taxonID	type	description	language	source
EA1F9FD0742C4FA4E7032F914DF69E61.taxon	description	Re-description (based on the specimens from Lushui County, Yunnan). Size: Body length 6.9 - 8.0 mm, width 3.5 - 3.9 mm. Color: Body black, slightly shiny on dorsum, labrum and bases of mandibles paler, reddish brown in some specimens. Antennae brown, with antennomere 1 more or less infuscated. Palpi yellowish brown. Legs dark brown to black, with tibiae basally and tarsi brown. Microsculpture: Head with dorsal microsculpture visible throughout, comprised of very fine, more or less isodiametric meshes. Pronotum with microsculpture clearly visible throughout, comprised of distinct isodiametric meshes. Elytra with microsculpture developed throughout, comprised of distinct, slightly transverse meshes. Head: Comparatively large (HWmax / PWmax = 0.69 - 0.72 and HWmin / PWmax = 0.59 - 0.61), impunctate, with small and moderately convex eyes; tempora flat, sloped to neck. Clypeus slightly concave and with slightly convex bead along anterior margin. Frontal suture distinct, slightly impressed. Frontal foveae small, faintly deepened; clypeo-ocular prolongations absent. Supraorbital setae situated slightly behind hind margins of eyes .. Antennae short, not extended to pronotal basal margin, with antennomeres 5 to 7 each 1.4 - 1.6 times as long as wide. Labrum deeply emarginate apically. Left mandible truncate at apex. Ligular sclerite more or less parallel-sided, with apex straight and apical angles slightly extended laterad. Pronotum (Fig. 1): Markedly transverse (PWmax / PL = 1.55 - 1.63), narrowed basad (PWmax / PWmin = 1.22 - 1.30) and widest in anterior third. Sides distinctly rounded in anterior two-thirds and very slightly rounded, almost rectilinearly convergent basad in posterior third, with one lateral seta on each side in apical third. Apical margin arcuately emarginate, bordered only laterally. Basal margin nearly straight, slightly oblique at basal angles, distinctly bordered throughout, slightly wider than apical margin and slightly shorter than elytral base between humeral angles. Apical angles slightly less than 90 °, produced anteriad and narrowly rounded at apices. Basal angles obtuse, sharp at apices, each with a small, obtuse denticle extended laterad. Pronotal disc moderately convex, slightly depressed basally, slightly sloped to sides and more abruptly sloped to apical angles. Lateral depressions present as narrow furrows behind apical angles, slightly widened behind lateral setigerous pore, and either extended basad to pronotal base or fused in basal quarter with large latero-basal depressions. Basal foveae distinct within latero-basal depressions, comparatively small, somewhat flat or slightly deepened. Areas between basal foveae and between basal foveae and lateral depressions slightly convex in most specimens. Pronotal surface coarsely and irregularly punctate along base and in lateral depressions, finer and sparser punctation present also along apical margin. Elytra: Oval, markedly rounded at sides, wide (EL / EW = 1.20 - 1.29, EL / PL = 2.58 - 2.69, EW / PWmax = 1.32 - 1.37), widest approximately at middle, not fused along suture. Humeri widely rounded at apices, each with indistinct denticle hardly visible from behind; sides just behind humeral angles markedly rounded. Subapical sinuations present, shallow. Sutural angles not separated from each other, less than 90 °, with apices blunt in both sexes. Basal borders faintly sinuate, joined with lateral margin at a markedly obtuse angle. Striae smooth, impunctate, faintly impressed along entire length. Parascutellar striae present, short, basal setigerous pores present; in some specimens, distal part of parascutellar stria connected with first (sutural) stria. Intervals faintly convex along entire length, impunctate, only slightly narrowed toward apices. Umbilicate setal series widely interrupted at middle, with anterior group comprised of 7 or 8 setigerous pores and posterior group comprised of 8 or 9 such pores. Hindwings: Reduced to small scales. Venter: Prosternum with scattered and short setae apically. Prosternal medial process not projected posteriad. Proepisterna smooth, impunctate. Metepisterna (Fig. 3) distinctly narrowed posteriad, approximately as wide as long. Sternum VII (anal) in both sexes with two pairs of setae along apical margin and widely rounded at apex. Tergum VII (anal) of female rounded at apex. Legs: Metacoxae (Fig. 2) with additional postero-medial setigerous pore and, in most specimens, with additional medial setigerous pore at least on one side. Metafemora with two setae along posterior margin. Protibiae with one ventroapical spine, the outer margin in both sexes with three or four uniform preapical spines. Tarsi glabrous dorsally, at most with a few setae on enlarged male protarsomeres; tarsomere 5 with three pairs of lateroventral setae. Metatarsi in both sexes notably shorter than minimum linear distance across neck constriction just behind eyes; tarsomere 1 longer than tarsomere 2 and shorter than tarsomeres 2 + 3. In males, protarsi moderately enlarged, with tarsomere 1 about as long as wide, tarsomeres 2 to 4 distinctly wider than long and tarsomeres 1 to 4 with adhesive vestiture ventrally (only apically on tarsomere 1); mesotarsi comparatively slightly enlarged, with tarsomere 1 slightly longer than wide, tarsomere 2 approximately as long as wide, tarsomere 3 slightly wider than long, tarsomere 4 much smaller than tarsomeres 2 and 3 and deeply concave apically; mesotarsomeres 2 and 3 with adhesive vestiture ventrally. Female genitalia (Figs 4 - 5): Apical stylomere comparatively slightly curved. Aedeagus (Figs 6 - 9): Median lobe markedly widened medially (dorsal aspect), markedly bent ventrad behind basal bulb and with terminal lamella slightly curved dorsad (lateral aspect), its ventral margin nearly straight medially. Terminal lamella (Fig. 6) flat, triangular in dorsal aspect, slightly wider than long, narrowly rounded at apex and without any apical capitulum. Apical orifice in dorsal position, prolonged to basal bulb. Internal sac with several spiny patches.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
EA1F9FD0742C4FA4E7032F914DF69E61.taxon	distribution	Distribution. Fig. 56. This species is known only from the Gaoligong Shan in Lushui and Tengchong counties, western Yunnan Province, China, at elevations of 2500 - 3200 m.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
C965766D3E1905C307D5E4FB31512D42.taxon	description	Description (male holotype only). Dorsal habitus as in Fig. 51. Size: Body length 8.2 mm, width 3.8 mm. Color: Body black, shiny on dorsum; mandibles basally, pronotum at basal angles and elytral epipleura slightly paler, reddish black. Antennae brown, with antennomeres 3 and 4 infuscated. Palpi light brown. Tarsi dark brown. Microsculpture: Head with dorsal microsculpture comprised of fine isodiametric meshes, visible throughout, except effaced on frons and vertex. Pronotum with microsculpture visible throughout, comprised of fine isodiametric meshes basally and laterally and of slightly transverse, nearly effaced meshes on remaining surface. Elytra with microsculpture distinct throughout, comprised of fine, slightly transverse meshes. Head: Comparatively large (HWmax / PWmax = 0.73 and HWmin / PWmax = 0.63), covered with micropunctures and very fine wrinkles; tempora slightly convex, almost flat, sloped to neck. Clypeus slightly concave and with slightly convex bead apically. Frontal suture distinct, slightly impressed. Frontal foveae small, oval, slightly deepened; clypeo-ocular prolongations absent. Supraorbital setae situated just behind hind margins of eyes. Eyes small and moderately convex. Antennae slightly extended beyond pronotal basal margin, with antennomeres 5 to 7 each about 1.6 times as long as wide. Labrum distinctly emarginate apically. Left mandible truncate at apex. Ligular sclerite nearly parallel-sided, straight at apex. Pronotum (Fig. 10): Markedly transverse (PWmax / PL = 1.64), narrowed basad (PWmax / PWmin = 1.26) and widest before middle. Sides rounded in anterior half and very faintly sinuate in basal half, with one lateral setigerous pore on each side before middle. Apical margin arcuately emarginate, bordered only laterally. Basal margin very broadly emarginate, almost straight, distinctly bordered throughout, slightly wider than apical margin and slightly narrower than elytral base between humeral angles. Apical angles slightly less than 90 °, extended anteriad and narrowly rounded at apices. Basal angles slightly more than 90 °, sharp at apices, each with a tiny denticle produced laterad. Pronotal disc moderately convex, slightly depressed basally, slightly sloped to sides and more abruptly sloped to apical angles. Lateral depressions present as narrow furrows behind apical angles, slightly widened behind lateral setigerous pore, and fused in basal quarter with large laterobasal depressions. Basal foveae elongate, moderately deep. Areas between basal foveae and between basal foveae and lateral depressions slightly convex. Pronotal surface coarsely and irregularly punctate along base and in lateral depressions, finer and sparser punctation present along apical margin and in central portion of disc. Elytra: Oval, rounded at sides, moderately wide (EL / EW = 1.34, EL / PL = 2.83, EW / PWmax = 1.29), widest approximately at middle, not fused along suture. Humeri slightly prominent, widely rounded at apices, each with indistinct denticle visible only from behind; sides just behind humeral angles markedly rounded. Subapical sinuations shallow. Sutural angles not separated from each other, less than 90 °, blunt at apices. Basal borders slightly sinuate, joined with lateral margin at very obtuse angle. Striae impunctate, faintly impressed along entire length. Parascutellar striae present, short, with basal setigerous pores present. Intervals slightly convex, almost flat, impunctate. Umbilicate setal series widely interrupted at middle, with anterior group comprised of six setigerous pores and posterior group comprised of seven setigerous pores on right side and of nine setigerous pores on left side. Hindwings: Reduced to small scales. Venter: Prosternum covered with scattered and very short setae. Prosternal medial process not projected posteriad. Proepisterna smooth, impunctate. Metepisterna (Fig. 11) distinctly narrowed posteriad, slightly wider than long. Sternum VII (anal) in male with two pairs of setae along apical margin and slightly truncate at apex. Legs: Metacoxae with additional posteromedial setigerous pore and with one or two additional medial setigerous pores. Metafemora with two setae along posterior margin. Protibiae with one ventroapical spine, the outer margin with one stouter spine and three slenderer spines apically. Tarsi glabrous dorsally, tarsomere 5 with three or four pairs of lateroventral setae. Metatarsi approximately equal to minimum linear distance across neck constriction just behind eyes; tarsomere 1 distinctly longer than tarsomere 2 and slightly shorter than tarsomeres 2 + 3. Protarsi (in male) moderately enlarged, with tarsomere 1 slightly longer than wide, tarsomeres 2 slightly wider than long, tarsomeres 3 and 4 distinctly wider than long and tarsomeres 1 to 4 with adhesive vestiture ventrally (only apically on tarsomere 1); mesotarsi comparatively slightly enlarged, with tarsomere 1 (not enlarged) distinctly longer than wide, tarsomere 2 slightly longer than wide, tarsomere 3 slightly wider than long, tarsomere 4 much smaller than tarsomeres 2 and 3 and deeply concave apically; mesotarsomeres 2 and 3 with adhesive vestiture ventrally. Aedeagus (Figs 12 - 15): Median lobe moderately widened medially (dorsal aspect), bent ventrad behind basal bulb and with terminal lamella slightly curved dorsad (lateral aspect) and with ventral margin nearly straight medially. Terminal lamella (Fig. 12) flat, in dorsal aspect triangular, slightly longer than wide, narrowly rounded at apex and without any apical capitulum. Apical orifice in dorsal position, prolonged to basal bulb. Internal sac with several spiny patches.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
C965766D3E1905C307D5E4FB31512D42.taxon	distribution	Distribution. Fig. 56. This new species is known only from the type locality (Shibali, Lishadi town, Fugong County) in the northern part of Gaoligong Shan, northwestern Yunnan Province, China.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
56F8823D68B08366D7ADB050DBFCC59F.taxon	description	Description (male). Dorsal habitus as in Fig. 52. Size: Body length 9.3 mm, width 4.0 mm. Color: Body black, slightly shiny on dorsum, almost matte due to fine micropunctation and distinct, granulate microsculpture; two penultimate abdominal sterna V and VI along posterior margin and sternum VII (anal) in apical portion clearly paler, light brown. Antennae, palpi, and legs light brown, femora slightly infuscated. Microsculpture: Head with dorsal microsculpture distinct throughout, comprised of very distinct, isodiametric, almost granulate meshes. Pronotum with microsculpture distinct throughout, comprised of distinct isodiametric and slightly transverse meshes. Elytra with microsculpture distinct throughout, comprised of isodiametric meshes, granulate on three lateral intervals. Head: Comparatively large (HWmax / PWmax = 0.70 and HWmin / PWmax = 0.59), covered with micropunctures and very fine wrinkles on dorsum; tempora short, slightly convex. Clypeus slightly depressed laterally and apically, with apical margin slightly concave and unbordered. Frontal suture superficial. Frontal foveae moderately wide and shallow, clypeo-ocular prolongations short and faintly impressed. Supraorbital setae situated at level of hind margins of eyes. Eyes slightly convex. Antennae short [apical antennomeres missing], with antennomeres 5 to 7 each about 1.8 - 2.0 times as long as wide. Labrum distinctly emarginate apically. Left mandible blunt at apex. Apex of ligular sclerite concave, with apical angles slightly projected laterally. Pronotum (Fig. 16): Relatively long (PWmax / PL = 1.36), distinctly narrowed basad (PWmax / PWmin = 1.27) and widest in anterior third. Sides gently rounded anteriorly, moderately sinuate in basal half and slightly divergent just before base; each side with one lateral setigerous pore in anterior third. Apical margin almost straight, bordered only near apical angles. Basal margin slightly emarginate medially, slightly oblique laterally, faintly bordered throughout, very slightly wider than apical margin and slightly narrower than elytral base between humeral angles. Apical angles acute, distinctly but not markedly produced anteriad. Basal angles almost 90 °, with sharp apices, not denticulate. Pronotal disc moderately convex, not depressed basally, moderately sloped to sides and abruptly sloped to apical angles. Lateral depressions narrow anteriorly, slightly widened behind middle and indistinct at basal angles. Basal foveae wide and very shallow. Area at basal angles flat. Pronotal surface throughout densely micropunctate and covered with very fine wrinkles; laterobasal areas slightly more coarsely punctate. Elytra: Oval, gradually rounded at sides, moderately wide (EL / EW = 1.40, EL / PL = 2.38, EW / PWmax = 1.25), widest at middle and fused along suture. Humeri rounded, without denticles at apices. Subapical sinuations very shallow. Sutural angles acute, sharp at apices, not separated from each other. Basal borders markedly sinuate, joined with lateral margin at an acute angle. Striae superficial, very fine, impunctate, slightly crenulate; inner striae nearly effaced at apex. Parascutellar striae rudimentary, each with a basal setigerous pore. Intervals absolutely flat, impunctate. Umbilicate setal series without distinct gap medially. Hindwings: Reduced to small scales. Venter: Prosternum finely punctate and pubescent. Medial prosternal process prominent and projected posteriad (Fig. 18). Proepisterna smooth. Metepisterna (Fig. 17) distinctly wider than long and slightly narrowed posteriad. Sternum VII (anal) with two pairs of setae along apical margin, rounded at apex. Legs: Metacoxae (Fig. 19) with posteromedial setigerous pore and without additional, setigerous or non-setigerous foveae medially. Metafemora with two setae along posterior margin. Protibiae with one preapical spine at outer distal margin and one ventroapical spine. Tarsi distinctly setose dorsally, tarsomere 5 with three pairs of lateroventral setae. Metatarsi slender, slightly longer than maximum width of head, with tarsomere 1 much longer than tarsomere 2, but distinctly shorter than tarsomeres 2 + 3. Male protarsi markedly enlarged (tarsomeres 2 and 3 much wider than long and tarsomeres 1 to 4 with adhesive vestiture ventrally); mesotarsi moderately enlarged (tarsomere 1 narrow, distinctly longer than wide and with only a pair of adhesive scales apically; tarsomere 2 about 1.2 times as long as wide, with adhesive vestiture ventrally; tarsomere 3 approximately as long as wide, with adhesive vestiture ventrally; and tarsomere 4 small, deeply concave apically and without adhesive vestiture ventrally). Aedeagus (Figs 20 - 22): Median lobe symmetrical, comparatively slender, markedly bent ventrad just behind basal bulb and with straight ventral margin. Sides rounded in middle portion and convergent in apical portion. Terminal lamella (Fig. 20) about 1.5 times as long as wide, triangular, narrowly rounded at apex, without apical capitulum. Apical orifice in dorsal position, prolonged to basal bulb. Internal sac without distinct sclerotic elements.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
56F8823D68B08366D7ADB050DBFCC59F.taxon	distribution	Distribution. Fig. 56. This new species is known only from the type locality (Gutang Township) in the eastern Himalaya, the southeastern part of Xizang Autonomous region (Tibet), Medog County, China.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
DFC67FB53E140134742F8529292FD27B.taxon	description	Description (male). Dorsal habitus as in Fig. 53. Size: Body length 8.7 mm, width 3.8 mm. Color: Body dark piceous; antennae brown, palpi slightly lighter brown; femora black, tibiae dark brown, tarsi brown. Microsculpture: Head with dorsal microsculpture very fine, distinct throughout, comprised of more or less isodiametric meshes. Pronotum with microsculpture comprised of distinct isodiametric and slighty transverse meshes, except effaced on disc. Elytra with microsculpture very fine, comprised of slightly transverse meshes on disc and of isodiametric meshes on two lateral intervals. Head: Comparatively large (HWmax / PWmax = 0.73 and HWmin / PWmax = 0.62), very finely micropunctate on dorsum and covered with fine wrinkles mainly in and around frontal foveae; tempora moderately long, slightly convex. Apex of clypeus concave, very faintly bordered. Frontal suture thin, faintly impressed, clypeo-ocular prolongations thin, not deepened, extended to supraorbital furrows. Supraorbital seta situated at level of hind margin of eye. Eyes moderately convex. Antennae short, not extended to pronotal basal margin, with antennomeres 5 to 7 each about 1.4 - 1.5 times as long as wide. Labrum distinctly emarginate apically. Left mandible truncate at apex. Ligular sclerite slightly widened anteriorly, with straight apex. Pronotum (Fig. 23): Slightly transverse (PWmax / PL = 1.45), distinctly narrowed basad (PWmax / PWmin = 1.24) and widest in anterior third. Sides rounded in anterior two-thirds and slightly sinuate before base; each side with one lateral setigerous pore in anterior third. Apical margin slightly emarginate, bordered only laterally. Basal margin almost straight, distinctly bordered throughout, approximately equal in width to apical margin and slightly narrower than elytral base between humeral angles. Apical angles slightly protruded anteriad, acute and very narrowly rounded at apices. Basal angles obtuse, with sharp, slightly denticulate apices. Pronotal disc moderately convex, not depressed basally, markedly sloped toward apical angles in apical half and slightly sloped toward sides in basal half. Lateral depressions very narrow from apical angles to pronotal base. Basal foveae oval and shallow, isolated from basal border. Areas at basal angles and between basal foveae slightly convex. Pronotal surface punctate almost throughout, very finely punctate medially and along apical margin and more densely and coarsely punctate basally and along sides, punctures confluent and particularly coarse within basal foveae. Elytra: Oval, markedly rounded at sides, comparatively short and wide (EL / EW = 1.34, EL / PL = 2.38, EW / PWmax = 1.22), widest at middle, not fused along suture. Humeri not prominent, with a tiny acute denticle at apices (visible only from behind). Subapical sinuations distinct, moderately deep. Sutural angle slightly less than 90 °, slightly blunted at apex. Basal borders slightly sinuate to humeri, joined with lateral margin at an obtuse angle. Striae impunctate, thin, superficial along their entire length. Parascutellar strioles present but short, basal setigerous pore present. Intervals flat up to apices, impunctate. Umbilicate setal series divided into two groups (humeral and apical), with an isolated setigerous pore medially. Hindwings: Reduced to small scales. Venter: Prosternum glabrous. Medial prosternal process slightly projected posteriad (Fig. 25). Proepisterna smooth. Metepisterna (Fig. 24) approximately as long as wide, markedly narrowed posteriad. Sternum VII (anal) with two pairs of setae along apical margin, widely rounded at apex. Legs: Metacoxae (Fig. 26) without posteromedial setigerous pore or any additional setigerous or nonsetigerous foveae medially. Metafemora with two setae along posterior margin. Protibiae with 1 stouter and 1 - 2 slenderer spines at outer distal margin and with one ventroapical spine. Tarsi glabrous dorsally, tarsomere 5 with three or four pairs of lateroventral setae. Metatarsi short, shorter than width of head just behind eyes, with tarsomere 1 about 1.4 times as long as tarsomere 2 and notably shorter than tarsomeres 2 + 3. Male protarsi markedly enlarged (tarsomeres 2 and 3 much wider than long and tarsomeres 1 - 4 with adhesive vestiture ventrally); mesotarsi moderately enlarged (tarsomere 1 slightly longer than wide and lacking adhesive vestiture ventrally; tarsomere 2 approximately as long as wide and with adhesive vestiture ventrally; tarsomere 3 about 1.2 times as wide as long and with adhesive vestiture ventrally; and tarsomere 4 distinctly smaller than tarsomeres 2 and 3, deeply concave apically and without adhesive vestiture ventrally). Aedeagus (Figs 27 - 29): Median lobe asymmetrical, markedly bent ventrad behind basal bulb, more faintly bent ventrad in apical portion, and slightly directed dorsad just at apex to form a faintly recognizable apical capitulum; middle portion of ventral margin convex. Terminal lamella (Fig. 27) about 2.0 times as long as wide, flat, triangular, slightly blunted at apex (dorsal aspect) and distinctly directed to right, its dorsal side with a triangular depression in basal quarter. Apical orifice in dorsal position, prolonged to basal bulb. Internal sac with two small and narrow spiny patches basally.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
DFC67FB53E140134742F8529292FD27B.taxon	distribution	Distribution. Fig. 56. Known only from the type locality (Hanmi) in the eastern Himalaya, in the southeastern part of Xizang Autonomous region (Tibet), Medog County, China.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
403F6C1D60AF146C5BC15592CB69794E.taxon	description	Description. Dorsal habitus as in Fig. 54. Size: Body length 8.8 - 10.8 mm, width 3.8 - 4.6 mm. Color: Body black, shiny on dorsum; labrum, also mandibles basally and lateral bead of pronotum in many specimens paler, reddish brown; antennae, palpi, tibiae, and tarsi reddish brown, femora blackish brown. Microsculpture: Head with dorsal microsculpture present throughout in most specimens, comprised of fine isodiametric meshes, more or less effaced on frons and vertex. Pronotum with microsculpture comprised of more or less effaced meshes, more distinct in females than in males. Elytral microsculpture in males comprised of more or less isodiametric meshes, distinct on two or three lateral intervals and in area along basal border, otherwise more or less effaced; in females comprised of isodiametric, nearly granulate meshes in basal half and on two or three lateral intervals apically and of slightly transverse meshes on inner intervals in apical half. Head: Comparatively large (HWmax / PWmax = 0.70 - 0.73 and HWmin / PWmax = 0.60 - 0.66), with micropunctures in areas near frontal foveae and around and behind supraorbital setae in most specimens, micropunctures also present on clypeus and frons in some specimens, micropunctation absent from head in a few specimens; tempora short, nearly flat, sloped to neck. Clypeus slightly concave and distinctly bordered apically. Frontal suture distinct, superficial or slightly impressed. Frontal foveae small and shallow, clypeo-ocular prolongations superficial, short in most specimens but distinct to supraorbital furrows in some specimens. Supraorbital seta situated at level of hind margin of each eye. Eyes small, moderately convex. Antennae short, not extended to pronotal basal margin, with antennomeres 5 to 7 each about 1.6 - 2.0 times as long as wide. Labrum distinctly emarginate apically. Left mandible truncate at apex. Ligular sclerite slightly widened and rounded at apex. Pronotum (Fig. 30): Slightly transverse (PWmax / PL = 1.40 - 1.51), narrowed basad (PWmax / PWmin = 1.25 - 1.36) and widest in anterior third. Sides rounded along their entire length, but in basal half less distinctly than in apical half; without any lateral setigerous pores. Apical margin very slightly concave or nearly straight medially, bordered only laterally. Basal margin more or less straight (very broadly rounded in some specimens), distinctly bordered throughout, width approximately equal to apical margin and slightly narrower than elytral base between humeral angles. Apical angles nearly 90 ° (lateral aspect), slightly protruded anteriad. Basal angles obtuse, each with small denticle at apex. Pronotal disc convex, only faintly depressed basally, markedly sloped to apical angles. Lateral depressions varied, from very narrow along entire length, indistinct, and with area at basal angles convex, to distinctly widened and deepened in basal third and depressed at basal angles. Basal foveae small, either flat or slightly impressed. Pronotal surface densely and distinctly punctate, mainly along sides laterobasally and lateroapically, with punctation more widely distributed over entire basal and apical portions in some specimens, or with very fine punctures also present in central portion in a few specimens, or with punctation restricted only to lateral and latero-basal areas in a few other specimens; in all specimens, punctures coarsest in latero-basal portion and in narrow area along sides. Elytra: Oval, rounded at sides, moderately wide (EL / EW = 1.33 - 1.44, EL / PL = 2.42 - 2.67, EW / PWmax = 1.22 - 1.30), widest at middle, not fused along suture. Humeri subangulate, rounded at apices, each with a tiny denticle visible only from behind. Subapical sinuations moderately deep. Sutural angles not separated from each other medially, slightly less than 90 °, with apices blunted in male and sharp in female. Basal borders slightly sinuate, joined with lateral margin at very obtuse angle. Striae impunctate, slightly impressed along entire length. Parascutellar striae present, short, basal setigerous pores present; in some specimens, first (sutural) striae interrupted basally with distal part of parascutellar striae connected to proximal part of isolated distal portion of first striae. Intervals slightly convex or nearly flat, in some specimens two or three lateral intervals very finely micro-punctate. Umbilicate setal series more or less widely interrupted at middle. Hindwings: Reduced to small scales. Venter: Prosternum smooth and glabrous, with at most a few very fine and barely evident setae apically. Prosternal medial process slightly prominent, not projected posteriad (Fig. 31). Proepisterna smooth, at most finely micropunctate. Metepisterna (Fig. 32) markedly narrowed posteriad, approximately as long as wide or slightly wider than long. Sternum VII (anal) in both sexes with two pairs of setae along apical margin and rounded at apex. Tergum VII (anal) of female rounded at apex. Legs: Metacoxae generally without posteromedial setigerous pore or any additional setigerous or nonsetigerous foveae medially [in one female collected at 0.3 km above Shibali on Yaping road, left metacoxa with an additional posteromedial setigerous pore]. Metafemora with two setae along posterior margin. Protibiae with one ventroapical spine, outer margin with one or two stouter spines and also two or three slenderer spines apically in most males, with four or five uniform, stout spines in females. Tarsi glabrous dorsally, tarsomere 5 with three (four in some specimens) pairs of lateroventral setae. Metatarsi approximately equal in length to minimum linear distance across neck constriction just behind eyes in males, slightly shorter in females; tarsomere 1 distinctly longer than tarsomere 2, but distinctly shorter than tarsomeres 2 + 3. In males, protarsi markedly enlarged (tarsomeres 2 - 4 much wider than long, tarsomere 1 about as long as wide, and tarsomeres 1 - 4 with adhesive vestiture ventrally); mesotarsi moderately enlarged (tarsomere 1 slightly longer than wide; tarsomere 2 approximately as long as wide; tarsomere 3 about 1.3 times as wide as long; and tarsomere 4 distinctly smaller than tarsomeres 2 and 3, and deeply concave apically, and tarsomeres 2 - 4 with adhesive vestiture ventrally). Female genitalia (Figs 33 - 34): Apical stylomere comparatively faintly curved. Aedeagus (Figs 35 - 37): Median lobe markedly bent ventrad behind basal bulb and convex on ventral side medially. Terminal lamella directed ventrad, triangular in dorsal aspect, about two times as long as wide and very narrowly rounded at apex (Fig. 35), dorsal side basally with large triangular depression prolonged apicad up to or beyond middle of terminal lamella; apical capitulum very small, slightly prominent ventrad and dorsad. Apical orifice slightly shifted to right, prolonged to basal bulb. Internal sac with two basal and (in most specimens) one medial spiny patches; medial spiny patch, if present, located on right side of medial lobe.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
403F6C1D60AF146C5BC15592CB69794E.taxon	distribution	Distribution. Fig. 57. This new species is known only from the northern part of the Gaoligong Shan (Fugong County) in northwestern Yunnan Province, China.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
DBD778FE61D0431E379B9FD1707675A7.taxon	description	Description (male). Dorsal habitus as in Fig. 55. Size: Body length 9.5 mm, width 4.1 mm. Morphological characters as described for Chydaeus asetosus except as follows: Color: Body black, shiny on dorsum; labrum, tibiae and tarsi paler, blackish brown; antennae dark brown, palpi lighter brown. Microsculpture: Head with dorsal microsculpture effaced. Pronotum with microsculpture more or less effaced. Head: Large (HWmax / PWmax = 0.71 and HWmin / PWmax = 0.62), covered with very fine micro-punctures on labrum and on areas around frontal foveae, above supraorbital furrows, and around and behind supraorbital setae. Frontal suture thin, slightly impressed. Frontal foveae very shallow, clypeo-ocular prolongations extended to supraorbital furrows. Ligular sclerite with apical angles slightly projected laterally. Pronotum (Fig. 38): Slightly transverse (PWmax / PL = 1.47), slightly narrowed basad (PWmax / PWmin = 1.23), with one lateral seta on each side before middle. Basal margin nearly straight medially and slightly oblique laterally, slightly wider than apical margin. Lateral depressions very narrow and barely evident along entire length. Basal foveae absent and area near basal angles slightly convex, almost flat. Pronotal surface punctate almost throughout, but punctures extremely fine medially. Elytra: Elytra relatively short (EL / EW = 1.40, EL / PL = 2.45, EW / PWmax = 1.19), widest just behind middle, without humeral denticles. Subapical sinuations slightly shallower than in Chydaeus asetosus. Parascutellar striae short, about 2.0 - 3.5 times as long as diameter of parascutellar setigerous pore, not connected distally with first (sutural) striae. Intervals nearly flat, lateral intervals very finely and indistinctly micro-punctate. Umbilicate setal series without distinct gap at middle. Venter: Metepisterna (Fig. 39) markedly narrowed posteriad, slightly wider than long. Legs: Metatarsi slightly shorter than minimum linear distance across neck constriction just behind eyes Aedeagus (Figs 40 - 43): Median lobe similar to that of Chydaeus asetosus, but terminal lamella (Fig. 40) narrower, slightly curved dorsad at apex and without apical capitulum; basal triangular depression on its dorsal side prolonged apicad up to middle of terminal lamella. Internal sac with two basal and one medial spiny patches; medial spiny patch located on left side of median lobe.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
DBD778FE61D0431E379B9FD1707675A7.taxon	distribution	Distribution. Fig. 57. This new species is known only from the type locality, N Baoshan (Baoshan City) in the southern part of the Nu Shan (Mountains) in western Yunnan Province, China.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
5D5AA06659020B34C930CEEDFFA45F4B.taxon	distribution	Distribution. Fig. 58. Chydaeus satoi was previously known only from the type locality (Zhonghe Feng) at an elevation of 2620 m in the Diangcang Shan (Mountains), situated north of Dali, western Yunnan Province, China (Ito, 2003). The additional material listed above was also collected in western Yunnan, but in the Gaoligong Shan, about 150 km to the west of the type locality.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
5D0911EADEBF6128FBF549FE97AB2D12.taxon	distribution	Distribution. Fig. 57. All the previous records for species were restricted to the Central Himalaya, from the eastern part of Nepal to West Bengal (Darjeeling) and Sikkim, at elevations of 1800 - 2900 m (Kataev and Schmidt 2006). Specimens recorded here extend the range of Chydaeus obtusicollis eastward to Xizang Autonomous Region (Cona and Medog counties) and northwestern Yunnan Province (Gongshan and Lushui counties), China. The elevational range of Yunnan records extends from 1400 to 2250 m.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
38BB9D0D5A4B477B5A271FA5FD5C486B.taxon	distribution	Distribution. Fig. 58. This species was previously known from several localities, all restricted to the southern parts of the Gaoligong Shan in western Yunnan Province, China, at elevations of 2200 - 2400 m (Kataev and Schmidt 2006). The new material was also collected on the southern parts of Gaoligong Shan, at elevations of 1800 - 2500 m, within Longling, Longyang, and Tengchong counties, western Yunnan Province, China.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
A5A6428190CC77AD7D128690FDA406C7.taxon	distribution	Distribution. Fig. 59. This species was described from a series collected in Kambaiti, northeastern Myanmar. The new material from Yunnan was collected about 100 km to the northeast of the type locality, in the Pianma area, near the Myanmar border, on the western slope of the Gaoligong Shan, at elevations of 2100 - 2500 m. These new records support Kataev and Schmidt's (2006) suggestion that the geographical range of Chydaeus malaisei is confined to the region west of the crest of the Gaoligong Shan.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
49C58940793551988BC5733F98D2ABB9.taxon	distribution	Distribution. According to Kataev and Schmidt (2006), Chydaeus semenowi is widely distributed over the Himalaya from Uttar Pradesh (India) to Bhutan, at elevations of 2400 - 3800 m. This species is recorded here from China (southern Xizang and northwestern Yunnan) for the first time (Fig. 59).	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
69C830C7CEC35D62A83DC16EBB888B71.taxon	distribution	Distribution. Fig. 60. The nominotypic subspecies of Chydaeus andrewesi was known previously from the eastern part of the Himalaya, from Central Nepal to Bhutan and Myanmar, at elevations of 1600 - 2700 m (Kataev and Schmidt 2006). Based on the new records presented here, this subspecies ranges farther east into western Yunnan Province, China, along the Myanmar border (the Gaoligong Shan in Fugong, Gongshan, Longyang, Lushui, and Tengchong counties, and the Bangma Shan and Ailao Shan (Mountains), at elevations of 1200 - 2720 m. This subspecies is also recorded here from the southern part of Xizang Autonomous Region.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
29B9E5C795DA533DAE33C4F3E055543C.taxon	distribution	Distribution. Fig. 60. Previously, this taxon was known only from the single male collected in Sa Pa, northern Vietnam. Based on the new records presented here, this species occurs not only in the mountains of northern Vietnam, but also in southern Yunnan Province (China) adjacent to the Vietnamese border.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
720B01107DFD555986008475DC12BDC2.taxon	distribution	Distribution. Chydaeus salvazae is distributed over northern Vietnam, southwestern China (Guizhou, Sichuan, and Yunnan provinces) and the Central Himalaya (Sikkim and Nepal) (Kataev and Schmidt 2006). In Yunnan, the species is common in southern (Pingbian and Jinping counties) and western (Longling, Longyang, Shuangjiang, Tengchong, and Yongping counties) parts of the province (Fig. 61).	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
ABB6D3FEEF65568B8943865C92903D21.taxon	distribution	Distribution. Fig. 62. Subspecies Chydaeus bedeli difficilis was known previously from only the type series, which was collected in the Hengduan Shan (Mountains) in northwestern Yunnan Province, China (Kataev and Schmidt 2002). Based on the new records presented here, it occurs also in southeastern Xizang Autonomous Region (Medog County) and western Yunnan (in the Gaoligong Shan and Nu Shan mountains in Fugong, Gongshan, Longyang, Lushui, and Tengchong counties).	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
CA49C16BE87E5732A49D8FD403F9D2F3.taxon	distribution	Distribution. Fig. 62. Chydaeus bedeli interjectus is common in the Eastern Himalaya, from the eastern part of Nepal to Bhutan (Kataev and Schmidt 2002). This taxon was not previously recorded from China; and new records presented here extend its known range to include the southern part Xizang Autonomous Region (Cona, Nyalam, and Yadong counties), at elevations of 2250 - 3900 m.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
BA371E0125FD5677844A9FAA0EDB2877.taxon	distribution	Distribution. Previously, Chydaeus bedeli vietnamensis was known from a single male collected in northern Vietnam (type locality: northern slope of Fansipan Mt., Sa Pa, 22 ° 17 ' N, 103 ° 44 ' E, 1525 m). Based on the new records reported here, this subspecies occurs also in southern Yunnan Province, China (Fig. 62).	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
7C045194BF28E6BFAC37AF44A2AB2A8B.taxon	distribution	Distribution. This species, a member of the irvinei group, was known previously from two isolated areas in the Central Himalaya: to the north of Sikkim in southern Xizang and in the upper Tama Koshi valley in southern Xizang and Central Nepal (Kataev and Schmidt 2002). The new record reported here (Fig. 63) is in the upper Bhote Koshi valley in southern Xizang, not far to the west from the Tama Koshi valley.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
CFC15643BF263C7CDB18E2162111B473.taxon	distribution	Distribution. The geographical range of Chydaeus similis in China extends from Central Sichuan to Yunnan. In Yunnan, it had been recorded previously only from the Gang Shan (Mountains) (Kataev and Schmidt 2002). Based on the new records reported here (Fig. 63), Chydaeus similis is more widely distributed in Yunnan.	en	Kataev, Boris M., Liang, Hongbin, Kavanaugh, David H. (2012): Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini). ZooKeys 171: 39-92, DOI: http://dx.doi.org/10.3897/zookeys.171.2306, URL: http://dx.doi.org/10.3897/zookeys.171.2306
