.0 mm. Body subcylindrical, elongate; uniformly black; antennae dark brown; legs including tarsi dark brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (Fig. 15). Head: Subspherical; integument closely punctate, somewhat shiny; frons convex, with median, round, shallow fovea above frontoclypeal region; apical margin of frontoclypeal region weakly trilobed, about 2 times wider than base; mandibles stout, bidentate, densely punctate. Antenna: Filiform to weakly serriform from antennomeres III – XI, attaining nearly 1 / 3 the length of the body; antennomere III longer than IV; antennomeres IV – VI each sub-equal, slightly longer than wide, each slightly shorter than VII; antennomeres VII – X each sub-equal, longer than wide; antennomere XI slightly longer than X. Pronotum: Integument somewhat shiny, densely punctate to rugose; longer than wide, with moderate, sharp hind angles; lateral sides basally constricted above pronotal hind angles, parallel-sided, cranially arcuate; disc convex with short, shallow median groove and pair of shallow, circular fovea; base sinuous. Scutellar shield: Quadrate, setose, shallowly punctate and distally truncated. Elytra: Striae absent to weakly indicated; interstices flattened; integument somewhat shiny, transversely rugose at basal 1 / 3, closely punctate at apical 2 / 3. Legs: First tarsomere shorter than the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I – III simple; metatarsomere IV excavate-emarginate; metatarsomere V elongate with simple claws. Venter (Fig. 16): Closely punctate, with short, recumbent yellowish setae; hypomeron with basally obliterated lateral antennal grooves; strongly-developed femoral ridge incomplete, elongate, extending almost to the lateral side of the pronotum; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially more than 3.0 – 6.0 times wider than laterally; last abdominal ventrite apically produced. Aedeagus (Fig. 17): Basal piece longer than wide, laterally parallel-sided, dorsally open, apically rounded; remaining parts elongate, parallel-sided; parameres elongate, apically pointed, triangular-shaped lateral tooth present half way between base of and apices of the parameres; secondary lateral lobes present, short, attached at base of parameres; median lobe elongate and narrow, apically pointed, deeply and narrowly bifid, as long as the parameres.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF4FFE44FF708DBDFAACF87B.taxon	distribution	Distribution. In Canada, this somewhat commonly collected, widespread eucnemid species has been taken in New Brunswick, Nova Scotia, Ontario, and Québec (Webster et al. 2012). In the United States, N. penetrans is known from Alabama, Arkansas (New State Record), Florida, Georgia, Illinois, Indiana, Iowa, Kansas (New State Record), Kentucky, Louisiana, Maine, Maryland, Michigan, Minnesota, Missouri, New Hampshire, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Texas, Vermont, Virginia, West Virginia, and Wisconsin (Otto and Karns 2017).	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF4FFE44FF708DBDFAACF87B.taxon	biology_ecology	Biology. Specimens were taken from a blacklight trap, white PVC trap, a Malaise trap, Lindgren funnel traps, a panel intercept trap, and cross-vane panel traps. Immature stages were documented by Otto (2017).	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF54FE40FF708FA3FDCBFDAD.taxon	description	Fig. 18 – 21	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF54FE40FF708FA3FDCBFDAD.taxon	diagnosis	Diagnosis. A well-developed, elongate, incomplete femoral ridge will readily distinguish N. rugosipennis new species from most Nematodes species in the region, except N. penetrans. A shorter antennomere V in relation to antennomere IV will further distinguish the new species from N. penetrans.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF54FE40FF708FA3FDCBFDAD.taxon	materials_examined	Type material. Female holotype: “ FLORIDA: ALACHUA COUNTY, GAINESVILLE, BEVILLE HTS, 19 - III- 80, L. A. STANGE, BLACKLIGHT TRAP ” / “ HOLOTYPE:, Nematodes, rugosipennis ♀, Otto, det. R. L. Otto, 2019 ” (red printed label). Male allotype: “ Florida: Marion Co., ~ 4 miles E Ft. McCoy, April 26 – June 15, 2015, Lindgren funnel trap, Kyle E. Schnepp ” / “ ALLOTYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2019 ” (yellow printed label). Holotype is deposited in FSCA. Allotype is deposited in KESC. Paratypes. 17 ♂♂, 22 ♀♀: FLORIDA: 2 ♂♂ 1 ♀, “ USA FLORIDA Columbia Co., O’Leno State Park, 01 - V- 96, Coll. Robert Vigneault ” / “ Collection of the Global, Eucnemid Research Project, (Robert L. Otto) ” (green framed white label) / “ PARATYPE:, Nematodes, rugosipennis ♂ (or ♀), Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (3, GERP); 1 ♂, “ USA FLORIDA Orange Co., Wekiwa Springs SP, 12 - V- 96, Coll. Robert Vigneault ” / “ Collection of the Global, Eucnemid Research Project, (Robert L. Otto) ” (green framed white label) / “ PARA- TYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (1, GERP); 1 ♀, “ FL: Polk Co., Green Swamp WMA, May 2001, Morris and Nigg, Malaise Trap ” / “ Collection of the Global, Eucnemid Research Project, (Robert L. Otto) ” (green framed white label) / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (1, GERP); 2 ♀♀, “ USA: FL: Wakulla Co.; Apalachicola; Nat. For.; FS 366; Pine / oak savanna; 15 – 20 - MAY- 2005; 30 ° 19.751 ’ N; 84 ° 30.309 ’ W; Deans & Murray; EX: “ ant heaven ” malaise trap ” / PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2020 ” (yellow printed label) (2, CDAE); 1 ♀, “ USA: FL: Leon Co.; Apalachicola; NF; FS 390; 30 ° 21.884 ’ N; 84 ° 40.774 ’ W; Mosquito Hell; Malaise trap; bottomlands; 21 – 28. V. 2005; S. Joshi ” / PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2020 ” (yellow printed label) (1, CDAE); 1 ♂, “ FLORIDA: Alachua, Co., Oleno State Park, 21 - IV- 2009, L. A. Wilby, EAB trap w / Manula oil ” / “ PARATYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (1, FSCA); 2 ♀♀, “ FLORIDA: Wakulla Co., Ochlockowee Riv. St. Pk., 11 - V- 2009, L. Wilby, EAB trap w / manuka oil ” / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (2, FSCA); 1 ♀, “ FLORIDA: Alachua Co., O’Leno St. Pk., 27 - V- 2009, L. Wilby, EAB trap w / manuka oil ” / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (1, FSCA); 2 ♂♂, “ FLORIDA: Highlands Co., Highlands Hammock St. Pk., 27.47070, - 81.55.254, 5 - IV- 2011, C. Carrero-Turnbull, D. Gaskilla, lindgren funnel, manuka oil ” / ““ PARATYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (2, FSCA); 2 ♂♂, “ Florida: Alachua Co., San Felasco Preserve, April 5 – May 6, 2014, Lindgren funnel trap, Kyle E. Schnepp ” / “ PARATYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2021 ” (yellow printed label) (2, KESC); 3 ♀♀, “ Florida: Alachua Co., 3.4 miles WNW Archer, March 30 – June 12, 2015, Lindgren funnel trap, Kyle E. Schnepp ” / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (3, KESC); 1 ♀, “ Florida: Marion Co., ~ 4 miles E Ft. McCoy, March 21 – April 26, 2015, Lindgren funnel trap, Kyle E. Schnepp ” / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (1, KESC); 2 ♀, “ Florida: Marion Co., ~ 4 miles E Ft. McCoy, April 26 – June 15, 2015, Lindgren funnel trap, Kyle E. Schnepp ” / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (2, KESC); 1 ♀, “ Florida: Suwanee Co., ~ 7 miles W White Springs, March 29 – April 19, 2017, Lindgren funnel trap, Kyle E. Schnepp ” / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2021 ” (yellow printed label) (1, KESC); 1 ♀, “ Florida: Suwanee Co., ~ 7 miles W White Springs, April 19 – May 16, 2017, Lindgren funnel trap, Kyle E. Schnepp ” / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2021 ” (yellow printed label) (1, KESC); 1 ♂, “ FL: Polk co., 2635 E Ewell Rd., Lakeland, 16 – 26 / II / 2018, R. Morris, WPT ” / “ Collection of the Global, Eucnemid Research Project, (Robert L. Otto) ” (green framed white label) / “ PARATYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (1, GERP); 3 ♂♂, “ FL: Polk co., 2635 E Ewell Rd., Lakeland, 18 – 29 / III / 2018, R. Morris, WPT ” / “ Collection of the Global, Eucnemid Research Project, (Robert L. Otto) ” (green framed white label) / “ PARATYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (3, GERP); 1 ♂, “ FL: Polk co., 2635 E Ewell Rd., Lakeland, V / 1 – 13, / 2018, R. Morris, Window pane Trap ” / “ Collection of the Global, Eucnemid Research Project, (Robert L. Otto) ” (green framed white label) / “ PARATYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (1, GERP); 1 ♂, “ Florida: Liberty Co., Torreya S. P. at light, May 11, 2018, Kyle E. Schnepp ” / “ PARATYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2021 ” (yellow printed label) (1, KESC); 1 ♂, “ Florida: Alachua Co., San Felasco H. P. S. P., April 24 – May 5, 2021, Lindgren funnel trap, Kyle E. Schnepp ” / “ PARATYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2021 ” (yellow printed label) (1, KESC); 3 ♀♀, “ Florida: Alachua Co., San Felasco H. P. S. P., April 24 – May 5, 2021, Lindgren funnel trap, Kyle E. Schnepp ” / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2021 ” (yellow printed label) (3, KESC). GEORGIA: 1 ♀, “ GEORGIA: Bibb County, 2.6 km SSW Walden, 32.68599, - 83.67626, 108 m, 29 Mar – 13 Apr 2018, LFT, C. Barnes, BSF # 78312 ” / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2020 ” (yellow printed label) (1, CMNH); 1 ♀, “ GEORGIA: Cook County, 4.7 km E of Ellenton, 76 m, 31.17202, - 83.53825, 25 Apr – 15 May 2018, LFT, N. Skaggs, BSF # 85831 ” / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2020 ” (yellow printed label) (1, CMNH). OKLA- HOMA: 1 ♀, “ OKLAHOMA Grady Co., Tuttle, VI / 10 / 05, Lindgren traps, Brian Baldwin ” / “ Collection of the Global, Eucnemid Research Project, (Robert L. Otto) ” (green framed white label) / “ PARATYPE:, Nematodes, rugosipennis ♀, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (1, GERP); 2 ♂♂, “ OKLAHOMA Grady Co., Tuttle, VI / 22 / 05, Blacklight traps, Brian Baldwin ” / “ Collection of the Global, Eucnemid Research Project, (Robert L. Otto) ” (green framed white label) / “ PARATYPE:, Nematodes, rugosipennis ♂, Otto, Det. R. L. Otto, 2019 ” (yellow printed label) (2, GERP). Paratypes are deposited in CDAE, CMNH, FSCA, GERP, and KESC.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF54FE40FF708FA3FDCBFDAD.taxon	description	Description. Female holotype: Length, 10.0 mm. Width, 2.5 mm. Body subcylindrical, elongate; uniformly pitch black; antennae dark brown; legs including tarsi dark brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (Fig. 18). Head: Subspherical; integument confluently rugose, somewhat dullish; frons convex, with median, round, shallow fovea above frontoclypeal region; apical margin of frontoclypeal region weakly trilobed, about 2 times wider than base; mandibles stout, bidentate, densely punctate. Antenna: Filiform to weakly serriform from antennomeres III – XI, attaining nearly 1 / 3 the length of the body; antennomere III longer than IV; antennomeres IV – V each sub-equal, longer than wide, each slightly shorter than VI; antennomeres VI – IX each sub-equal, longer than wide; antennomere X slightly shorter than IX; antennomere XI stout, slightly longer than X, less than 2.0 times longer than wide. Pronotum: Integument dullish, densely granulose to rugose; longer than wide, with moderate, sharp hind angles; lateral sides basally constricted above pronotal hind angles, parallel-sided, cranially arcuate; disc convex with deep median groove and 2 pairs of deep, circular fovea; base sinuous. Scutellar shield: Elongate, sub-triangular, setose, shallowly punctate and distally rounded. Elytra: Distinctly striate; interstices elevated; integument somewhat shiny, transversely rugose. Legs: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I – III simple; metatarsomere IV excavate-emarginate; metatarsomere V elongate with simple claws. Venter (Fig. 19): Very closely punctate, with short, recumbent yellowish setae; hypomeron with basally obliterated lateral antennal grooves; well-developed femoral ridge incomplete, elongate, extending near the lateral side of the pronotum; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially more than 3.0 – 6.0 times wider than laterally; last abdominal ventrite apically produced. Male allotype (Fig. 20). Length, 9.0 mm. Width, 2.0 mm. Antennae filiform to weakly serriform, reaching at least 1 / 3 the length of the body; habitus similar to holotype; legs and antennae dark brown; antennomere X is as long as antennomere IX; pronotum similar to holotype, except second pair of circular fovea posterolaterad of the first pair are absent; integument similar to holotype. Aedeagus (paratype) (Fig. 21). Basal piece longer than wide, laterally parallel-sided, dorsally open, apically rounded; remaining parts elongate, constricted laterally just above the basal piece, laterally sinuous; parameres elongate, apically rounded, triangular-shaped lateral tooth present near apices; median lobe elongate and narrow, apically pointed, deeply and narrowly bifid, longer than parameres. Variation. Seventeen male and 22 female paratypes were examined. Male paratypes measured 7.0 – 10.0 mm long and 1.5 – 2.0 mm wide. Female paratypes measured 6.0 – 11.5 mm long and 1.5 – 2.5 mm wide. Fifteen of the 22 female paratypes are shorter and narrower than the holotype. The three female paratypes are larger and wider than the holotype. The four remaining female paratypes are just as long as but slightly narrower than the holotype. All male paratypes are smaller and narrower than the holotype. Four of the 17 male paratypes is as long as and as wide as the allotype. Five male paratypes are longer than the allotype. Eight male paratypes are shorter than the allotype. Circular fovea on the pronotal disc are much deeper in seven of the 17 male paratypes and eight of the 22 female paratypes compared to the other paratypes, holotype, and allotype. All paratypes lack the second pair of circular fovea on the pronotum. Two male paratypes are brownish in color, possibly teneral at the time they were collected. There are no other exoskeletal difference between any paratypes, allotype, and the holotype.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF54FE40FF708FA3FDCBFDAD.taxon	distribution	Distribution. This eucnemid species is known from 41 specimens taken from a number of locations in Florida, two locales in Georgia and a single location in Oklahoma.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF54FE40FF708FA3FDCBFDAD.taxon	biology_ecology	Biology. The holotype was taken from a blacklight trap placed in Gainesville, Florida. Eleven female paratypes, three male paratypes and male allotype were taken from Lindgren funnel traps deployed in Alachua, Marion, and Suwanee Counties, Florida. Five male paratypes were taken from a window pane trap deployed in Polk County, Florida. Two female paratypes were taken from “ ant heaven ” Malaise trap placed within a pine / oak savanna in Wakulla County, Florida. One female paratype was taken from a Malaise trap placed in bottomlands within Leon County, Florida. One female paratype was taken from a Malaise trap placed in Polk County, Florida. One male and three female paratypes were taken from Emerald Ash Borer trap baited with Manuka oil in two Florida Counties. Two male paratypes were taken from Lindgren funnel trap baited with Phoebe oil in Highlands County, Florida. One male paratype was taken from light in Liberty County, Florida. Two female specimens were taken from Lindgren funnel traps placed in two locations in Georgia. Two male paratypes were taken from blacklight in Oklahoma. One female paratype was taken from Lindgren funnel traps in Oklahoma. Immature stages remain unknown.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
0F7C87E7EF54FE40FF708FA3FDCBFDAD.taxon	etymology	Etymology. The specific epithet is derived from its transversely rugose surfaces present throughout the elytral interstices. Note. Basal sex combs present in male Nematodes species are often difficult to observe when examining specimens under magnification. Protarsomere I was observed to be shorter in male specimens compared to the length of protarsomere I in female specimens.	en	Otto, Robert L. (2021): A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species. Insecta Mundi 2021 (881): 1-19, DOI: 10.5281/zenodo.5352354
