taxonID	type	description	language	source
7606A5B8245F52F7ADF39BA5978B68C1.taxon	distribution	Distribution. The genus Erhaia is mainly known from China, where it has been recorded with various species from the province of Yunnan in the west to the provinces of Hunan, Hubei and Fujian in the east (Davis et al. 1985; Davis and Kang 1995; Davis and Rao 1997; Wilke et al. 2000, 2001; Liu et al. 2014). One species was described from northern India (Davis and Rao 1997), three from Nepal (Nesemann et al. 2007) and one from Bhutan (Gittenberger et al. 2017 a). Here we deal with the systematics of only the species occurring in the southern Himalayan foothills in Bhutan, India, and Nepal.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
C16E201EF7AA5B7BB2E1D9D2C4050BD6.taxon	materials_examined	Material examined. Holotype. (Fig. 2) Bhutan • District Thimphu, c. 5 km E of Chhuzom, W of Geneykha; in brooklet with a prayer wheel along the road; 2750 m a. s. l.; 27 ° 18 ' 43 " N, 89 ° 36 ' 10 " E; E. Gittenberger, Choki Gyeltshen & Pema Leda leg. 25. X. 2018; NBCB 1057. Paratypes. (Fig. 3) 6 shells and 2 animals in ethanol 70 %; same data as for holotype; NBCB 1058.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
C16E201EF7AA5B7BB2E1D9D2C4050BD6.taxon	diagnosis	Diagnosis. Shell large for the genus (H> 2 mm), ovoid, with a relatively large aperture.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
C16E201EF7AA5B7BB2E1D9D2C4050BD6.taxon	description	Measurements. (N = 9): H 2.2 - 2.4 mm, B 1.5 - 1.6 mm. Holotype 2.2 x 1.6 mm.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
C16E201EF7AA5B7BB2E1D9D2C4050BD6.taxon	etymology	Etymology. The epithet jannei refers to Mr Janne Clewing, the son of the last author.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
819F33E69E015A96978B52B692112C7D.taxon	materials_examined	Material examined. Holotype. (Fig. 4) Bhutan • District Thimphu, c. 5 km E of Chhuzom, W of Geneykha; in brooklet with a prayer wheel along the road; 2750 m a. s. l.; 27 ° 18 ' 43 " N, 89 ° 36 ' 10 " E; E. Gittenberger, Choki Gyeltshen & Pema Leda leg. 25. X. 2018; NBCB 1059. Paratype. (Fig. 5) 1 shell; same data as for holotype; NBCB 1060.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
819F33E69E015A96978B52B692112C7D.taxon	diagnosis	Diagnosis. Shell with a partly reflected peristome, teleoconch with spiral lirae.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
819F33E69E015A96978B52B692112C7D.taxon	description	Measurements. (N = 2): holotype and paratype H 1.9 mm, B 1.1 mm.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
819F33E69E015A96978B52B692112C7D.taxon	etymology	Etymology. The epithet pelkiae refers to Ms. Pelki Yangdon, the daughter of the fourth author.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
636A6BE75F255DBF8BD0CC9B5C347E4A.taxon	materials_examined	Material examined. Holotype. Bhutan • District Wangdue Phodrang, Gangchhu, 2883 m a. s. l.; 27 ° 26 ' N, 90 ° 11 ' E; Jigme Wangchuk leg. 21. III. 2015; shell; NBCB 1013. Paratypes. 2 shells; same data as for holotype; NBCB 1014. Additional specimens from the type locality: 23 shells and 88 specimens in ethanol 70 %, 10 specimens in ethanol 97 %, E. Gittenberger, Choki Gyeltshen & Pema Leda leg. 22. X. 2018; NBCB 1072.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
636A6BE75F255DBF8BD0CC9B5C347E4A.taxon	description	Measurements. (N = 124): H 1.6 - 2.1 mm, B 1.3 - 1.7 mm.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
BFB7EE714706502FB3A17F4E2A794F00.taxon	materials_examined	Material examined (photo only). Holotype. (NHMW 104172).	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
BFB7EE714706502FB3A17F4E2A794F00.taxon	discussion	Discussion. A flaring final half of the body whorl, though not as extreme as in the holotype of E. sugurensis, may also occur as an individual variation in E. nainitalensis and the width of the umbilical chink may vary, as is shown by Davis and Rao (1997: 277, figs 2 A-F).	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
17AF80062CC55BF8B55476DE58D31D70.taxon	materials_examined	Material examined (photo only). Holotype. (NHMW 1033159).	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
E2D86FBA658857FC93211203D47458A8.taxon	materials_examined	Material examined (photos only). Holotype and 4 paratypes of Erhaia chandeshwariensis (NHMW 103315 and 103316).	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
E2D86FBA658857FC93211203D47458A8.taxon	discussion	Discussion. According to Davis and Rao (1997: 277, legends to figure 2) the holotype of E. nainitalensis is 2.28 mm high. However, elsewhere in the same article (Davis and Rao 1997: 279, table 1) the shell height of a single male animal is reported as 1.62 mm whereas 1.88 - 1.95 - 2.14 mm (minimum-mean-maximum) is indicated for five female snails. The difference in measurements between the single male and the five females might be indicative of sexual dimorphism. Shells of the species are also supposed to be " minute ", what is defined as 2.0 mm high or smaller (Davis and Rao 1997: 275, 289, table 5). We suppose that the dimensions of the holotype of E. chandeshwariensis, i. e. H 1.94 mm, fall within the range of the measurements of E. nainitalensis. Three of the four paratypes of E. chandeshwariensis (NHMW 103316 / 4) are c. 1.9 mm high, whereas the fourth shell is damaged, missing the apical whorls. Davis and Rao (1997: 278, fig. 3) figure a smooth columella for E. nainitalensis, but mention (p. 289) a " discernable glassy thickening of the columella ". Judging the nominal taxa on the basis of photographs of shells and additional data in the literature, we conclude that in general shape and apertural characters, i. e. a narrow umbilical chink, a virtually smooth columella, and a thickened outer and inner lip, the holotype of E. chandeshwariensis cannot be distinguished from the shells of E. nainitalensis that are figured by Davis and Rao (1997: 277, figs 2 A-F, 280, figs 4 A-D). The fact that Davis and Rao (1997: 276) indicate 33 / 4 - 4 whorls for E. nainitalensis, whereas Nesemann and Sharma (2007: 64) mention 31 / 2 whorls for E. chandeshwariensis, might be due to different counting methods.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
65E57C37BD1F572AB522BC1491C34D65.taxon	materials_examined	Material examined. Bhutan • District Lhuentse: Khardungchhu; 27 ° 31 ' 56 " N, 91 ° 12 ' 19 " E; 1634 m a. s. l.; J. Wangchuk leg. 28. IV. 2017; 3 shells; NBCB 1061. Same data except for 27. III. 2019; 8 specimens in ethanol 70 %; NBCB 1064. District Lhuentse: Jarkangchhu; 27 ° 32 ' 27 " N, 91 ° 12 ' 25 " E; 1333 m a. s. l.; J. Wangchuk leg. 28. IV. 2017; 2 shells; NBCB 1063. Same data except for 27. III. 2019; 7 specimens in ethanol 70 %; NBCB 1066. District Lhuentse: Songkhangchhu; 27 ° 31 ' 54 " N, 91 ° 11 ' 17 " E; 1152 m a. s. l.; J. Wangchuk leg. 27. III. 2019; 3 specimens in ethanol 70 %; NBCB 1068. District Lhuentse: Fawan; 27 ° 29 ' 22 " N, 91 ° 10 ' 57 " E; 940 m a. s. l.; J. Wangchuk leg. 27. III. 2019; 3 specimens in ethanol 70 %; NBCB 1069. District Mongar: Chhuburee; 27 ° 15 ' 41 " N, 91 ° 09 02 " E; 818 m a. s. l.; J. Wangchuk leg. 3. V. 2017; 2 shells; NBCB 1062. Same data except for 26. III. 2019; 5 specimens in ethanol 70 %; NBCB 1065. District Mongar: Rekpalung; 27 ° 19 ' 34 " N, 91 ° 13 ' 28 " E; 885 m a. s. l.; J. Wangchuk leg. 27. III. 2019; 3 specimens in ethanol 70 %; NBCB 1070. District Trongsa: Chendebji; 27 ° 29 ' 24 " N, 90 ° 20 ' 18 " E; 2631 m a. s. l., J. Wangchuk photographed 12. I. 2018. District Wangdue Phodrang: 40 km SSE of Wangdue Phodrang; 27 ° 09 ' 25 " N, 90 ° 04 ' 05 " E; 527 m a. s. l.; E. Gittenberger, Choki Gyeltshen & Kezang Tobgay leg. 24. IX. 2019; 23 shells; 23 specimens in ethanol 70 %; 10 specimens in ethanol 97 %; NBCB 1084. District Zhemgang, Kekhar, 27 ° 12 ' 37 " N, 90 ° 46 ' 28 " E; 1540 m a. s. l., J. Wangchuk leg. photographed 17. I. 2018.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
65E57C37BD1F572AB522BC1491C34D65.taxon	description	Measurements. According to Davis et al. (1986: 431) the shell height of males and females combined (N = 10) is 3.32 - 3.72 mm. However, for the lectotype a larger shell height is indicated, i. e. 3.92 mm (Davis et al. 1986: 429, fig. 3 A, 430). That shell is not even a relatively large specimen. Additional shells figured by Davis et al. (1986: 429, 430, fig. 3 F, G, H, K, L) and printed at the same scale, are larger. The shells that are known from Bhutan (N = 73) are relatively small, with 5 - 51 / 2 whorls measuring H 2.8 - 3.6 mm, B 1.3 - 1.7 mm.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
65E57C37BD1F572AB522BC1491C34D65.taxon	distribution	Distribution. (Fig. 1). According to Subba Rao (1989: 68) this species occurs in the Indian states of Himachal Pradesh (" Jhiri valley ") and Uttarakhand (= Uttaranchal). Nesemann et al. (2007: 62) refers to it as widely distributed in the western and central Himalaya, in Nepal mainly at 1300 - 2100 m a. s. l. The records for Bhutan, at altitudes of 527 - 2631 m a. s. l., extend its range eastwards.	en	Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki, Bjoern Stelbrink, (2020): The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya. ZooKeys 929: 1-17, DOI: http://dx.doi.org/10.3897/zookeys.929.49987, URL: http://dx.doi.org/10.3897/zookeys.929.49987
