identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0C081779B91B7842C5B84045FC6523E3.text	0C081779B91B7842C5B84045FC6523E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameroconcava Perkins 2022	<div><p>Hydraena cameroconcava, new species</p><p>Figs. 1, 6 (habitus), 6 (aedeagus), 43 (map)</p><p>Type Material. Holotype (male): Cameroon: Northwest Province. “ BRITISH CAMEROONS / Matute, / Tiko Plantation / 5.v.1949, B. Malkin ” (UKNHM).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the absence of a distinct pronotal macula, the frons distinctly concave between median area and eye, the widely separated metaventral plaques that are markedly converging anteriorly (ratios P2/w/l/s ca. 1/1/4/5), and the aedeagus (Fig. 6). Also refer to the diagnosis of H. camerocorona for comments on the male genitalia.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.20/0.50; head width 0.32; pronotum 0.30/0.42, PA 0.36, PB 0.37; elytra 0.67/0.50. Habitus as illustrated (Fig. 6). Dorsum dark brown to black, frons and pronotal disc darker, light brown clypeus contrasting with black frons, palpi testaceous, tip of last palpomere not darker. Frons distinctly concave between median area and eye. Dorsum of head and pronotum finely sparsely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1–3xpd; clypeus dull, punctures very fine and dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral, and posterosubmedial pronotal foveae rather deep</p><p>Elytra with summit of posterior declivity at or very near posterior 1/3, where elytra transversely quite convex; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 1–2xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another. Abdominal apex with slightly off-center apical notch.</p><p>Venter: Mentum and postmentum densely finely punctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/4/5. P1 laminate; median carina sinuate in profile. P2 l/w ca. 4/1, sides slightly converging toward rather narrow blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques narrow, straight, markedly converging anteriorly, posteriorly separated by ca. 4x anterior separation, not raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS concave. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight.</p><p>Distribution. Currently known only from the type locality (Fig. 43).</p><p>Etymology. Named in reference to the concavities of the frons.</p></div>	https://treatment.plazi.org/id/0C081779B91B7842C5B84045FC6523E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B91B7843C5B8451DFE4C217E.text	0C081779B91B7843C5B8451DFE4C217E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocontrasta Perkins 2022	<div><p>Hydraena camerocontrasta, new species</p><p>Figs. 1, 7 (habitus), 7 (aedeagus), 43 (map)</p><p>Type Material. Holotype (male): Cameroon: Northwest Province. “ BRITISH CAMEROONS. / Mamfe, 7-11.i.1949 / B. Malkin coll. // Rain forest; clear stream / Gravel and sand” (UKNHM).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the light brown clypeus contrasting with dark brown frons, the frons shallowly concave between median area and eye, and the male genitalia (Fig. 7). Also refer to the diagnosis of H. camerocorona for comments on the male genitalia.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.22/0.54; head width 0.30; pronotum 0.29/0.44, PA 0.36, PB 0.39; elytra 0.76/0.54. Habitus as illustrated (Fig. 7). Dorsum brown to dark brown, frons and pronotal disc darker, light brown clypeus contrasting with dark brown frons, palpi testaceous, tip of last palpomere not darker. Frons shallowly concave between median area and eye. Dorsum of head and pronotum finely sparsely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1–3xpd; clypeus dull, punctures very fine and dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral, and posterosubmedial pronotal foveae moderately deep</p><p>Elytra with summit of posterior declivity at or very near posterior 1/3, where elytra transversely quite convex; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 1–2xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another. Abdominal apex with slightly off-center apical notch.</p><p>Venter: Mentum and postmentum moderately finely punctulate, shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel or slightly diverging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques narrow, straight, parallel, raised very slightly, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, protibia slightly arcuate, meso- and metatibiae straight. Abdominal apex with off center apical notch.</p><p>Distribution. Currently known only from the type locality (Fig. 43).</p><p>Etymology. Named in reference to the known distribution and the contrasting sizes of the parameres: right very elongate, left just a nub.</p></div>	https://treatment.plazi.org/id/0C081779B91B7843C5B8451DFE4C217E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B91A7840C5B846E1FF5524F7.text	0C081779B91A7840C5B846E1FF5524F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocorona Perkins 2022	<div><p>Hydraena camerocorona, new species</p><p>Figs. 1, 8 (habitus), 8 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon, Akonolinga, moist secondary forest and plantation, at light, 7.I.1978, Loc. #17 // Gardenforhall—Samuelson ” (LUM).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the very fine and widely spaced elytral punctures, the comparatively wide lateral explanate margins of the elytra, the pronotal macula, and the aedeagus (Fig. 8).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.26/0.56; head width 0.34; pronotum 0.31/0.4, PA 0.37, PB 0.38; elytra 0.75/0.56. Habitus as illustrated (Fig. 8). Dorsum brown to dark brown or black, frons and diffusely margined pronotal macula darker, frons almost black, contrasting with light brown clypeus, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum sparcely finely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1–3xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae moderately deep, shallowly confluent, posterosubmedial foveae very shallow.</p><p>Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins wide; punctures fine, on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 3–4xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</p><p>Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely punctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.7/4/4. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately wide, slightly converging toward one another anteriorly, weakly raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 2x P2. Tibiae slender, protibial very slightly arcuate, meso- and metatibiae straight. Abdominal apex symmetrical; last tergite with slightly off-center apical notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution, and the wide lateral explanate margins of the elytra.</p></div>	https://treatment.plazi.org/id/0C081779B91A7840C5B846E1FF5524F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B9197840C5B84069FAF02377.text	0C081779B9197840C5B84069FAF02377.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerojocula Perkins 2022	<div><p>Hydraena camerojocula, new species</p><p>Figs. 1, 9 (habitus), 9 (aedeagus), 41 (map)</p><p>Type Material. Holotype (male): “ Cameroon: C. S. Prov., 15 km. S. Mbalmayo, Nyong River, ex. leaves at margin, 22.v.1982, leg. P. D. Perkins ” (MCZ).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the posterosubmedian foveae moderately deep, medially confluent forming U-shaped depression, the metaventral plaques moderately narrow posteriorly, tapering to point anteriorly, and the aedeagus (Fig. 9). Dorsal habitus similar to other species in Aedeagal Group 1; reliable determinations will require examination of the male genitalia.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.35/0.60; head width 0.36; pronotum 0.32/0.46, PA 0.37, PB 0.41; elytra 0.82/0.60. Habitus as illustrated (Fig. 9). Dorsum generally dark brown, except labrum, clypeus and anterolateral angles of pronotum lighter, palpi testaceous, tip of last palpomere not darker. Frons moderately finely and moderately densely punctulate, pronotum moderately coarsely, moderately densely punctate, punctures much larger than those of frons, interstices shining, ca. 1–2xpd. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae deep, posterolateral foveae very shallow, posterosubmedian foveae moderately deep, medially confluent forming U-shaped depression.</p><p>Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or slightly greater, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming moderately strong angle with one another. Venter: Mentum sparsely finely punctulate, shining; postmentum very finely densely micropunctulate, surrounding areas smooth, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow posteriorly, tapering to point anteriorly, slightly converging toward one another anteriorly, weakly raised, located at sides of shallow median depression. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, straight. Abdominal apex asymmetrical, with deep off-center notch.</p><p>Distribution. Currently known only from the type locality (Fig. 41).</p><p>Etymology. Named in reference to the known distribution, and to the smile shaped pronotal fovea.</p></div>	https://treatment.plazi.org/id/0C081779B9197840C5B84069FAF02377	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B9197841C5B844E9FB622113.text	0C081779B9197841C5B844E9FB622113.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameroovala Perkins 2022	<div><p>Hydraena cameroovala, new species</p><p>Figs. 1, 10 (habitus), 10 (aedeagus), 41 (map)</p><p>Type Material. Holotype (male): “ Cameroon: C. S. Prov., 15 km. S. Mbalmayo, Nyong River, ex. leaves at margin, 22.v.1982, leg. P. D. Perkins ” (MCZ) . Paratypes: Same data as holotype (4 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the oval body form, the small body size (ca. 1.18 mm), the posterosubmedial foveae moderately deep, confluent medially forming U-shaped impression, and the aedeagus (Fig. 10). Similar in habitus to some other members of Aedeagal Group 1; reliable determinations will require dissection of males.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.18/0.56; head width 0.34; pronotum 0.31/0.44, PA 0.36, PB 0.39; elytra 0.70/0.56. Habitus as illustrated (Fig. 10). Body form quite oval. Dorsum generally brown to dark brown, maxillary palpi, labrum and clypeus testaceous, pronotum with diffusely margined weakly defined macula. Dorsum of head and pronotum moderately coarsely, often very densely punctulate, frons punctures distinctly finer and less dense than pronotal punctures, interstices shining; punctures of frons disc and pronotal disc ca. 2–3xef, interstices ca. 1xpd or less; punctures of clypeus very fine, very dense, dull. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae deep, posterolateral absent, posterosubmedial foveae moderately deep, confluent medially forming U-shaped impression.</p><p>Elytra with summit of posterior declivity at about posterior 1/3; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect very slightly separately rounded, in posterior aspect margins forming strong angle with one another.</p><p>Venter: Mentum sparsely finely punctulate, shining; postmentum very finely densely micropunctulate, surrounding areas smooth, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/3. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow posteriorly, tapering to point anteriorly, slightly converging toward one another anteriorly, weakly raised, located at sides of shallow median depression. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, straight. Abdominal apex asymmetrical, with deep off-center notch.</p><p>Distribution. Currently known only from the type locality (Fig. 41).</p><p>Etymology. Named in reference to the known distribution, and to the oval body form.</p></div>	https://treatment.plazi.org/id/0C081779B9197841C5B844E9FB622113	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B9187846C5B8464DFA78241F.text	0C081779B9187846C5B8464DFA78241F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerosicula Perkins 2022	<div><p>Hydraena camerosicula, new species</p><p>Figs. 1, 11 (habitus), 11 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon: N. Prov., 35 km N. Ngaoundere, stream, 1.v.1982, leg. P. D. Perkins ” (MCZ) . Paratypes: Same data as holotype (3 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the comparatively large body size (ca. 1.49 mm), the deep pronotal impressions, the comparatively wide P2, the narrow median depression between the metaventral plaques, and the aedeagus (Fig. 11).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.49/0.69; head width 0.39; pronotum 0.37/0.51, PA 0.41, PB 0.46; elytra 0.93/0.69. Habitus as illustrated (Fig. 11). Dorsum brown to dark brown, frons and pronotal macula darker, maxillary palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately finely moderately sparsely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1–3xpd; clypeus dull, punctures very fine and dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae deep, confluent with shallow posterolateral; posterosubmedial pronotal foveae moderately deep, markedly punctate.</p><p>Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 1–2xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/1. P1 laminate; median carina sinuate in profile. P2 comparatively wide, l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow, slightly arcuate, parallel, tapering anteriorly, weakly raised, located at sides of moderately deep, narrow median depression. AIS width at straight posterior margin ca. 1.5x P2. Protibia very slender, straight. All tibiae slender, straight. Abdominal apex asymmetrical, with deep off-center notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution, and to the dagger-shaped process of the aedeagus.</p></div>	https://treatment.plazi.org/id/0C081779B9187846C5B8464DFA78241F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B91F7846C5B84399FDA2232F.text	0C081779B91F7846C5B84399FDA2232F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameroclandestina Perkins 2022	<div><p>Hydraena cameroclandestina, new species</p><p>Figs. 3, 12 (habitus), 12 (aedeagus), 40 (map)</p><p>Type Material. Holotype (male): “ Cameroon: N. W. Prov., 7 km. E. Bamenda, stream, 30.i.1982, leg. P. D. Perkins ” (MCZ) . Paratypes: Same data as holotype (5 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the moderately coarsely, very densely punctulate head and pronotum, and the aedeagus (Fig. 12). The dorsal habitus is very similar to that of H. camerocrebrata and H. cameropetila; all three species are in Aedeagal Species Group 2. H.cameroclandestina is larger in body size than H. cameropetila (ca. 1.32 vs. 1.20 mm); also, the P2 is wider than that of the two compared species. Reliable determinations of all three species will require examination of the male genitalia, which distinctly differ from one another.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.32/0.58; head width 0.33; pronotum 0.32/0.47, PA 0.38, PB 0.40; elytra 0.79/0.58. Habitus as illustrated (Fig. 12). Dorsum dark brown to black, pronotum without macula, maxillary palpi brown. Dorsum of head and pronotum moderately coarsely, very densely punctulate, each puncture with short, distinctive whitish seta, interstices dull; punctures of frons disc and pronotal disc ca. 2xef, interstices ca. 1xpd or less; punctures of clypeus fine, dense. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral and posterosubmedial foveae shallow.</p><p>Elytra with summit of posterior declivity at about midlength; lateral explanate margins narrow to moderately wide; on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with short and fine conspicuous whitish seta. Intervals not raised, width ca. 0.5–1xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum and postmentum very densely very finely punctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow, parallel, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 2x P2. All tibiae moderately stout. Protibia very slightly arcuate. Meso- and metatibiae straight. Abdominal apex symmetrical, with very small and shallow apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 40).</p><p>Etymology. Named in reference to the known distribution, and to the close dorsal similarity to H. camerocrebrata n. sp. and H. cameropetila n. sp.</p></div>	https://treatment.plazi.org/id/0C081779B91F7846C5B84399FDA2232F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B91F7847C5B84451FB28263B.text	0C081779B91F7847C5B84451FB28263B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocomplexa Perkins 2022	<div><p>Hydraena camerocomplexa, new species</p><p>Figs. 3, 13 (habitus), 13 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon: Yaounde: Mt. Febe, 950 m., hygropetric, 10.iv.1982, P. D. Perkins collector” (MCZ) . Paratype: Same data as holotype (1 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the small size (ca. 1.16 mm), the relatively large and wide pronotum (length/width ratio ca. 17/25), the moderately deep pronotal foveae, and the aedeagus (Fig. 13).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.16/0.54; head width 0.33; pronotum 0.31/0.46, PA 0.39, PB 0.41; elytra 0.81/0.54. Habitus as illustrated (Fig. 13). Dorsum light brown, head and pronotal macula darker (holotype is teneral), palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum densely moderately finely punctulate, interstices weakly shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1–2xpd; punctures of clypeus very fine and dense. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum relatively large and wide, length/width ratio ca. 17/25. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae shallow, posterosubmedial foveae moderately deep.</p><p>Elytra with posterior declivity very gradual, summit slightly beyond midlength; lateral explanate margins moderately wide; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another. Venter: Mentum and postmentum finely sparsely punctulate, weakly shining. Genae very slightly raised, shining, with weak posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt. Plaques moderately narrow, straight, slightly convergent anteriorly, weakly raised, located at sides of moderately deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 1.5x P2. Pro- meso- and metatibiae moderately stout, straight. Abdominal apex symmetrical, without apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution and the very complex aedeagus.</p></div>	https://treatment.plazi.org/id/0C081779B91F7847C5B84451FB28263B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B91E7844C5B841A5FACF25D6.text	0C081779B91E7844C5B841A5FACF25D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocrebrata Perkins 2022	<div><p>Hydraena camerocrebrata, new species</p><p>Figs. 3, 14 (habitus), 14 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon: Yaounde: Mt. Febe, 950 m., hygropetric, 10.iv.1982, P. D. Perkins collector” (MCZ) . Paratypes (120): Same data as holotype (28 MCZ); same data except 7.iv.1982 (5 MCZ); same data except 18.vi.1983 (64 MCZ); same data except 14.vi.1983 (12 MCZ); Yaounde, Mt. Mbankolo, 900m, hygropetric microhabitat, 20.v.1982, P. D. Perkins collector (11 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the very densely punctate dorsum, the very narrow P2 (l/w ca. 4/1) and the aedeagus (Fig. 14). Also refer to the diagnosis of H. camerocompressa . Although the aedeagi of H. camerocrebrata and H. cameropetila distinctly differ in several characters, a relationship of the two species is indicated by the shapes of the parameres: large, wide and with similar clusters of setae (Figs. 14, 18).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.33/0.57; head width 0.31; pronotum 0.33/0.48, PA 0.41, PB 0.42; elytra 0.85/0.57. Habitus as illustrated (Fig. 14). Dorsum dark brown to black, pronotum without macula, palpi brown. Dorsum of head and pronotum moderately coarsely, very densely punctulate, each puncture with short, distinctive whitish seta, interstices dull; punctures of frons disc and pronotal disc ca. 2–3xef, interstices ca. 1xpd or less; punctures of clypeus fine, dense. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral and posterosubmedial foveae moderately deep.</p><p>Elytra with summit of rather abrupt posterior declivity at about posterior 1/3; lateral explanate margins narrow to moderately wide; on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 0.5xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect very slightly separately rounded, in posterior aspect margins forming very weak angle with one another.</p><p>Venter: Mentum and postmentum nitid, dull. Genae very slightly raised, very weakly shining, with posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/6/6. P1 laminate; median carina weakly sinuate in profile. P2 very narrow, l/w ca. 4/1, sides parallel, apex blunt. Plaques narrow, straight or very slightly arcuate, slightly convergent anteriorly, weakly raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at arcuate posterior margin ca. 2.5x P2. Pro- meso- and metatibiae straight, moderately stout.Abdominal apex symmetrical, without apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution and to the very densely punctate dorsum.</p></div>	https://treatment.plazi.org/id/0C081779B91E7844C5B841A5FACF25D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B91D7844C5B84309FE6A209E.text	0C081779B91D7844C5B84309FE6A209E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocubita Perkins 2022	<div><p>Hydraena camerocubita, new species</p><p>Figs. 3, 15 (habitus), 15 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon: 35 km N. Ngaoundere, stream, 1.v.1982, P. D. Perkins collector” (MCZ) . Paratypes: 12 km. N. Ngaoundere, lake margin, 1.v.1982, P. D. Perkins collector (2 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the small size (ca. 1.26 mm), the metaventral plaques that are very small, very narrow, subcarinate lines, and the aedeagus (Fig. 15). The dorsal habitus is similar to that of H. cameroconfinis n. sp., but the present species is much smaller (ca. 1.26 vs. 1.66 mm), and has different ventral proportions (P2/w/l/s ca. 1/0.15/0.15/2 vs. 1/0.25/1.5/1). The aedeagi of the two species are markedly dissimilar.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.26/0.52; head width 0.33; pronotum 0.30/0.41, PA 0.33, PB 0.37; elytra 0.76/0.52. Habitus as illustrated (Fig. 15). Dorsum brown to light brown, head and pronotal macula dark brown to black, palpi light brown to testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately densely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of frons disc slightly smaller than 1xef, on pronotal disc slightly larger than 1xef, interstices ca. 2–3xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae moderately deep, posterolateral pronotal foveae very shallow, posterosubmedial foveae very shallow.</p><p>Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect slightly separately rounded, in posterior aspect margins forming very shallow angle with one another.</p><p>Venter: Mentum and postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.15/0.15/2. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2.5/1, sides almost parallel, apex blunt. Plaques very small, very narrow, subcarinate lines, straight, convergent anteriorly, very weakly if at all raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 2x P2. Pro-, meso- and metatibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution and the arm-like shape of the gonopore bearing process of the aedeagus.</p></div>	https://treatment.plazi.org/id/0C081779B91D7844C5B84309FE6A209E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B91D7845C5B847C7FEEC263B.text	0C081779B91D7845C5B847C7FEEC263B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocurvata Perkins 2022	<div><p>Hydraena camerocurvata, new species</p><p>Figs. 3, 16 (habitus), 16 (aedeagus), 40 (map)</p><p>Type Material. Holotype (male): “ Cameroon: N. W. Prov., 7 km. E. Bamenda, stream, 30.i.1982, leg. P. D. Perkins ” (MCZ) . Paratype: Same data as holotype (1 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the ovate body form, the transversely convex pronotum, the posterosubmedial foveae shallow, confluent medially forming U-shaped impression, the very narrow P2, and the aedeagus (Fig. 16). Differentiation from other similarly shaped members of Aedeagal Species Group 2 will require careful examination of the aedeagus.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.30/0.57; head width 0.34; pronotum 0.31/0.49, PA 0.39, PB 0.41; elytra 0.80/0.57. Habitus as illustrated (Fig. 16). Body form ovate. Dorsum generally brown to dark brown, maxillary palpi, labrum and clypeus testaceous, pronotum with diffusely margined weakly defined macula. Dorsum of head and pronotum moderately coarsely, moderately densely punctulate, frons punctures distinctly finer and less dense than pronotal punctures, interstices shining; punctures of frons disc and pronotal disc ca. 1–2xef, interstices ca. 1–2xpd; punctures of clypeus fine, moderately dense, interstices shining medially, dull laterally. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum transversely convex, anterior margin weakly emarginate over middle 1/2; anterolateral and posterolateral foveae shallow, posterosubmedial foveae shallow, confluent medially forming U-shaped impression, anteromedial fovea moderately deep.</p><p>Elytra with summit of posterior declivity at about posterior 1/3; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum and postmentum sparsely very finely punctulate, shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/3/3. P1 laminate; median carina sinuate in profile. P2 very narrow, l/w ca. 4/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques narrow, very slightly arcuate, tapering anteriorly and slightly converging toward one another anteriorly, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 3x P2. All tibiae stout. Protibia widened near midlength, very slightly arcuate. Meso- and metatibiae straight. Abdominal apex symmetrical, last tergite rounded, without apicomedian notch, last ventrite emarginate.</p><p>Distribution. Currently known only from the type locality (Fig. 40).</p><p>Etymology. Named in reference to the known distribution, and to the ovate body form and very curvaceous aedeagus.</p></div>	https://treatment.plazi.org/id/0C081779B91D7845C5B847C7FEEC263B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B91C7845C5B841A5FB2022DF.text	0C081779B91C7845C5B841A5FB2022DF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerofoveata Perkins 2022	<div><p>Hydraena camerofoveata, new species</p><p>Figs. 3, 17 (habitus), 17 (aedeagus), 41 (map)</p><p>Type Material. Holotype (male): “ Cameroon: C. S. Prov., 15 km. NW Yaounde, Isali River nr. Nouma, 2.i.1983, leg. P. D. Perkins ” (MCZ).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the small body size (ca. 1.24 mm), the densely coarsely punctate dorsum, the anterolateral and posterolateral foveae deep, shallowly confluent, the posterosubmedian foveae deep, the plaques absent or very small and very faintly indicated by less dense hydrofuge pubescence, located at sides of very shallow median depression, and the aedeagus (Fig. 17). The habitus is quite similar to that of H. camerocrebrata, but that species has well developed metaventral plaques, and a different aedeagus.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.24/0.54; head width 0.31; pronotum 0.31/0.44, PA 0.37, PB 0.38; elytra 0.73/0.54. Habitus as illustrated (Fig. 17). Dorsum dark brown to black, maxillary palpi brown, tip of last palpomere not darker. Frons densely moderately coarsely, pronotum coarsely densely punctate, punctures much larger than those of frons, interstices shining, ca. 0.5–1xpd. Labrum apicomedially excised, free margins very weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral and posterolateral foveae deep, shallowly confluent; posterosubmedian foveae deep; anteromedian fovea shallow.</p><p>Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1–2xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with short and fine seta. Intervals not raised, width ca. 0.5–1xpd, as are interstices between punctures of a row. Apices in dorsal aspect separately rounded, in posterior aspect margins forming moderately strong angle with one another.</p><p>Venter: Mentum and postmentum sparsely finely punctulate, shining. Genae raised, without posterior ridge. P1 laminate; median carina sinuate in profile. P2 l/w ca. 4/1, sides parallel or slightly diverging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques absent or very small and very faintly indicated by less dense hydrofuge pubescence, located at sides of very shallow median depression. AIS width at straight posterior margin ca. 2.5x P2. All tibiae straight, moderately stout. Abdominal apex symmetrical, with very small apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 41).</p><p>Etymology. Named in reference to the known distribution, and to the deep pronotal foveae.</p></div>	https://treatment.plazi.org/id/0C081779B91C7845C5B841A5FB2022DF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B903785AC5B842B8FA6121EB.text	0C081779B903785AC5B842B8FA6121EB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameropetila Perkins 2022	<div><p>Hydraena cameropetila, new species</p><p>Figs. 3, 18 (habitus), 18 (aedeagus), 43 (map)</p><p>Type Material. Holotype (male): Cameroon: Northwest Province. “ BRITISH CAMEROONS. / Mamfe, 7-11.i.1949 / B. Malkin coll. // Rain forest; clear stream / Gravel and sand” (UKNHM) . Paratypes: Same data as holotype (2 UKNHM) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of absence of metaventral plaques, the very narrow P2, and the aedeagus (Fig. 18). For characters differentiating the two species refer to the diagnosis of H. camerosetosa . Although H. cameropetila is smaller than H. camerosetosa in body length (ca. 1.20 vs. 1.33 mm), the width of the mesocoxae are the same in the two species, making the ratio of P2 width/mesocoxa width approximately 0.12 for H. cameropetila and 0.40 H. camerosetosa . Also refer to the diagnoses of H. millenaria and H. camerocrebrata .</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.20/0.50; head width 0.30; pronotum 0.29/0.42, PA 0.32, PB 0.36; elytra 0.74/0.50. Habitus as illustrated (Fig. 18). Dorsum dark brown to black, pronotum without macula, palpi brown. Dorsum of head and pronotum moderately coarsely, very densely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 2–3xef, interstices ca. 1xpd or less; punctures of clypeus moderately fine, dense. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral and posterosubmedial foveae moderately deep, anteromedian fovea shallow.</p><p>Elytra with summit of posterior declivity at about posterior 1/3; lateral explanate margins wide; on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 0.5xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect very slightly separately rounded, in posterior aspect margins forming weak angle with one another.</p><p>Venter: Mentum and postmentum densely finely punctulate, dull. Genae raised, dull, without posterior ridge.</p><p>Metaventral plaques absent. P1 laminate; median carina sinuate in profile. P2 very narrow, l/w ca. 4/1, sides parallel, blunt apex raised slightly above mesoventral intercoxal process.. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 4x P2. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight. Abdominal apex with very small indistinct apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 43).</p><p>Etymology. Named in reference to the distribution and the very narrow mesoventral intercoxal process (P2).</p></div>	https://treatment.plazi.org/id/0C081779B903785AC5B842B8FA6121EB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B903785BC5B847ACFA7E273F.text	0C081779B903785BC5B847ACFA7E273F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameroancylis Perkins 2022	<div><p>Hydraena cameroancylis, new species</p><p>Figs. 4, 19 (habitus), 19 (aedeagus), 41 (map)</p><p>Type Material. Holotype (male): “ Cameroon: C. S. Prov., 15 km. NW Yaounde, Isali River nr. Nouma, 2.i.1983, leg. P. D. Perkins ” (MCZ) . Paratype: Same data as holotype (1 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the anterolateral pronotal foveae large and deep, posterolateral pronotal foveae absent, the brown to dark brown dorsum, and the aedeagus (Fig. 19). Similar in dorsal habitus to H. cameropalmula and H. millenaria; reliable determinations will require examination of the aedeagus.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.35/0.60; head width 0.36; pronotum 0.33/0.48, PA 0.40, PB 0.43; elytra 0.85/0.60. Habitus as illustrated (Fig. 19). Dorsum brown to dark brown, frons and diffusely margined pronotal disc darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately sparcely moderately coarsely punctulate, more sparsely punctulate on pronotal disc than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 1–2xef, interstices ca. 1–2xpd or slightly larger; punctures of clypeus very fine, very dense, dull. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae large and deep, posterolateral pronotal foveae absent, posterosubmedial foveae very shallow.</p><p>Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins wide; punctures moderately coarse, on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, each puncture with very short and very fine seta. Intervals not raised, width ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming moderate angle with one another.</p><p>Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2.5. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques narrow, short, slightly converging toward one another anteriorly, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. All tibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 41).</p><p>Etymology. Named in reference to the known distribution, and to the strong hook-shaped process of the aedeagus.</p></div>	https://treatment.plazi.org/id/0C081779B903785BC5B847ACFA7E273F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B902785BC5B840A1FB5023E3.text	0C081779B902785BC5B840A1FB5023E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocompacta Perkins 2022	<div><p>Hydraena camerocompacta, new species</p><p>Figs. 4, 20 (habitus), 20 (aedeagus), 40 (map)</p><p>Type Material. Holotype (male): “ Cameroon: N. W. Prov., 7 km. E. Bamenda, stream, 30.i.1982, leg. P. D. Perkins ” (MCZ) . Paratypes: Same data as holotype (12 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the relatively large body size (ca. 1.60 mm), the broad body form, the antero- and posterolateral pronotal foveae moderately deep, shallowly confluent, the wide lateral explanate margins of the elytra, and the aedeagus (Fig. 20). Somewhat similar to H. camerocontinua in body size and form, but differing therefrom in denser dorsal punctation, ventral morphology, and aedeagus.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.59/0.71; head width 0.40; pronotum 0.40/0.55, PA 0.45, PB 0.48; elytra 0.93/0.71. Habitus as illustrated (Fig. 20). Dorsum brown to dark brown or black, frons and diffusely margined pronotal disc darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately sparcely moderately coarsely punctulate, more sparsely punctulate on pronotal disc than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 1–2xef, interstices ca. 2–3xpd or slightly larger; punctures of clypeus very fine. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae moderately deep, shallowly confluent, posterosubmedial foveae very shallow.</p><p>Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins wide; punctures moderately coarse, on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, each puncture with very short and very fine seta. Intervals not raised, width ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/3/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques narrow, very slightly arcuate, converging toward one another anteriorly, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. Protibia slender, very slightly arcuate. Meso- and metatibiae slender, straight. Abdominal apex asymmetrical, with off-center deep notch.</p><p>Distribution. Currently known only from the type locality (Fig. 40).</p><p>Etymology. Named in reference to the known distribution, and to the compact body form.</p></div>	https://treatment.plazi.org/id/0C081779B902785BC5B840A1FB5023E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B9027858C5B8451DFF262136.text	0C081779B9027858C5B8451DFF262136.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocompressa Perkins 2022	<div><p>Hydraena camerocompressa, new species</p><p>Figs. 4, 21 (habitus), 21 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon: 35 km N. Ngaoundere, stream, 1.v.1982, P. D. Perkins collector” (MCZ) . Paratypes: Same data as holotype (6 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the very narrow P2 (l/w ca. 4/1), the unusual ratios of P2 width and plaque shape (P2/w/l/s ca. 1/1/6/5), and the aedeagus (Fig. 21). Although the P2 is very narrow, as is that of H. camerocrebrata n. sp., the habitus and aedeagi of the two species markedly differ. H. cameropetila (CAM13) also has a very narrow P2, but lacks metaventral plaques.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.40/0.61; head width 0.33; pronotum 0.33/0.48, PA 0.37, PB 0.39; elytra 0.83/0.61. Habitus as illustrated (Fig. 21). Dorsum brown to light brown, head and pronotal macula darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately densely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1–3xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae shallow, posterosubmedial foveae very shallow.</p><p>Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 1–2xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum and postmentum nitid, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/6/5. P1 laminate; median carina weakly sinuate in profile. P2 narrow, l/ w ca. 4/1, sides parallel, apex blunt. Plaques narrow, straight, parallel, weakly raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 1.5x P2. Pro- meso- and metatibiae slender, straight. Abdominal apex with deep asymmetrical slightly off-center notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution and the very narrow mesoventral intercoxal process (P2).</p></div>	https://treatment.plazi.org/id/0C081779B9027858C5B8451DFF262136	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B9017859C5B846A9FC6C24D3.text	0C081779B9017859C5B846A9FC6C24D3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameroconfinis Perkins 2022	<div><p>Hydraena cameroconfinis, new species</p><p>Figs. 4, 22 (habitus), 22 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon: 35 km N. Ngaoundere, stream, 1.v.1982, P. D. Perkins collector” (MCZ) . Paratypes: Same data as holotype (122 MCZ); 12 km. N. Ngaoundere, lake margin, 1.v.1982, leg. P. D. Perkins (1 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the comparatively large size (ca. 1.62 mm vs. 1.10–1.55 for other known species), the narrow metaventral plaques, and the aedeagus (Fig. 9). The dorsal habitus is similar to that of H. camerocubita, but H. cameroconfinis is much larger (ca. 1.62 vs. 1.26 mm), and has different ventral proportions (P2/w/l/s ca. 1/0.25/1.5/1 vs. 1/0.15/0.15/2). The aedeagi of the two species are markedly dissimilar (Figs. 15, 22).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.62/0.68; head width 0.38; pronotum 0.39/0.52, PA 0.44, PB 0.45; elytra 0.94/0.68. Habitus as illustrated (Fig. 22). Dorsum brown to light brown, head and pronotal macula darker, palpi light brown to testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately densely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 2–3xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral, and posterosubmedial foveae very shallow.</p><p>Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row.Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum and postmentum nitid, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.25/1.5/1. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt. Plaques narrow, straight, slightly converging toward one another anteriorly, weakly raised, located at sides of moderately deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS concave, width at straight posterior margin ca. 1x P2. Pro- and mesotibiae slender, straight. Metatibia slender, straight, inner margin with row of short setae. Abdominal apex with deep asymmetrical slightly off-center notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution, near the northern limit of the Sudano-Guinean zone (zone 2), just south of the very dry Sudano-Sahelian zone (zone 1).</p></div>	https://treatment.plazi.org/id/0C081779B9017859C5B846A9FC6C24D3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B9007859C5B8400DFEED2207.text	0C081779B9007859C5B8400DFEED2207.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerolatula Perkins 2022	<div><p>Hydraena camerolatula, new species</p><p>Figs. 4, 23 (habitus), 23 (aedeagus), 43 (map)</p><p>Type Material. Holotype (male): Cameroon: Northwest Province. “ BRITISH CAMEROONS. / Mamfe, 7-11.i.1949 / B. Malkin coll. // Rain forest; clear stream / Gravel and sand” (UKNHM) . Paratype: Same data as holotype (1 UKNHM) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the moderately densely punctate dorsum, the relatively wide mesoventral intercoxal process (P2, l/w ca. 2/1), and the aedeagus (Fig. 23). Hydraena cameropetila is also densely punctate, but much smaller (1.20), and lacks plaques. H. camerocrebrata is also densely punctate, but much smaller (ca. 1.33 vs. 1.50), and the plaques are larger (P2/w/l/s ratios ca. 1/1/6/6 vs. 1/0.2/2/2). Also refer to the diagnosis of H. camerocrebrata .</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.49/0.62; head width 0.36; pronotum 0.37/0.50, PA 0.42, PB 0.44; elytra 0.88/0.62. Habitus as illustrated (Fig. 23). Dorsum light brown to dark brown, head, pronotal macula and elytra dark brown, pronotum rather widely margined in light brown to testaceous, palpi testaceous, tip of last palpomere not darker. Dorsum of head moderately finely sparsely punctulate, pronotum moderately coarsely, moderately densely punctate, punctures much larger than those of frons, interstices shining, ca. 1–2xpd. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral and posterolateral foveae moderately deep, posterosubmedial foveae very shallow.</p><p>Elytra with summit of posterior declivity slightly beyond midlength, where elytra transversely quite convex; lateral explanate margins moderately wide; on basal 1/3 punctures larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</p><p>Venter: Mentum and postmentum very finely punctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.2/2/2. P1 laminate; median carina sinuate in profile. P2 relatively wide, l/w ca. 2/1, sides parallel or slightly diverging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques very narrow, very indistinct, straight, parallel, not raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 1x P2, or slightly narrower than P2. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight. Abdominal apex with slightly off center shallow notch in tergite, into which minute apical process of ventrite fits.</p><p>Distribution. Currently known only from the type locality (Fig. 43).</p><p>Etymology. Named in reference to the known distribution and the relatively wide mesoventral intercoxal process (P2).</p></div>	https://treatment.plazi.org/id/0C081779B9007859C5B8400DFEED2207	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B907785EC5B842B8FAE62198.text	0C081779B907785EC5B842B8FAE62198.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameropalmula Perkins 2022	<div><p>Hydraena cameropalmula, new species</p><p>Figs. 4, 24 (habitus), 24 (aedeagus), 43 (map)</p><p>Type Material. Holotype (male): Cameroon: Northwest Province. “ BRITISH CAMEROONS / Matute, / Tiko Plantation / 5.v.1949, B. Malkin ” (UKNHM).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the pronotum with a macula, the absence of posterosubmedial pronotal foveae, and the aedeagus (Fig. 24). Reliable determinations to distinguish this species from other similar members in Aedeagal Species Group 3 will require dissection of males.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.33/0.58; head width 0.34; pronotum 0.34/0.47, PA 0.38, PB 0.38; elytra 0.82/0.58. Habitus as illustrated (Fig. 24). Dorsum testaceous to dark brown, head, pronotal macula and elytra dark brown, pronotum rather widely margined in light brown to testaceous, palpi testaceous, tip of last palpomere not darker. Dorsum of head finely sparsely punctulate, pronotum moderately coarsely, densely punctate, punctures much larger than those of frons, interstices shining, ca. 1–2xpd. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae moderately deep, posterolateral foveae very shallow, posterosubmedial foveae absent.</p><p>Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</p><p>Venter: Mentum and postmentum finely punctulate, shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/4/4. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques narrow, straight, parallel or nearly so, not raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 2.5x P2. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight. Abdominal apex with apicomedian shallow notch in tergite, into which minute apical process of ventrite fits.</p><p>Distribution. Currently known only from the type locality (Fig. 43).</p><p>Etymology. Named in reference to the known distribution and the palmate shape of the aedeagus.</p></div>	https://treatment.plazi.org/id/0C081779B907785EC5B842B8FAE62198	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B907785FC5B846C4FDD424AF.text	0C081779B907785FC5B846C4FDD424AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena millenaria Perkins 2022	<div><p>Hydraena millenaria, new species</p><p>Figs. 4, 25 (habitus), 25 (aedeagus), 40 (map)</p><p>Type Material. Holotype (male): “ Cameroon: N. W. Prov., Bamenda, stream nr. Skyline Hotel, 1.ii.1982, leg. P. D. Perkins ” (MCZ) . Paratype: Same data as holotype (1 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the absence of metaventral plaques, the frons and pronotum finely sparsely punctulate, the punctures of the pronotum very consistent in size throughout, and the aedeagus (Fig. 25). The dorsal punctation is much finer and sparser than that of H. cameropetila and H. camerosetosa, two species which also lack metaventral plaques. All three species are in different Aedeagal Species Groups.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.20/0.53; head width 0.33; pronotum 0.31/0.44, PA 0.34, PB 0.39; elytra 0.75/0.53. Habitus as illustrated (Fig. 25). Dorsum brown to dark brown, head, pronotal fascia and elytra dark brown, pronotum rather widely bordered anteriorly and posteriorly slightly lighter brown, maxillary palpi testaceous, tip of last palpomere not darker. Frons and pronotum finely sparsely punctulate, punctures of pronotum slightly larger than those of frons, very consistent in size throughout, interstices shining, ca. 1–3xpd. Clypeus medially very finely sparsely punctulate, shining; laterally densely micropunctulate, dull. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae deep, posterolateral and posterosubmedian foveae shallow.</p><p>Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals shining, not raised, width ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, surrounding areas smooth, shining. Genae raised, shining, without posterior ridge. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques absent. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, straight. Abdominal apex slightly asymmetrical, with deep and wide apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 40).</p><p>Etymology. Named in reference to the fact that the author’s goal of describing more than 1000 new species of water beetles is now happily attained.</p></div>	https://treatment.plazi.org/id/0C081779B907785FC5B846C4FDD424AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B906785FC5B84069FB1223BF.text	0C081779B906785FC5B84069FB1223BF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocarinata Perkins 2022	<div><p>Hydraena camerocarinata, new species</p><p>Figs. 4, 26 (habitus), 26 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon: 35 km N. Ngaoundere, stream, 1.v.1982, P. D. Perkins collector” (MCZ).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the nearly semicordate pronotum, the widely separated and carinate mesoventral plaques, and the aedeagus (Fig. 26). Refer also to the diagnosis of H. camerosemicordata n. sp.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.35/0.57; head width 0.33; pronotum 0.35/0.44, PA 0.37, PB 0.33; elytra 0.81/0.57. Habitus as illustrated (Fig. 26). Pronotum almost semicordiform, anterior margin much wider than posterior margin. Dorsum brown except head and pronotal fascia black, palpi light brown to testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately punctulate, more sparsely punctulate on fascia than on surrounding areas, interstices shining; punctures of frons disc fine and sparse, less than 1xef; punctures of pronotal disc ca. 2xef, interstices ca. 1–3xpd or slightly larger; punctures of clypeus fine, moderately dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae moderately deep, posterosubmedial foveae very shallow.</p><p>Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures 2–3xpd and deeper than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, very narrow, width ca. 1/2xpd or less, as are interstices between punctures of a row, each puncture with very short and very fine seta. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum and postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.15/0.15/2. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2.5/1, sides almost parallel, apex blunt. Plaques very small, very narrow, carinate lines, straight, convergent anteriorly, very weakly raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 2x P2. Protibiae slender, very weakly arcuate. Mesotibiae slender, straight. Metatibiae slender, straight, very slightly wider at midlength than apically. Abdominal apex symmetrical, with apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution and the carinate metaventral plaques.</p></div>	https://treatment.plazi.org/id/0C081779B906785FC5B84069FB1223BF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B906785CC5B84521FB2421A3.text	0C081779B906785CC5B84521FB2421A3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerosemicordata Perkins 2022	<div><p>Hydraena camerosemicordata, new species</p><p>Figs. 4, 27 (habitus), 27 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon: 35 km N. Ngaoundere, stream, 1.v.1982, P. D. Perkins collector” (MCZ) . Paratypes: Same data as holotype (6 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the chordate and convex pronotum, the very narrow metaventral plaques, and the aedeagus (Fig. 27). Differentiated from H. camerocarinata n. sp., a species which also has a semichordate pronotum, by the more convex and more chordate pronotum, the deeper pronotal foveae, the area between the antero- and posterolateral pronotal foveae more strongly convex, the more acuminate elytra, and the different proportions of the ventral characters (P2/w/l/s ca. 1/0.2/2/2 vs. 1/0.15/0.15/2).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.37/0.54; head width 0.31; pronotum 0.37/0.44, PA 0.35, PB 0.31; elytra 0.81/0.54. Habitus as illustrated (Fig. 27). Pronotum semicordiform, anterior margin much wider than posterior margin. Dorsum brown except head and pronotal fascia black, palpi light brown to testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately punctulate, more sparsely punctulate on fascia than on surrounding areas, interstices effacedly microreticulate, weakly shining; punctures of frons disc fine and sparse, 1xef or less; punctures of pronotal disc ca. 1–2xef, interstices ca. 1–3xpd; punctures of clypeus fine, moderately dense. Labrum apicomedial excision deep and wide, free margins weakly upturned, angulate laterally and medially. Pronotum distinctly convex, especially transversely, anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae deep, posterosubmedial foveae deep, area between antero- and posterolateral foveae distinctly convex.</p><p>Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures 1–2xpd and deeper than largest pronotal punctures, punctures becoming gradually slightly smaller toward posterior. Intervals not raised, width ca. 1xpd, interstices between punctures of a row ca. 2–3xpd, each puncture with very short and very fine seta. Apices in dorsal aspect conjointly rather sharply rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum and postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.2/2/2. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2.5/1, sides almost parallel, apex blunt. Plaques very small, very narrow, lines, straight, convergent anteriorly, very weakly if at all raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 2x P2. Pro-, meso- and metatibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution and the semi-chordate pronotum.</p></div>	https://treatment.plazi.org/id/0C081779B906785CC5B84521FB2421A3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B905785DC5B84715FBD6271B.text	0C081779B905785DC5B84715FBD6271B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameroarida Perkins 2022	<div><p>Hydraena cameroarida, new species</p><p>Figs. 5, 28 (habitus), 28 (aedeagus), 39 (map)</p><p>Type Material. Holotype (male): “ Cameroon: N. Prov., 22 km. N. Maroua, desert stream, 14.i.1982, leg. P. D. Perkins ” (MCZ).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the brown to dark brown or black dorsum, the deep anterolateral pronotal foveae, and the aedeagus (Fig. 28). Similar in habitus and size to H. camerocontinua, but differing therefrom in denser dorsal punctation, ventral morphology, and aedeagal characters.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.37/0.57; head width 0.34; pronotum 0.32/0.45, PA 0.38, PB 0.40; elytra 0.80/0.57. Habitus as illustrated (Fig. 28). Dorsum brown to dark brown or black, frons and pronotal disc darker, maxillary palpi light brown, tip of last palpomere not darker. Frons finely moderately densely punctulate, ca. 1xef, interstices ca. 1–2xpd, interstices shining. Pronotum moderately coarsely moderately densely punctate, ca. 2xpd of frons punctures; interstices ca. 1xpd or slightly greater, shining. Clypeus medially very finely sparsely punctulate, shining; laterally very finely densely punctulate, dull. Labrum apicomedially excised, free margins not upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae deep, posterolateral shallow, posterosubmedial moderately deep.</p><p>Elytra with summit of posterior declivity at or very near posterior 1/3; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</p><p>Venter: Mentum sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques narrow, straight or very slightly arcuate, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. Protibia very slender, straight. All tibiae slender, straight. Abdominal apex symmetrical, with deep apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 39).</p><p>Etymology. Named in reference to the known distribution, and to the arid area.</p></div>	https://treatment.plazi.org/id/0C081779B905785DC5B84715FBD6271B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B904785DC5B84045FAD62353.text	0C081779B904785DC5B84045FAD62353.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerobuloba Perkins 2022	<div><p>Hydraena camerobuloba, new species</p><p>Figs. 5, 29 (habitus), 29 (aedeagus), 43 (map)</p><p>Type Material. Holotype (male): Cameroon: Northwest Province. “ BRITISH CAMEROONS / Matute, / Tiko Plantation / 5.v.1949, B. Malkin ” (UKNHM) . Paratypes: Same data as holotype (48 UKNHM) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the pronotum color, mostly black, contrasting with light brown to testaceous color narrowly along anterior and posterior margins, the elytral margins in posterior aspect forming very strong angle with one another, and the aedeagus (Fig. 29).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.25/0.54; head width 0.33; pronotum 0.30/0.43, PA 0.38, PB 0.39; elytra 0.75/0.54. Habitus as illustrated (Fig. 29). Dorsum with head and most of pronotum black, pronotum narrowly along anterior and posterior margins light brown to testaceous, palpi testaceous. Frons and clypeus finely, moderately densely punctulate, interstices shining. Pronotum moderately coarsely, densely punctulate, interstices shining; punctures of pronotal disc ca. 2–3xef, interstices ca. 1xpd or less. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral and posterosubmedial foveae moderately deep.</p><p>Elytra with summit of posterior declivity at about posterior 1/3; lateral explanate margins moderately wide; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect slightly separately rounded, in posterior aspect margins forming very strong angle with one another.</p><p>Venter: Mentum and postmentum moderately finely punctulate, shining. Genae raised, dull, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow, straight, parallel or nearly so, not raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight.</p><p>Distribution. Currently known only from the type locality (Fig. 43).</p><p>Etymology. Named in reference to the distribution and the relatively large distal lobe of the aedeagus.</p></div>	https://treatment.plazi.org/id/0C081779B904785DC5B84045FAD62353	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B9047852C5B8448DFB2126A6.text	0C081779B9047852C5B8448DFB2126A6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocameola Perkins 2022	<div><p>Hydraena camerocameola, new species</p><p>Figs. 5, 30 (habitus), 30 (aedeagus), 39 (map)</p><p>Type Material. Holotype (male): “ Cameroon: N. Prov., 22 km. N. Maroua, desert stream, 14.i.1982, leg. P. D. Perkins ” (MCZ) . Paratypes: Same data as holotype (9 MCZ); Same locality, 27.v.1982, leg. P. D. Perkins (5 MCZ)</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the rather elongate body form, the very coarse and very dense pronotal and elytral punctures, interstices narrow walls, shining, the narrow, long plaques, located at sides of deep longitudinal groove-like median sulcus, and the aedeagus (Fig. 30).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.48/0.60; head width 0.36; pronotum 0.34/0.44, PA 0.40, PB 0.42; elytra 0.92/0.60. Habitus as illustrated (Fig. 30). Body form elongate. Dorsum generally brown to dark brown, maxillary palpi, testaceous, pronotum with diffusely margined weakly defined dark brown macula. Dorsum of head and pronotum coarsely, very densely punctulate. Frons punctures distinctly smaller and less dense than pronotal punctures, interstices shining; punctures of frons disc and ca. 2–3xef, interstices ca. 1–2xpd. Pronotal punctures very coarse and very dense, interstices narrow walls, shining. Clypeus very finely sparsely punctulate in median 1/3, shining; lateral areas densely micropunctulate, dull. Labrum apicomedially excised, free margins rounded, very weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral foveae deep, confluent, posterosubmedial foveae deep, very coarsely punctate.</p><p>Elytra with summit of posterior declivity at about midlength; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures. Intervals not raised, width ca. 0.5xpd or less, sometimes narrow walls, as are interstices between punctures of a row. Apices in dorsal aspect jointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum and postmentum rather coarsely densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2.3/1.3. P1 laminate; median carina sinuate in profile. P2 narrow, l/w ca. 3/1, sides slightly converging toward blunt apex raised slightly above mesoventral intercoxal process. Plaques narrow, long compared to P2, straight, slightly converging toward one another anteriorly, weakly raised, located at sides of deep longitudinal groove-like median sulcus. AIS width at straight posterior margin ca. 2x P2. Protibia moderately arcuate, slightly widened on inner margin over distal 1/3. Meso- and metatibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 39).</p><p>Etymology. Named in reference to the known distribution, and to the cameo-like aedeagus.</p></div>	https://treatment.plazi.org/id/0C081779B9047852C5B8448DFB2126A6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B90B7853C5B841D9FE8C2443.text	0C081779B90B7853C5B841D9FE8C2443.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocontinua Perkins 2022	<div><p>Hydraena camerocontinua, new species</p><p>Figs. 5, 31 (habitus), 31 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon: N. Prov., 35 km N. Ngaoundere, stream, 1.v.1982, leg. P. D. Perkins ” (MCZ).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the relatively wide and short P2, l/w ca. 1/1, sides slightly diverging toward apex, apex very indistinct, fused with and at same level as mesoventral intercoxal process., the anterolateral pronotal foveae moderately deep, posterolateral pronotal foveae very shallow or absent, and the aedeagus (Fig. 31). Somewhat similar in habitus to H. cameroarida, but differing in finer, sparser dorsal punctation, ventral characters, and aedeagal morphology.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.47/0.67; head width 0.39; pronotum 0.37/0.52, PA 0.42, PB 0.47; elytra 0.91/0.67. Habitus as illustrated (Fig. 31). Dorsum brown to dark brown or black, frons and diffusely margined pronotal macula darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum sparcely moderately finely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1–3xpd or slightly larger; punctures of clypeus obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae moderately deep, posterolateral pronotal foveae very shallow or absent, posterosubmedial foveae very shallow.</p><p>Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins wide; punctures fine, on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 2–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</p><p>Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.3/1/1.3. P1 laminate; median carina sinuate in profile. P2 relatively wide and short, l/w ca. 1/1, sides slightly diverging toward apex, apex very indistinct, fused with and at same level as mesoventral intercoxal process. Plaques narrow, slightly converging toward one another anteriorly, very weakly raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.3x P2. Protibia slender, arcuate. Meso- and metatibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution, and to the fused intercoxal processes of the meso- and metaventrites.</p></div>	https://treatment.plazi.org/id/0C081779B90B7853C5B841D9FE8C2443	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B90A7853C5B843BDFB62209F.text	0C081779B90A7853C5B843BDFB62209F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameroconvexa Perkins 2022	<div><p>Hydraena cameroconvexa, new species</p><p>Figs. 5, 32 (habitus), 32 (aedeagus), 41 (map)</p><p>Type Material. Holotype (male): “ Cameroon: C. S. Prov., 15 km. NW Yaounde, Isali River nr. Nouma, 2.i.1983, leg. P. D. Perkins ” (MCZ).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the large body size (ca. 1.74 mm, largest known in Cameroon), the pronotum with sides markedly arcuate over middle 1/3, the oval, transversely quite convex elytra, the very wide P1 and P2, the plaques widely separated carinate lines, and the aedeagus (Fig. 32).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.74/0.79; head width 0.42; pronotum 0.42/0.58, PA 0.47, PB 0.48; elytra 1.05/0.79. Habitus as illustrated (Fig. 32). Pronotum with sides markedly arcuate over middle 1/3, sides more strongly sinuate posterior to middle than anterior to middle. Frons finely and moderately sparsely punctulate, punctures ca. 1xef, interstices ca. 1–3xpd, shining. Pronotum coarsely densely punctate in anterior and posterior 1/3, interstices ca. 1xpd, shining; disc convex, very finely very sparsely punctulate, interstices ca. 3–6xpd, shining. Punctures of clypeus very fine, sparse on disc, interstices shining, very dense, dull laterally. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin emarginate over middle 1/2; anterolateral pronotal foveae deep; posterolateral and posterosubmedial pronotal foveae very shallow.</p><p>Elytra oval, transversely quite convex, posterior declivity abrupt, summit of at or very near midlength; lateral explanate margins wide; punctures coarse, on basal 1/3 punctures ca. 2xpd of largest pronotal punctures, each puncture with very short and very fine seta. Intervals not raised, width ca. 2–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum finely sparsely punctulate, shining. Postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.2/1/1. P1 and P2 very wide. P2 l/w ca. 1/1, sides parallel, apex blunt. Plaques widely separated carinate lines, located at sides of deep median depression. AIS very wide, width at straight posterior margin ca. 1.2x P2. All tibiae slender. Protibia very slightly arcuate. Meso- and metatibiae straight. Abdominal apex asymmetrical, with deep apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 41).</p><p>Etymology. Named in reference to the known distribution, and to the convex dorsum.</p></div>	https://treatment.plazi.org/id/0C081779B90A7853C5B843BDFB62209F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B90A7850C5B847C1FADD263B.text	0C081779B90A7850C5B847C1FADD263B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerocordata Perkins 2022	<div><p>Hydraena camerocordata, new species</p><p>Figs. 5, 33 (habitus), 33 (aedeagus), 41 (map)</p><p>Type Material. Holotype (male): “ Cameroon: C. S. Prov., Etoug Ebe, stream with dead leaves, 22.iv.1982, leg. P. D. Perkins ” (MCZ) . Paratypes: Same data as holotype (2 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the large body size (ca. 1.53 mm), the markedly cordiform pronotum, the very wide P1 and P2, and the aedeagus (Fig. 33).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.53/0.69; head width 0.36; pronotum 0.38/0.52, PA 0.41, PB 0.38; elytra 0.90/0.69. Habitus as illustrated (Fig. 33). Pronotum markedly cordiform, width at midlength much wider than anterior and posterior margins. Dorsum dark brown to black, except labrum, clypeus, and maxillary palpi light brown to testaceous, tip of last palpomere not darker. Frons moderately finely moderately sparsely punctate, punctures ca. 1xef. Pronotum coarsely densely punctate, especially anterior and posterior to moderately convex disc, interstices in densely punctate areas narrow walls; interstices on disc ca. 1xpd, shining; punctures laterally much smaller and sparser. Punctures of clypeus very fine, moderately dense on disc, very dense laterally. Labrum apicomedian excision deep and wide, free margins weakly upturned, angulate laterally and medially. Pronotum anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae deep, shallowly confluent; posterosubmedial foveae deep, area between antero- and posterolateral foveae distinctly convex.</p><p>Elytra with posterior declivity rather abrupt, summit at or very near midlength; lateral explanate margins wide; on basal 1/3 punctures 1xpd or slightly larger and deeper than largest pronotal punctures, punctures becoming gradually slightly smaller laterally and toward posterior. Intervals not raised, width on disc ca. 1xpd, much wider laterally and posteriorly, interstices between punctures of a row ca. 1–6xpd, each puncture with very short and very fine seta. Apices in dorsal aspect conjointly rather sharply rounded, in posterior aspect margins forming shallow angle with one another.</p><p>Venter: Mentum finely sparsely punctulate, shining. Postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.1/0.2/1.5. P1 and P2 very wide. P2 l/w ca. 1/1, sides parallel, apex blunt. Plaques very small, very short, widely separated carinate lines, located at sides of deep median depression. AIS very wide, weakly concave, width at straight posterior margin ca. 1.2x P2. Tibiae slender, straight. Protibia very slightly widened and bearing short spine at ca. distal 2/3. Mesotibia slightly widened at ca. distal 2/3. Abdominal apex symmetrical, with rather wide and deep apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 41).</p><p>Etymology. Named in reference to the known distribution, and to the markedly cordiform pronotum.</p></div>	https://treatment.plazi.org/id/0C081779B90A7850C5B847C1FADD263B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B9097851C5B841A5FF0425D7.text	0C081779B9097851C5B841A5FF0425D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerodidacta Perkins 2022	<div><p>Hydraena camerodidacta, new species</p><p>Figs. 5, 34 (habitus), 34 (aedeagus), 40 (map)</p><p>Type Material. Holotype (male): “ Cameroon: N. W. Prov., 7 km. E. Bamenda, stream, 30.i.1982, leg. P. D. Perkins ” (MCZ).</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the comparatively large body size (ca. 1.65 mm), the finely sparsely punctulate frons and pronotum, punctures very consistent in size throughout, interstices shining, the anterolateral foveae moderately deep, posterolateral and posterosubmedian foveae very shallow or absent, the very narrow plaques, with margins very ill-defined, and the aedeagus (Fig. 34).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.65/0.69; head width 0.38; pronotum 0.38/0.53, PA 0.42, PB 0.48; elytra 0.98/0.69. Habitus as illustrated (Fig. 34). Dorsum brown to dark brown, head, pronotal fascia and elytra dark brown, pronotum rather widely bordered anteriorly and posteriorly in light brown, maxillary palpi testaceous, tip of last palpomere not darker. Frons and pronotum finely sparsely punctulate, punctures of pronotum slightly larger than those of frons, very consistent in size throughout, interstices shining, ca. 1–3xpd. Clypeus medially very finely sparsely punctulate, shining; laterally densely micropunctulate, dull. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae moderately deep, posterolateral and posterosubmedian foveae very shallow or absent.</p><p>Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals shining, not raised, width ca. 2–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect rather sharply conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely very densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.3/1.3/1.3. P1 laminate; median carina sinuate in profile. P2 moderately wide, l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques very narrow, margins very ill-defined, slightly arcuate, slightly converging toward one another anteriorly, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1x P2.All tibiae slender. Protibia and mesotibia very weakly arcuate. Metatibia straight. Abdominal apex symmetrical, with deep, narrow apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 40).</p><p>Etymology. Named in reference to the known distribution, and to the two finger-like processes of the aedeagus.</p></div>	https://treatment.plazi.org/id/0C081779B9097851C5B841A5FF0425D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B9087851C5B84309FA652033.text	0C081779B9087851C5B84309FA652033.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerodigita Perkins 2022	<div><p>Hydraena camerodigita, new species</p><p>Figs. 5, 35 (habitus), 35 (aedeagus), 40, 42 (maps)</p><p>Type Material. Holotype (male): “ Cameroon: N. Prov., 35 km N. Ngaoundere, stream, 1.v.1982, leg. P. D. Perkins ” (MCZ) . Paratypes: Same data as holotype (2 MCZ); N. W. Prov., Bamenda, stream nr. Skyline Hotel, 1.ii.1982, leg. P. D. Perkins (4 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the narrow P2 (l/w ca. 4/1), the very small, short and narrow plaques, the deep anterolateral pronotal foveae, and the aedeagus (Fig. 35). Similar in habitus to H. camerosetosa, but differing therefrom in ventral and aedeagal characters.</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.25/0.56; head width 0.34; pronotum 0.32/0.43, PA 0.37, PB 0.38; elytra 0.76/0.56. Habitus as illustrated (Fig. 35). Dorsum brown to dark brown, frons and pronotal disc often darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately finely and moderately sparsely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1–2xpd; clypeus dull, punctures very fine and dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae deep, posterolateral and posterosubmedial pronotal foveae moderately deep</p><p>Elytra with posterior declivity very gradual, summit of at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 1–2xpd or slightly greater, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</p><p>Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/ w/l/s) ca. 1/0.5/1/1.5. P1 laminate; median carina sinuate in profile. P2 l/w ca. 4/1, sides parallel, blunt apex raised slightly above mesoventral intercoxal process. Plaques very small, short, narrow, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. All tibiae slender, straight. Abdominal apex symmetrical, with apicomedian notch.</p><p>Distribution. Currently known only from the type locality and a site at Bamenda (Figs. 40, 42).</p><p>Etymology. Named in reference to the known distribution, and to the finger-like main-piece of the aedeagus.</p></div>	https://treatment.plazi.org/id/0C081779B9087851C5B84309FA652033	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B9087856C5B847ADFACD2763.text	0C081779B9087856C5B847ADFACD2763.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameroflagellata Perkins 2022	<div><p>Hydraena cameroflagellata, new species</p><p>Figs. 5, 36 (habitus), 36 (aedeagus), 42 (map)</p><p>Type Material. Holotype (male): “ Cameroon: C. S. Prov., 15 km. S. Mbalmayo, Nyong River, ex. leaves at margin, 22.v.1982, leg. P. D. Perkins ” (MCZ) . Paratypes: Same data as holotype (2 MCZ); Akonolinga, moist secondary forest and plantation, at light, 7.i.1978, leg. Gardensforhall &amp; Samuelson (1 LUM) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the small size (ca. 1.05 mm, smallest species in Cameroon), the very narrow, very short, carinate or subcarinate, and widely separated plaques (P2/w/l/s ca. 1/0.2/2/2), and the male genitalia (Fig. 36).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.05/0.45; head width 0.27; pronotum 0.27/0.37, PA 0.31, PB 0.33; elytra 0.77/0.45. Habitus as illustrated (Fig. 36). Dorsum light brown to brown, head, pronotal macula and elytra dark brown, pronotum rather widely margined in light brown to testaceous, palpi testaceous, tip of last palpomere not darker. Dorsum of head finely moderately sparsely punctulate, pronotum moderately finely, moderately densely punctate, punctures ca. 1xpd of largest punctures of frons, interstices shining, ca. 1–3xpd. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae moderately deep, posterolateral shallow or absent, posterosubmedian foveae shallow, medially confluent forming U-shaped impression.</p><p>Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row.Apices in dorsal aspect separately rounded, in posterior aspect margins forming weak angle with one another.</p><p>Venter: Mentum moderately sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.2/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques very narrow, slightly arcuate, slightly converging toward one another anteriorly, raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. Protibia slender, very weakly arcuate. Meso- metatibiae slender, straight. Abdominal apex symmetrical, with very small apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 42).</p><p>Etymology. Named in reference to the known distribution, and to the very long aedeagal flagellum.</p></div>	https://treatment.plazi.org/id/0C081779B9087856C5B847ADFACD2763	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B90F7856C5B8409DFB1823E3.text	0C081779B90F7856C5B8409DFB1823E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena cameroradiata Perkins 2022	<div><p>Hydraena cameroradiata, new species</p><p>Figs. 5, 37 (habitus), 37 (aedeagus), 41 (map)</p><p>Type Material. Holotype (male): “ Cameroon: C. S. Prov., Yaounde: Bastos, puddle, 12.xii.1981, leg. P. D. Perkins ” (MCZ) . Paratypes: Same data as holotype (10 MCZ) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the very sparcely and very finely punctulate head and pronotum, interstices strongly shining, the anterolateral pronotal foveae moderately deep, posterolateral and posterosubmedial foveae absent, the absence of metaventral plaques, and the aedeagus (Fig. 37).</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.33/0.59; head width 0.34; pronotum 0.35/0.47, PA 0.38, PB 0.43; elytra 0.80/0.59. Habitus as illustrated (Fig. 37). Dorsum brown to light brown, frons and diffusely margined pronotal macula slightly darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum sparcely finely punctulate, pronotum more finely sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of pronotal disc ca. 1xef, interstices ca. 3–5xpd or slightly larger; punctures of frons disc slightly larger and more closely spaced than those of pronotal disc; punctures of clypeus very fine, nearly obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae moderately deep, posterolateral and posterosubmedial foveae absent.</p><p>Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins narrow; punctures fine, on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 2–4xpd, as are interstices between punctures of a row. Apices in dorsal aspect separately rounded, in posterior aspect margins forming moderately strong angle with one another.</p><p>Venter: Mentum and postmentum sparsely finely punctulate, shining. Genae raised, shining, without posterior ridge. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques absent, median depression deep. AIS width at straight posterior margin ca. 2x P2. All tibiae slender. Pro- and mesotibiae very slightly arcuate, metatibia straight. Abdominal apex symmetrical, with shallow apicomedian notch.</p><p>Distribution. Currently known only from the type locality (Fig. 41).</p><p>Etymology. Named in reference to the known distribution, and to the highly reflective dorsum.</p></div>	https://treatment.plazi.org/id/0C081779B90F7856C5B8409DFB1823E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
0C081779B90F7857C5B84555FB43217E.text	0C081779B90F7857C5B84555FB43217E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraena camerosetosa Perkins 2022	<div><p>Hydraena camerosetosa, new species</p><p>Figs. 5, 38 (habitus), 38 (aedeagus), 43 (map)</p><p>Type Material. Holotype (male): Cameroon: Northwest Province. “ BRITISH CAMEROONS / Matute, / Tiko Plantation / 5.v.1949, B. Malkin ” (UKNHM) . Paratypes: Same data as holotype (10 UKNHM) .</p><p>Differential Diagnosis. Differentiated from other members of the genus in Cameroon by the combination of the densely punctate dorsum, the absence of metaventral plaques, and the aedeagus (Fig. 38). H. camerosetosa differs from H. cameropetila, another species that lacks metaventral plaques, by a larger body size (ca. 1.33 vs. 1.20 mm), a markedly different aedeagus, and a wider mesoventral intercoxal process (P2), which is twice as wide as the P2 of H. cameropetila . Although H. cameropetila is smaller than H. camerosetosa in body length, the width of the mesocoxae are the same in the two species, making the ratio of P2 width/mesocoxa width approximately 0.12 for H. cameropetila and 0.40 H. camerosetosa .</p><p>Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.33/0.55; head width 0.34; pronotum 0.34/0.46, PA 0.39, PB 0.42; elytra 0.78/0.55. Habitus as illustrated (Fig. 38). Dorsum testaceous to black, head and pronotal macula dark brown to black, pronotum rather widely margined in light brown to testaceous, elytra brown, palpi testaceous, tip of last palpomere not darker. Dorsum of head finely sparsely punctulate, pronotum moderately coarsely, densely punctate, punctures much larger than those of frons, interstices shining, ca. 1xpd or less. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral, posterolateral, and posterosubmedial foveae very shallow.</p><p>Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect separately rounded, in posterior aspect margins forming distinct angle with one another.</p><p>Venter: Mentum finely punctulate, shining. Postmentum densely micropunctulate, dull. Genae raised, dull, without posterior ridge. Metaventral plaques absent. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, protibia very slightly arcuate, meso- and metatibiae straight. Abdominal apex with apicomedian indentation.</p><p>Distribution. Currently known only from the type locality (Fig. 43).</p><p>Etymology. Named in reference to the entirely setose metaventrite, which lacks plaques.</p></div>	https://treatment.plazi.org/id/0C081779B90F7857C5B84555FB43217E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2022): Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). Zootaxa 5203 (1): 1-66, DOI: 10.11646/zootaxa.5203.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5203.1.1
