identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
7606A5B8245F52F7ADF39BA5978B68C1.text	7606A5B8245F52F7ADF39BA5978B68C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Erhaia Davis & Kuo 1985	<div><p>Genus Erhaia Davis &amp; Kuo, 1985</p><p>Type species by original designation.</p><p>Erhaia daliensis Davis &amp; Kuo, in Davis et al. 1985.</p><p>Shells.</p><p>The shells vary from conical to more or less ovoid, rarely with a flaring final part of the body whorl. The apex is flattened as in the European amnicolid genus Bythinella Moquin-Tandon, 1856, because of the very low spiral of the protoconch. The peristome is continuous and may be more or less protruding. The parietal and the columellar side of the aperture are about equally long and the regularly curved palatal side of the aperture gradually passes into the basal side, forming a single, regularly curved border. Bythinella cannot be distinguished from Erhaia conchologically, but in some Erhaia species from China the columella has one or two spiral lamellae, that are not known from Bythinella. In Erhaia the protoconch may have spiral striae, which have not been described for any of the Bythinella species.</p><p>Distribution.</p><p>The genus Erhaia is mainly known from China, where it has been recorded with various species from the province of Yunnan in the west to the provinces of Hunan, Hubei and Fujian in the east (Davis et al. 1985; Davis and Kang 1995; Davis and Rao 1997; Wilke et al. 2000, 2001; Liu et al. 2014). One species was described from northern India (Davis and Rao 1997), three from Nepal (Nesemann et al. 2007) and one from Bhutan (Gittenberger et al. 2017a). Here we deal with the systematics of only the species occurring in the southern Himalayan foothills in Bhutan, India, and Nepal.</p></div>	https://treatment.plazi.org/id/7606A5B8245F52F7ADF39BA5978B68C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gittenberger, Edmund;Leda, Pema;Wangchuk, Jigme;Gyeltshen, Choki;Bjoern Stelbrink,	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
C16E201EF7AA5B7BB2E1D9D2C4050BD6.text	C16E201EF7AA5B7BB2E1D9D2C4050BD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Erhaia jannei Gittenberger & Stelbrink 2020	<div><p>Erhaia jannei Gittenberger &amp; Stelbrink sp. nov. Figs 2, 3, 11</p><p>Erhaia sp. Gittenberger et al. 2017a: 25, fig. 3; 2017c: 900, 903, fig. 8.</p><p>Material examined.</p><p>Holotype. (Fig. 2) Bhutan • District Thimphu, c. 5 km E of Chhuzom, W of Geneykha; in brooklet with a prayer wheel along the road; 2750 m a.s.l.; 27°18'43"N, 89°36'10"E; E. Gittenberger, Choki Gyeltshen &amp; Pema Leda leg. 25.X.2018; NBCB 1057. Paratypes. (Fig. 3) 6 shells and 2 animals in ethanol 70%; same data as for holotype; NBCB 1058.</p><p>Diagnosis.</p><p>Shell large for the genus (H&gt; 2mm), ovoid, with a relatively large aperture.</p><p>Shell.</p><p>Shell obliquely ovoid, with 3½ -4 convex, shouldered whorls that are separated by a deep suture; clearly higher than broad; yellowish brown with fine irregular growth lines and some blackish brown periostracal ridges, one of which runs from the apertural columellar border into the umbilicus. Peristome not reflected. Parietal, columellar and a short part of the adjoining basal apertural border thickened by a whitish callus. Most specimens with a continuous peristome and a narrow umbilical chink. Protoconch encrusted in all specimens; teleoconch without spiral sculpture.</p><p>Measurements.</p><p>(N = 9): H 2.2-2.4 mm, B 1.5-1.6 mm. Holotype 2.2 × 1.6 mm.</p><p>Ecology</p><p>(Fig. 11). The species is uncommon on the rocks in the shaded streamlet uphill of the prayer wheel, where it occurs with Galba truncatula ( Müller, 1774), Physa sp. (new for Bhutan), and Erhaia pelkiae sp. nov.</p><p>DNA data</p><p>(Fig. 18). The three individuals shared the same haplotype for both 16S rRNA (GenBank acc. no. MT239078) and COI (GenBank acc. no. MT237716). The uncorrected genetic p-distances between this species and E. wangchuki were 0.84% for 16S rRNA and 4.87% for COI. The distances were considerably higher when compared to Erhaia sp. from China, Guangxi, viz. 2.74% for 16S rRNA and 10.25% for COI (GenBank acc. nos. KC832722 and KC832701, respectively).</p><p>Notes.</p><p>This species was discovered in 2012, but since only a single shell was collected then, a description was considered premature.</p><p>Etymology.</p><p>The epithet jannei refers to Mr Janne Clewing, the son of the last author.</p></div>	https://treatment.plazi.org/id/C16E201EF7AA5B7BB2E1D9D2C4050BD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gittenberger, Edmund;Leda, Pema;Wangchuk, Jigme;Gyeltshen, Choki;Bjoern Stelbrink,	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
819F33E69E015A96978B52B692112C7D.text	819F33E69E015A96978B52B692112C7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Erhaia pelkiae Gittenberger & Gyeltshen 2020	<div><p>Erhaia pelkiae Gittenberger &amp; Gyeltshen sp. nov. Figs 4, 5, 11</p><p>Material examined.</p><p>Holotype. (Fig. 4) Bhutan • District Thimphu, c. 5 km E of Chhuzom, W of Geneykha; in brooklet with a prayer wheel along the road; 2750 m a.s.l.; 27°18'43"N, 89°36'10"E; E. Gittenberger, Choki Gyeltshen &amp; Pema Leda leg. 25.X.2018; NBCB 1059. Paratype. (Fig. 5) 1 shell; same data as for holotype; NBCB 1060.</p><p>Diagnosis.</p><p>Shell with a partly reflected peristome, teleoconch with spiral lirae.</p><p>Shell.</p><p>Shell elongated ovoid, with 3½ convex, shouldered whorls that are separated by a deep suture; clearly higher than broad; light yellowish brown with fine growth lines and some brown periostracal ridges. Peristome reflected at the columellar and the basal side. Parietal and columellar side of the aperture thickened by a whitish callus. An irregular umbilical chink only in the paratype might be represent a malformation resulting from repair of the shell wall. Protoconch encrustated; teleoconch with fine spiral lines.</p><p>Measurements.</p><p>(N = 2): holotype and paratype H 1.9 mm, B 1.1 mm.</p><p>Ecology</p><p>(Fig. 11). The snails are rare on the rocks in the shaded streamlet uphill the prayer wheel. See also the data for E. jannei .</p><p>Notes.</p><p>The differences between the sympatric E. pelkiae and E. jannei are too large to regard as sexual dimorphism.</p><p>Etymology.</p><p>The epithet pelkiae refers to Ms. Pelki Yangdon, the daughter of the fourth author.</p></div>	https://treatment.plazi.org/id/819F33E69E015A96978B52B692112C7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gittenberger, Edmund;Leda, Pema;Wangchuk, Jigme;Gyeltshen, Choki;Bjoern Stelbrink,	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
636A6BE75F255DBF8BD0CC9B5C347E4A.text	636A6BE75F255DBF8BD0CC9B5C347E4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Erhaia wangchuki Gittenberger, Sherub & Stelbrink 2017	<div><p>Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017 Figs 6, 7, 12</p><p>Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017a: 23 ("district Wangdue Phodrang, Gangchhu, 2883 m a.s.l.; 27°26'N, 90°11'E "). Gittenberger et al. 2017b: 43, fig. 28. Gittenberger et al. 2017c: 900, 903, figs 9, 10.</p><p>Material examined.</p><p>Holotype. Bhutan • District Wangdue Phodrang, Gangchhu, 2883 m a.s.l.; 27°26'N, 90°11'E; Jigme Wangchuk leg. 21.III.2015; shell; NBCB1013. Paratypes. 2 shells; same data as for holotype; NBCB1014. Additional specimens from the type locality: 23 shells and 88 specimens in ethanol 70%, 10 specimens in ethanol 97%, E. Gittenberger, Choki Gyeltshen &amp; Pema Leda leg. 22.X.2018; NBCB 1072.</p><p>Shell.</p><p>Shell conical, with 3- 3½ convex, broadly shouldered whorls, that are separated by a deep suture; a little higher than broad; pale yellowish grey with fine irregular growth lines and some dark brown periostracal ridges, one of which sometimes running from a slightly angled site of the apertural columellar border into the umbilicus. Peristome not reflected. Parietal, columellar and about half the adjoining basal apertural border strongly thickened by a whitish callus. Most specimens with a continuous peristome and a broad umbilical chink. Protoconch with faint spiral lirae; teleoconch without spiral sculpture.</p><p>Measurements.</p><p>(N = 124): H 1.6-2.1 mm, B 1.3-1.7 mm.</p><p>Ecology</p><p>(Fig. 12). See Gittenberger et al. (2017a) for data about the Gangzetem brooklet and its surroundings. The snails are very common on the pebbles and rocks in the open area near the road, next to the water powered prayer wheel.</p><p>DNA data</p><p>(Fig. 18). A single individual (GenBank acc. nos. KY798003 and MT237715, for 16S rRNA and COI, respectively) is genetically distinct from E. jannei (see data for that species) and showed genetic distances of 2.74% for 16S rRNA and 11.43% for COI compared to Erhaia sp. from China, Guangxi (GenBank acc. nos. KC832722 and KC832701).</p><p>Notes.</p><p>Only three relatively large shells form the type series of this species. Many more specimens, none of which exceed 1.7 mm in breadth and over 2.0 mm in height, were collected recently. This necessitated some adaptations in the description of the shells. Contrary to the original description, the shell should be described as higher than broad.</p><p>Erhaia in Nepal and northern India.</p><p>In their monograph on the aquatic molluscs of the Ganga River system Nesemann et al. (2007: 64-65) published short descriptions with drawings only of three Erhaia species from Nepal. We acquired photos of the holotypes of these nominal species, which are compared with the congeneric species from Bhutan and northern India.</p></div>	https://treatment.plazi.org/id/636A6BE75F255DBF8BD0CC9B5C347E4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gittenberger, Edmund;Leda, Pema;Wangchuk, Jigme;Gyeltshen, Choki;Bjoern Stelbrink,	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
BFB7EE714706502FB3A17F4E2A794F00.text	BFB7EE714706502FB3A17F4E2A794F00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Erhaia sugurensis Nesemann, Shah & Tachamo 2007	<div><p>Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007 Fig. 10</p><p>Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, in Nesemann et al. 2007: 65 ("Nepal, Central Zone, Lalitpur District, Godawari, upper reaches of Sugure Khola forrest stream, elevation of 1700 m a.s.l.").</p><p>Shell.</p><p>According to Nesemann et al. (2007: 65) the shells are 1.6-1.9 mm high, with 3½ whorls that are "not convex", the aperture is "rounded, widened and enlarged, the inner and outer lip is thickened". The species is said to differ most conspicuously by "the separation of the last half whorl from the shell".</p><p>Material examined</p><p>(photo only). Holotype. (NHMW 104172).</p><p>Discussion.</p><p>A flaring final half of the body whorl, though not as extreme as in the holotype of E. sugurensis, may also occur as an individual variation in E. nainitalensis and the width of the umbilical chink may vary, as is shown by Davis and Rao (1997: 277, figs 2A-F).</p><p>Notes.</p><p>Erhaia sugurensis occurs sympatrically with E. banepaensis at the type locality.</p></div>	https://treatment.plazi.org/id/BFB7EE714706502FB3A17F4E2A794F00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gittenberger, Edmund;Leda, Pema;Wangchuk, Jigme;Gyeltshen, Choki;Bjoern Stelbrink,	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
17AF80062CC55BF8B55476DE58D31D70.text	17AF80062CC55BF8B55476DE58D31D70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Erhaia banepaensis Nesemann & S. Sharma 2007	<div><p>Erhaia banepaensis Nesemann &amp; S. Sharma, 2007 Fig. 8</p><p>Erhaia banepaensis Nesemann &amp; S. Sharma, in Nesemann et al. 2007: 64 ("Nepal, Central Zone, Kavre District, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa"; the altitude is not indicated).</p><p>Shell.</p><p>The shells are described by Nesemann and Sharma (2007: 64) as 1.6-2.0 mm high, with 4- 4½ “convex” whorls, an aperture that is "ovate but not widened and not enlarged", with an inner lip that is "thin and fused to the body whorl"; it can be distinguished from the other Nepalese Erhaia species by the "conical and compact shape" and “convex” whorls (2007: 65).</p><p>Material examined</p><p>(photo only). Holotype. (NHMW 1033159).</p><p>Notes.</p><p>Nesemann and S. Sharma are mentioned as authors for this species, without specifying for what part of the text in Nesemann et al (2007) they have responsibility.</p><p>According to Nesemann et al. (2007: 64) E. banepaensis occurs sympatrically with E. chandeshwariensis at the shared type locality of these species. Erhaia banepaensis is supposed to be more widely distributed in Nepal between 1400 and 2085 m a.s.l., but the type series is restricted to the holotype (NHMW 103319 [not 1033159]) and a paratype (NHMW 103320). The location of the additional material is not indicated.</p></div>	https://treatment.plazi.org/id/17AF80062CC55BF8B55476DE58D31D70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gittenberger, Edmund;Leda, Pema;Wangchuk, Jigme;Gyeltshen, Choki;Bjoern Stelbrink,	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
E2D86FBA658857FC93211203D47458A8.text	E2D86FBA658857FC93211203D47458A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Erhaia nainitalensis Davis & Rao 1997	<div><p>Erhaia nainitalensis Davis &amp; Rao, 1997 Fig. 9</p><p>Erhaia nainitalensis Davis &amp; Rao, 1997: 276 ("India, Uttar Pradesh, Nainital District, Padampuiri"; " 29°23'N, 79°30'E ")</p><p>Erhaia chandeshwariensis Nesemann and S. Sharma, in Nesemann et al. 2007: 64, 78 fig. 4 ("Nepal, Central Zone, Kavre District, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa"; the altitude is not indicated).</p><p>Material examined</p><p>(photos only). Holotype and 4 paratypes of Erhaia chandeshwariensis (NHMW 103315 and 103316).</p><p>Discussion .</p><p>According to Davis and Rao (1997: 277, legends to figure 2) the holotype of E. nainitalensis is 2.28 mm high. However, elsewhere in the same article (Davis and Rao 1997: 279, table 1) the shell height of a single male animal is reported as 1.62 mm whereas 1.88-1.95-2.14 mm (minimum-mean-maximum) is indicated for five female snails. The difference in measurements between the single male and the five females might be indicative of sexual dimorphism. Shells of the species are also supposed to be “minute”, what is defined as 2.0 mm high or smaller (Davis and Rao 1997: 275, 289, table 5). We suppose that the dimensions of the holotype of E. chandeshwariensis, i.e. H 1.94 mm, fall within the range of the measurements of E. nainitalensis . Three of the four paratypes of E. chandeshwariensis (NHMW 103316/4) are c. 1.9 mm high, whereas the fourth shell is damaged, missing the apical whorls. Davis and Rao (1997: 278, fig. 3) figure a smooth columella for E. nainitalensis, but mention (p. 289) a "discernable glassy thickening of the columella".</p><p>Judging the nominal taxa on the basis of photographs of shells and additional data in the literature, we conclude that in general shape and apertural characters, i.e. a narrow umbilical chink, a virtually smooth columella, and a thickened outer and inner lip, the holotype of E. chandeshwariensis cannot be distinguished from the shells of E. nainitalensis that are figured by Davis and Rao (1997: 277, figs 2A-F, 280, figs 4A-D). The fact that Davis and Rao (1997: 276) indicate 3¾ -4 whorls for E. nainitalensis, whereas Nesemann and Sharma (2007: 64) mention 3½ whorls for E. chandeshwariensis, might be due to different counting methods.</p><p>Family Pomatiopsidae Stimpson, 1865</p></div>	https://treatment.plazi.org/id/E2D86FBA658857FC93211203D47458A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gittenberger, Edmund;Leda, Pema;Wangchuk, Jigme;Gyeltshen, Choki;Bjoern Stelbrink,	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
D99BC01B5F17555A8AFDB555B1CC6CB1.text	D99BC01B5F17555A8AFDB555B1CC6CB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricula Benson 1843	<div><p>Genus Tricula Benson, 1843</p><p>Type species by monotypy.</p><p>Melania (Tricula) montana Benson, 1843</p></div>	https://treatment.plazi.org/id/D99BC01B5F17555A8AFDB555B1CC6CB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gittenberger, Edmund;Leda, Pema;Wangchuk, Jigme;Gyeltshen, Choki;Bjoern Stelbrink,	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
65E57C37BD1F572AB522BC1491C34D65.text	65E57C37BD1F572AB522BC1491C34D65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricula montana (Benson 1843)	<div><p>Tricula montana (Benson, 1843) Figs 1, 16-17</p><p>Melania (Tricula) montana Benson, 1843: 467 ( “Bhimtal”, Nainital District, Uttarakhand, India; 1370 m a.s.l.). Lectotype in The Natural History Museum, London no. 1964426 (design. Davis et al. 1986: 428, fig. 3A).</p><p>Tricula montana; Preston 1915: 68. Davis et al. 1986: 428-433, figs 3-4 (shell), 4 (operculum), 5-8 (anatomy), 9-10 (radula). Nesemann et al. 2007: 62, 78 pl. 15, fig. 1.</p><p>Material examined.</p><p>Bhutan • District Lhuentse: Khardungchhu; 27°31'56"N, 91°12'19"E; 1634 m a.s.l.; J. Wangchuk leg. 28.IV.2017; 3 shells; NBCB 1061. Same data except for 27.III.2019; 8 specimens in ethanol 70%; NBCB 1064.</p><p>District Lhuentse: Jarkangchhu; 27°32'27"N, 91°12'25"E; 1333 m a.s.l.; J. Wangchuk leg. 28.IV.2017; 2 shells; NBCB 1063. Same data except for 27.III.2019; 7 specimens in ethanol 70%; NBCB 1066.</p><p>District Lhuentse: Songkhangchhu; 27°31'54"N, 91°11'17"E; 1152 m a.s.l.; J. Wangchuk leg. 27.III.2019; 3 specimens in ethanol 70%; NBCB 1068.</p><p>District Lhuentse: Fawan; 27°29'22"N, 91°10'57"E; 940 m a.s.l.; J. Wangchuk leg. 27.III.2019; 3 specimens in ethanol 70%; NBCB 1069.</p><p>District Mongar: Chhuburee; 27°15'41"N, 91°09 02"E; 818 m a.s.l.; J. Wangchuk leg. 3.V.2017; 2 shells; NBCB 1062. Same data except for 26.III.2019; 5 specimens in ethanol 70%; NBCB 1065.</p><p>District Mongar: Rekpalung; 27°19'34"N, 91°13'28"E; 885 m a.s.l.; J. Wangchuk leg. 27.III.2019; 3 specimens in ethanol 70%; NBCB 1070.</p><p>District Trongsa: Chendebji; 27°29'24"N, 90°20'18"E; 2631 m a.s.l., J. Wangchuk photographed 12.I.2018.</p><p>District Wangdue Phodrang: 40 km SSE of Wangdue Phodrang; 27°09'25"N, 90°04'05"E; 527 m a.s.l.; E. Gittenberger, Choki Gyeltshen &amp; Kezang Tobgay leg. 24.IX.2019; 23 shells; 23 specimens in ethanol 70%; 10 specimens in ethanol 97%; NBCB 1084.</p><p>District Zhemgang, Kekhar, 27°12'37"N, 90°46'28"E; 1540 m a.s.l., J. Wangchuk leg. photographed 17.I.2018.</p><p>Shell.</p><p>Shell slender conical, with up to c. 5 shouldered, moderately convex whorls, separated by an incised suture; with obsolete growth lines and poorly discernible dense spiral lirae. Pale yellowish grey, with a light brown apertural border when fully grown. Apex not flattened, often decollate. Aperture triangular with broadly rounded edges, its parietal side about double the length of the columellar side; palatal side straight, passing into the slightly curved basal border with a more strongly curved transitional part. Parietal border of the aperture attached, at least in the middle and not or only slightly protruding. Umbilicus closed or nearly so.</p><p>Measurements.</p><p>According to Davis et al. (1986: 431) the shell height of males and females combined (N = 10) is 3.32-3.72 mm. However, for the lectotype a larger shell height is indicated, i.e. 3.92 mm (Davis et al. 1986: 429, fig. 3A, 430). That shell is not even a relatively large specimen. Additional shells figured by Davis et al. (1986: 429, 430, fig. 3F, G, H, K, L) and printed at the same scale, are larger.</p><p>The shells that are known from Bhutan (N = 73) are relatively small, with 5- 5½ whorls measuring H 2.8-3.6 mm, B 1.3-1.7 mm.</p><p>Distribution.</p><p>(Fig. 1). According to Subba Rao (1989: 68) this species occurs in the Indian states of Himachal Pradesh ("Jhiri valley") and Uttarakhand (= Uttaranchal). Nesemann et al. (2007: 62) refers to it as widely distributed in the western and central Himalaya, in Nepal mainly at 1300-2100 m a.s.l. The records for Bhutan, at altitudes of 527-2631 m a.s.l., extend its range eastwards.</p><p>Habitat.</p><p>This species was found in Bhutan without accompanying Erhaia species mostly in densely vegetated, shaded areas with more or less overgrown springs and streamlets (Figs 1, 13 - 15). Davis et al. (1986: 427) describe a similar habitat for the Nainital District near the type locality of T. montana . The locality in the district of Wangdue Phodrang is an overgrown, dripping wet, vertical, rocky wall along the road.</p><p>The shells from Mongar, Chhuburee, and from Lhuentse, Jarkangchhu, are all decollate (Fig. 17), whereas shells from the other localities still have their apical whorls present (Fig. 16). This might be a consequence of unknown differences in water quality at those different localities.</p><p>DNA data.</p><p>Two snails from Chhuburee and two snails from Khardungchhu were sequenced. These specimens shared the same haplotype per population for both 16S rRNA (GenBank acc. nos. MT239080 and MT239079, for Chhuburee and Khardungchhu,) and COI (GenBank acc. nos. MT237718 and MT237717, for Chhuburee and Khardungchhu). The two populations differed genetically by 1.0% and 4.9% for 16S rRNA and COI, respectively. Because the monophyly of Tricula remains uncertain (see e.g., Liu et al. 2014), we compared these sequences with additional data available from GenBank. For 16S rRNA, the lowest genetic distances, i.e., 3.1% and 3.3%, were identified between snails from Chhuburee and Khardungchhu, respectively, and Tricula sp. from China, Hunan, Xiangxi, Fenghuang (GenBank acc. no. EU311736), and 3.3% and 3.5% between snails from Chhuburee and Khardungchhu, respectively, and T. ludongbini Davis &amp; Y.-H. Guo, 1986 from China, Yunnan, Panlong River, Hei Long Tan (GenBank acc. no. KC832717).</p><p>The genetic distances between T. montana from Chhuburee and Khardungchhu were considerably higher for COI, with 8.9% and 9.3%, respectively, between snails from Chhuburee and Khardungchhu and Tricula sp. from China, Sichuan, Dayi, Tian Gong Mia, Huang Ba (GenBank acc. no. AF253070), and Tricula hortensis Attwood &amp; Brown, 2003 from China (GenBank acc. no. JQ082621).</p><p>Notes.</p><p>The species was identified conchologically by using the data provided by Benson (1843), Davis et al. (1986) and Nesemann et al. (2007), taking also the distributional data (Nesemann et al. 2007: 62) into account.</p><p>Some species of Tricula may transmit schistosomes that could in principle infect humans and other mammals. No data in respect of this are known for T. montana .</p><p>See Davis et al. (1986) for a detailed account on this species, with data on shell morphology, anatomy of males and females, biogeography, and systematic relationships.</p></div>	https://treatment.plazi.org/id/65E57C37BD1F572AB522BC1491C34D65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gittenberger, Edmund;Leda, Pema;Wangchuk, Jigme;Gyeltshen, Choki;Bjoern Stelbrink,	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
