taxonID	type	description	language	source
1312774BFFD1FFE3FF04FF70FCA9FBE8.taxon	materials_examined	Type species: Denticulobasis garrisoni sp. nov., by present designation.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD1FFE3FF04FF70FCA9FBE8.taxon	etymology	Etymology: From the Latin denticulus – meaning diminutive of tooth and – basis, a Greek feminine noun for pedestal or foundation; a common suffix used in Coenagrionidae. The name Denticulobasis is an allusion to the presence of a plate with denticles on female sternum S 8.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD1FFE3FF04FF70FCA9FBE8.taxon	distribution	Distribution: Brazil and Peru (Fig. 101). Generic characterization. Medium size damselflies (HW 18.2 – 19.2 mm). Top of head brown with blue postocular spots. Thorax dominantly yellow, with three longitudinal pale blue stripes: antehumeral, metepisternal, and metepimeral. S 1 – 7 dorsally brown, laterally yellow, S 8 – 10 dorso-laterally yellow. Frons rounded. Hind prothoracic lobe in males with very small lateral lobes, median lobe well-developed, projected caudally and two-lipped (Figs 4 – 6); females with well-developed lateral and median lobes (Figs 8 – 10), the latter single-lipped. Male mesepisternum with an antero-medial horn with a basal smooth portion and a distal portion ridged as a screw. Female mesepisternum with an antero-medial mammiliform tubercle. Metafemoral spines on distal 1 / 2 as long as width of femur (Fig. 3 a); metatibial spurs shorter than intervening spaces (Fig. 3 b). Supplementary tooth of tarsal claw well-developed (Figs 3 c, d). Wings petiolated distal to Ac for a distance as long as or twice as long as Ac length (Fig. 3 e). R 3 in FW and HW originating closer to Px 6 (Fig. 3 e). Penis (Figs. 1) distal segment expanded laterally along distal 1 / 3 forming a large lateral terminal lobe and with two lateral lobes on each side. No terminal fold. Internal fold with a filiform process directed disto-laterally (Fig. 1). Male cercus with a dorso-lateral membranous depression at base (Figs. 12 – 13) and no ventral process. Cercus in lateral view subequal to S 10 (excluding distal lobe). Female sternum S 8 with a denticulated vulvar plate (Figs. 18, 20, 22). Ovipositor extending to about level of tip of cercus (Fig. 11). Ventral border of lateral valve of ovipositor with one row of denticles each one subrectangular, directed ventro-posteriorly and associated with a robust short truncated seta (Fig. 2 a).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD1FFE2FF04FB40FC50FE41.taxon	etymology	Etymology: Named after the Ariken indian people who inhabited the municipality of Ariquemes, Rondônia, and whose descendants live now scattered throughout the region.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD1FFE2FF04FB40FC50FE41.taxon	materials_examined	Type specimen: Holotype (Ƥ). BRAZIL, Rondônia State: Rio Pardo and tributaries, ca. 13 Km NW of Fazenda Rancho Grande (62 Km SW of Ariquemes 10 º 50´S, 63 º 7´W), 187 m., 5 – 10 / XI / 1989, R. W. Garrison leg. (MNRJ).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD1FFE2FF04FB40FC50FE41.taxon	description	Description. Female holotype Head. Labium white, base of mandibles and genae yellowish white, anteclypeus and labrum grey, the latter surrounded by yellowish white. Postclypeus, antefrons, and top of head brown, the latter with rounded pale blue postocular spots. Thorax. Prothorax: Propleuron and anterior lobe light blue; remaining parts orange yellow. Pterothorax orange yellow with blue stripes on mesepisternum (antehumeral), metepisternum, and metepimeron, the latter occupying entire lateral part of sclerite. Legs yellow. Wings hyaline, venation brown, pterostigma brown surrounded by a yellow line. Px in FW 12, HW 11; R 3 arising near Px 6 in FW, and 5 in HW. Anal vein separating from wing margin distally to Ac for a distance equal to ½ of Ac length in FW and ¼ in HW. Abdomen. S 1 – 2 dorsally dark brown, laterally light blue. S 3 dorsally dark brown, dorso-laterally blue, ventrally yellow. S 4 – 6 dorsally dark brown, laterally yellow; S 7 – 8 dorso-laterally yellowish orange, ventrally whitish yellow, S 9 – 10 orange yellow. A black ring occupying distal 1 / 7 of S 3 – 6. Structural characters. Hind lobe of prothorax (Fig. 8) with lateral lobes well-developed. Median lobe subrectangular, single-lipped with a longitudinal depression at mid-line. Mesostigmal plate (Fig. 8) with anterior and posterior margins convergent and connected to the one on other side. Mid-dorsal carina with cleft at level of mesepisternal tubercle. Mesepisternal tubercles (Fig. 8) low and mammiliform. Ovipositor reaching to level of tip of cercus (as in Fig. 11). Vulvar plate (Fig. 18) with a short neck and straight distal margin with six denticles of regular size arranged side by side (Fig. 18). Dimensions (mm). Abdomen 32.10; HW 19.45.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD1FFE2FF04FB40FC50FE41.taxon	discussion	Remarks. General color pattern, and especially presence of a vulvar plate, leaves no doubt that D. ariken is a Denticulobasis. It differs from the females of the other two congeners by the convergent margins of mesostigmal plate and the morphology of vulvar plate with distal border straight (curved in D. dunklei) and denticles arranged regularly side by side (in three groups in D. garrisoni).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD3FFE0FF04FB3BFD5EFD92.taxon	etymology	Etymology: Named after my colleague Sidney W. Dunkle, who collected the specimens in Peru, in recognition of his important contributions to Odonatology.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD3FFE0FF04FB3BFD5EFD92.taxon	materials_examined	Type specimens: Holotype (3) and allotype (Ƥ). PERU, Loreto Department. Explorama Lodge, 50 miles NE of Iquitos on Amazon River at the junction with Yanomano River (3 º 21´59 ´´ S, 72 º 47´56 ´´ W), 24 - VIII- 1989, S. W. Dunkley leg. (FSCA).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD3FFE0FF04FB3BFD5EFD92.taxon	description	Description. Male holotype. Head. Labium whitish yellow. Labrum, base of mandibles, and genae greyish brown. Anteclypeus, postclypeus, frons, antennae, and top of head brown with large blue postocular spots. Rear of head whitish yellow. Thorax. Prothorax: Propleuron and anterior lobe light blue. Pronotum light brown. Pterothorax: mesopleuron dominantly yellowish brown. Horns and area between them black, except at their latero-basal region, which is bluish. Metepisternum yellowish brown and metepimeron yellowish white. Three blue stripes; antehumeral, metepisternal, and a poorly defined metepimeral. Legs yellow. Wings hyaline, venation brown, pterostigma brownish yellow surrounded by yellow. Px in FW 12 – 13 in HW 11. R 3 arising near Px 6 in FW and 5 in HW. Petiolation distal to Ac by a distance 1.5 times length of Ac in FW and equal to it in HW. Abdomen. S 1 and S 2 dorsally brown, laterally light blue. S 3 – 6 dorsally brown, laterally yellow with black ring at distal 1 / 10. S 7 dorsally reddish brown, laterally yellow with black ring at proximal ¼. S 8 – 10 dorsally brownish yellow, laterally yellow. Cercus and paraproct brownish yellow. Structural characters. Hind prothoracic lobe (Fig. 4) with lateral lobes minute, median lobe projecting caudally, subquadrangular and two-lipped; ventral lip continuous, and dorsal lip reduced to two flaps separated by an incision at mid-line. Mesepisternal horns in dorsal view (Fig. 4) strongly diverging, each tapering to a blunt tip, mostly smooth with a transversely ridged portion at distal 1 / 5. Cercus in dorsal view slightly wider at apex than at base (excluding distal lobe) (Fig. 12) with a whitish shallow subtriangular membranous depression at base (Fig. 12). In ventro-posterior view (Fig. 16) cercus is spoon-shaped, and continues apically as a distal lobe. A ventral transverse digitiform structure present at limit between main portion and distal lobe of cercus (Fig. 16) which is not visible in dorsal view (Fig. 12). Penis (Fig. 87) as described for the genus. Dimensions (mm). Abdomen: 32.9; HW 18.82.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD2FFE6FF04FA06FB2AFF59.taxon	etymology	Etymology: Named after my colleague and friend Rosser W. Garrison, who collected the types in Rondônia and has helped me in the form of specimens and advice for over 20 years.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD2FFE6FF04FA06FB2AFF59.taxon	materials_examined	Type specimen: Holotype (3) and allotype (Ƥ). BRAZIL, Rondônia State: Rio Pardo and tributaries, ca 13 Km NW of Fazenda Rancho Grande, 62 Km SW of Ariquemes (10 º 50´S, 63 º 7´W), 187 m.; 5 – 10 November 1989, R. W. Garrison leg. (taken at ground level deep in forest in the company of Megapodagrion megalopus Selys, rare) (MNRJ). Paratypes: 2 3, same data as holotype (RWG); 1 3, same data as holotype (ABMM).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD2FFE6FF04FA06FB2AFF59.taxon	description	Description. Male holotype. Head. Labium yellowish white. Labrum, base of mandibles, genae, and anteclypeus whitish yellow. Postclypeus olive. Antefrons and top of head brown with blue postocular spots (Fig. 101 d). Thorax. Prothorax: laterally blue, dorsally orange yellow, except for the anterior lobe, which has an orange yellow spot surrounded by blue. Pterothorax: meso and metapleuron orange yellow with blue, antehumeral, mesepisternal, and metepimeral stripes. Legs yellow, wings hyaline, venation and pterostigma brown. Px in FW 11 – 12 in HW 10 – 11, R 3 arising near Px 6 in FW and near 5 in HW. Petiolation distal to Ac by a distance ½ of its length in both wings. Abdomen. S 1, S 2 and proximal part of S 3 laterally blue, dorsally yellow; remaining part of S 3, S 4 – 6 laterally yellow, dorsally dark brown; S 7 reddish brown with a black ring at mid-length, S 8 – 9 dorso-laterally yellowish orange, cercus orange yellow, distal lobe whitish yellow. Structural characters. Hind prothoracic lobe (Fig. 6) with a small lateral lobe, median lobe projecting caudally, semicircular, two-lipped; ventral lip continuous, dorsal lip disappearing antero-laterally. Mesepisternal horns (Fig. 6) with a basal and a distal portion, each one occupying about 1 / 2 horn's length; basal portion smooth and directed antero-laterally, distal portion transversally ridged like a screw and directed postero-laterally. Cercus in dorsal view (Fig. 13) subrectangular, about as wide at base as at the apex (excluding distal lobe), with a white, approximately rounded, basal membranous depression. Medial borders with a basal convexity and a distal concavity; a large whitish distal lobe present. After relaxing and spreading paraprocts, a digitiform transverse process directed medially is clearly visible in ventro-posterior and dorsal views (Figs 13, 17). Penis (Fig. 90) as in D. dunklei. Dimensions (mm). Abdomen 27.8 – 31.4 (mean 30.2); HW 18.2 - 19 - 4 (mean 18.7).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD4FFE5FF04FB4BFE04FE83.taxon	etymology	Etymology: From the Latin tuberculum – a tubercle and – basis, a Greek feminine noun for pedestal or foundation; a common suffix used in Coenagrionidae. The name Tuberculobasis refers to the presence of paired tubercles on mesepisterna of males and most females.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD4FFE5FF04FB4BFE04FE83.taxon	distribution	Distribution. South America (Brazil, Colombia, Peru, Suriname, Venezuela) (Fig. 101). Generic characterization. Small to medium size damselflies (HW 15.5 – 22.0 mm). Top of head black or brown (reddish yellow in T. mammilaris), with blue postocular spots (sometimes absent in T. mammilaris). Thorax usually with three longitudinal pale blue or silver stripes: antehumeral, metepisternal, and sometimes metepimeral. S 1 – 6 variously brownish or yellowish, S 7 – 10 red, yellow or orange brown. Frons rounded. Hind prothoracic lobe in males with median lobe well-developed projected caudally and two-lipped (Figs 25 – 45), in females single-lipped (Figs 47 – 56). Mesepisterna with an antero-medial pair of tubercles present in all males and 64 % of known females. Cercus in lateral view much shorter than S 10 in both sexes. Metafemoral spines on distal 1 / 2 as long as width of femur (Fig. 24 a); metatibial spurs shorter than intervening spaces (Fig. 24 b). Supplementary tooth of tarsal claw well-developed (Figs 24 c, d). Wings (Fig. 24 e) petiolated distally to Ac for a distance as long as Ac length (88.25 %) or 1 / 2 as long (Fig. 24 e). R 3 in FW originating closer to Px 5 – 6 (more frequently at 6) in HW closer to Px 4 – 6 (more frequently at 5). Penis distal segment with distal 1 / 3 laterally expanded into a large lateral terminal lobe and with two lateral lobes on each side (Figs 89 – 95). No terminal fold. Internal fold with a filiform process directed disto-laterally (Figs. 89 – 93). Male cercus domeshaped (Fig. 68), the distal part sharply decumbent forming a ventral process (ventro-medial in T. arara and T. costalimai), which is perpendicular to the long axis of cercus. Female sternum S 8 with a vulvar spine (except in T. cardinalis, see under that species). Ovipositor extending to about level of tip of cercus (Figs 95 – 99). Ventral border of lateral valve of ovipositor with one row of subtriangular denticles fine tipped directed ventro-posteriorly associated with a long seta tapering into a fine tip (Fig. 2 b).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD4FFE5FF04FB4BFE04FE83.taxon	description	Larvae unknown.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD6FFE4FF04FBFBFA3BF874.taxon	etymology	Etymology: The name refers to the Arara indian people who inhabit the municipality of Ji-Paraná.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD6FFE4FF04FBFBFA3BF874.taxon	materials_examined	Type specimen: Holotype (3), BRAZIL, Rondônia State, Ji-Paraná (8 º 03´N, 62 º 52´W), 25 - I- 1961, Machado & Pereira leg. (inside forest, away from water) (ABMM).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD6FFE4FF04FBFBFA3BF874.taxon	description	Description. Male holotype. Head. Missing. Thorax. Prothorax: Anterior lobe grey, median lobe brown with brown-yellowish lateral areas. Posterior lobe reddish brown. Pterothorax: mesopleuron and metepisternum ventrally brownish yellow, dorsally brown with a poorly defined antehumeral grey stripe and a well defined metepisternal stripe continuing on to mesinfrepisternum. Legs yellow, wings hyaline, venation brown, pterostigma reddish brown. Px in FW 11 – 12 in HW 10; R 3 in FW originating near Px 6, in HW near Px 4 – 5. Petiolation distal to Ac by a distance twice or 1.5 times the length of Ac in FW and 1.5 times in HW. Abdomen. S 1 – 2 dorsally brown, laterally whitish blue. S 3 – 6 dorso-laterally dark brown with a pre-apical lateral yellow marking scarcely visible. S 7 – 10 and cercus yellowish orange. Paraproct brown. Structural characters. Hind prothoracic lobe (Figs 25, 26) without lateral lobes, median lobe subrectangular, dorsal lip fused medially with ventral lip resulting a median depression (Fig. 25). Mesepisternal tubercles (Figs 25, 26) high (2.36 mm) and divergent in dorsal view (Fig. 25) with the apex subtruncated (Fig. 25). Cercus (Figs. 57 – 59) shorter than S 10, about 1 / 3 shorter than paraprocts, with short ventro-medial subtriangular process (Figs 58 – 59). Penis typical for Tuberculobasis (as in Fig. 89). Dimensions (mm). Abdomen 31.2; HW 19.2	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFD6FFE4FF04FBFBFA3BF874.taxon	discussion	Remarks. By having high metepisternal tubercles with their bases adjacent to mid-dorsal carina, paraproct much longer than cercus, and no lateral lobes in hind prothoracic lobe, T. arara belongs in the costalimai species group together with T. guarani, T. inversa, T. costalimai, and T. williamsoni. It is closer to	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC9FFFBFF04FEBEFE37FB26.taxon	distribution	Distribution. PERU, Loreto Department.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC9FFFBFF04FEBEFE37FB26.taxon	discussion	Remarks. T. cardinalis was described by Fraser (1946) under Oxyagrion based on one female specimen from Peru. Costa (1978) suggested that the species might belong in Acanthagrion due to the presence of thoracic stripes and De Marmels (1984 a) provided a partial redescription of the holotype, including drawings and photographs. Von Ellenrieder & Garrison (2007) examined the holotype in BMNH and concluded that the species is not an Oxyagrion and should be assigned to Leptobasis. They provided drawings of its main diagnostic characters, some of which are reproduced here (Figs 47, 95). The presence of mound-like mesepisternal tubercles and ovipositor not surpassing tip of cercus (Fig. 95), plus wing petiolation subequal to Ac justifies its transfer to Tuberculobasis. Von Ellenrieder & Garrison (2007) found some discrepancies between Fraser's description and the holotype that they regarded as unintentional mistakes by Fraser (1946). The most significant discrepancy was the presence of a vulvar spine mentioned by Fraser (1946) but missing in the holotype.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC9FFFBFF04FEBEFE37FB26.taxon	description	Male. Unknown.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC9FFFAFF04FB3DFA3BFE80.taxon	distribution	Distribution. BRAZIL, States of Rio de Janeiro, Minas Gerais, and Espírito Santo.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC9FFFAFF04FB3DFA3BFE80.taxon	discussion	Remarks. T. costalimai can be easily identified by its large size (abdomen 32 – 34; HW 19 – 20), male mesepisternal tubercle adjacent to mid-dorsal carina (Fig. 27), and ventral process of male cercus curved and directed ventro-medially (Fig. 62). The description of male and female by Santos (1957) is adequate, and I provide here new illustrations of cerci (Figs 60 – 62) and thorax (Figs 27 – 28, 101 a). Santos (1957) stated that female thorax does not have the dorsal prominences described for male. Although considerably smaller than in males, tubercles could be detected in all 20 females I examined including the allotype and eight specimens of the type series. In ten specimens (50 %) tubercles were well-developed, in six (30 %) very small, and in four (20 %) poorly-defined. Since in all specimens of the female type series examined by Santos (1957) mesepisternal tubercles could be distinguished it becomes evident that he overlooked these tubercles, certainly much less conspicuous than the " prominences " he had described in the males. In seven females (35 %) tubercles were connected with mid-dorsal carina and in 15 (75 %) posterior margin of mesostigmal plates curved back and penetrated between tubercle of mesostigmal plate and mesepisternal tubercle (Fig. 48).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC8FFFAFF04FE98FF54F9E2.taxon	etymology	Etymology: Named after the late Dr. Dirk Cornelis Geijskes, collector of the type specimen, in recognition for his enormous contribution to the knowledge of Neotropical Odonata.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC8FFFAFF04FE98FF54F9E2.taxon	materials_examined	Type specimens: Holotype (3). SURINAME, Republiek, Lelydorp (5 º 22´N, 57 º 30´W) 15 - IV- 1938, D. K. Geijskes leg. (MNRJ).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC8FFFAFF04FE98FF54F9E2.taxon	description	Description. Male holotype. Head. Labium yellow. Face green. Antefrons and top of head black with blue postocular spots. Traces of a yellow spot in front of postocular spot and a transverse yellow band behind it. Rear of head yellow. Thorax. Prothorax: Anterior lobe dark metallic green, median and posterior lobes brown with areas metallic copper and gold. Pterothorax: mesepisternum metallic copper with an argentous longitudinal stripe surrounded by metallic gold. Mesepisternal tubercles and mesepimeron metallic gold. Metapleuron brown with two broad argentous stripes on metepisternum and metepimeron. Legs yellow, tarsi brownish, wings hyaline, venation and pterostigma yellow. Px in FW 11; in HW 10. R 3 originating near Px 5 in FW and 4 in HW. Petiolation in FW distal to Ac by a distance equal to Ac length in FW and twice its length in HW. Abdomen. S 1 – 2 brown with a wide argentous lateral stripe. S 3 – 7 missing, S 9 – 10 reddish brown, cercus brown. Paraproct brown with apex dark. Structural characters. Hind prothoracic lobe (Figs 35, 36) with lateral lobes well-developed, median lobe projected posteriorly, semicircular, two-lipped; dorsal lip connected medially with ventral one by a small bridge (Fig. 75). Mesepisternal tubercle (Figs 35, 36) low (0.82 mm), in dorsal view elliptical. Hind margin of S 10 slightly elevated (Fig. 77). Cercus in lateral view (Fig. 76) with a slender tapering curved ventral process ending in a small hook-like tip. Penis as described for genus, but filiform process extending beyond lateral border (Fig. 90). Dimensions (mm). HW 19.0 Female. Unknown	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC8FFFAFF04FE98FF54F9E2.taxon	discussion	Remarks. This is the second species of Tuberculobasis recorded from Suriname after T. mammilaris (BELLE, 2002). By its cercus subequal to paraproct, low and mammiliform mesepisternal tubercles with bases well separated from mid-dorsal carina, and well-developed lateral lobes on prothoracic hind margin, it belongs in T. mammilaris species group. It differs from the other four species of this group by characters given in the key. Thoracic color with metallic luster and S 10 elevated dorso-distally are unique characters for this species.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC8FFFEFF04F979FC8CFCE5.taxon	etymology	Etymology: The name refers to the Guarani indian people who still inhabit various areas of the State of São Paulo.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC8FFFEFF04F979FC8CFCE5.taxon	materials_examined	Type specimens: Holotype (3). BRAZIL, São Paulo State, Ibitinga (21 º 45´S, 48 º 49´W), Fazenda Itaguaré, 17 - VI- 1965, K. Lenko leg. (ABMM).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC8FFFEFF04F979FC8CFCE5.taxon	description	Description. Male holotype. Head. Labium whitish yellow; labrum dark green; antefrons and top of head dark brown with rounded pale postocular spots. Thorax. Prothorax yellowish brown except for dark brown anterior lobe. Thorax, pterothorax: mesopleuron brown; metapleuron yellowish. Poorly defined grey stripes at posterior part of mesepisternum and metepisternum. Legs yellowish, wings hyaline, venation and pterostigma brown, Px in FW 11, in HW 10. R 3 arising near Px 5 in FW, near Px 4 in HW. Petiolation distal to Ac by a distance equal to Ac length in both wings. Abdomen. S 1 – 2 dorsally brownish yellow ventrally yellowish with a grey dorso-lateral stripe. S 3 – 6 dorsally brown, laterally yellow, with a dark brown ring at distal 1 / 7. S 70 – 10 yellowish orange. Structural characters. Hind prothoracic lobe with very small lateral lobes (Figs 29, 30), median lobe projecting caudally, semicircular, and two-lipped; both lips continuous (Fig. 29). Mesepisternal tubercles (Figs 29, 30) very high (2.83 mm), conical, with the apex directed laterally and bases adjacent to mid-dorsal carina (Fig. 29). Cercus 1 / 3 shorter than paraproct (Fig. 64) in posterior view (Fig. 65) with ventral process subtriangular. Penis as usual for the genus (as in Fig. 1). Dimensions (mm). Abdomen 32.5; HW 19.8. Female. Unknown.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC8FFFEFF04F979FC8CFCE5.taxon	discussion	Remarks. Tuberculobasis guarani has the highest mesepisternal tubercle within the genus (Fig. 30). It belongs in the costalimai species group and it is close to T. arara. It shares with this species the presence of subtriangular short ventral process of cercus but can be separated from it by characters given in the key. Together with T. costalimai, T. guarani is the only Tuberculobasis so far reported from SE Brazil, and the first species of the genus recorded for the State of São Paulo.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCCFFFDFF04F913FE9BFBF4.taxon	distribution	Distribution. Brazil, Amazonas, Pará, and Rondônia States. Peru, Loreto Department.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCCFFFDFF04F913FE9BFBF4.taxon	discussion	Remarks. Tuberculobasis inversa was described by Selys (1876) based on a single male from Santarém, Pará, Brazil. Santos (1961) described and illustrated both sexes of Leptobasis tuberculata from Porto Velho, Rondônia, Brazil but after examining the holotype of T. inversa synonymized L. tuberculata with L. inversa (Santos, 1965). The holotype lacks head and right wings, but color is well preserved. The descriptions made by Selys (1876) and Santos (1961) are adequate. Until recently the only illustrations known for females specimen of T. inversa were those provided by Santos (1961) under L. tuberculata. I had the opportunity to study and provide an illustration of the thoracic structures of a specimen from Peru that agreed well with the specimen described by Santos (1961) especially on the unique shape of the hind prothoracic lobe (Fig. 49). R. W. Garrison identified as T. inversa a male in his collection found in the same month and at the same place where the female illustrated herein had been collected and suggested, and I agree, that it is the female of T. inversa. However, the Peruvian specimen has a pair of small but very evident mesepisternal tubercles that are apparently lacking in the Brazilian specimen described by Santos. I believe these small tubercle might have been overlooked by Santos (1961) as he did with the tubercles of the females of T. costalimai (see Remarks under T. costalimai). Besides, by its male character T. inversa belongs in the costalimai group in which the known females (T. costalimai, T. williamsoni) have tubercles, whereas the known females (T. mammilaris, T. tirio and T. yanomami) of the mammilaris group the known females (T. mammilaris, T. tirio and T. yanomami) have no tubercles. In the costalimai species group T. inversa can be easily separated from the other species by its subparallel mesepisternal tubercles, with anterior borders connected to mesostigmal plate by a curved plate (Figs. 31, 32).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCFFFFCFF04FB4BFB27FD24.taxon	etymology	Etymology: The name refers to the Karitiana indian people who inhabit the municipality of Porto Velho where the holotype was collected.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCFFFFCFF04FB4BFB27FD24.taxon	materials_examined	Type specimens: Holotype (3): BRAZIL, Rondônia State, Porto Velho (8 º 46´S, 63 º 54´W), III 1963, W. C. A. Bokermann leg. (in a forest creek) (ABMM).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCFFFFCFF04FB4BFB27FD24.taxon	description	Description. Male holotype. Head. Labium whitish yellow. Genae, base of mandibles and labrum greyish green; anteclypeus and postclypeus black; antefrons greyish green; top of head black except for remnants of a pale stellate marking in front of the median ocellus, a poorly defined pale marking between the eyes and the lateral ocelli a yellowish occipital and postocular areas and a blue rounded postocular spot. Rear of the head yellowish white turning to yellowish blue near eyes. Thorax. Prothorax: Dorsally brown, laterally greyish blue with a bluish area on anterior lobe. Pterothorax: mesopleuron brown, metapleuron yellowish. Blue stripes as follows: at posterior part of mesepisternum, at metepisternum and lateral part of metepimeron. Legs yellow, wings hyaline, venation light brown. Pterostigma brown surrounded with yellow; Px in FW 12; in HW 11 – 12; R 3 originating near Px 6 in FW and near 5 in HW. Petiolation distal to Ac by a distance ½ length of Ac in FW and only slightly distal in HW. Abdomen. Missing. Structural characters. Hind prothoracic lobe with lateral and median lobes well-developed (Fig. 37), the latter subtriangular and two-lipped. Posterior border of mesostigmal plate connected with mid-dorsal carina (Fig. 37). Mesostigmal plates curved and directed anteriorly (Fig. 37). Mesepisternal tubercles (Figs 37 – 38) well separated from mid-dorsal carina, low (0.78 mm), in lateral view nipple-shaped, in dorsal view tearshaped with antero-lateral extremity tapering into a fine tip (Fig. 37). Dimensions (mm). HW 17.1 Female. Unknown.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCFFFFCFF04FB4BFB27FD24.taxon	discussion	Remarks. I hesitated in describing this species because of the missing abdomen. However, its thorax offers sufficient characters to distinguish it from males of all other known species of this genus. None of the 11 species of Tuberculobasis whose males have been described have tear-shaped mesepisternal tubercles and mesostigmal plates curved and directed anteriorly as does T. karitiana. Since the male of T. cardinalis from Peru remains unknown, T. karitiana might represent the male sex of T. cardinalis. However, I think this unlikely because T. cardinalis is much bigger. The size difference between female of T. cardinalis (HW 20 mm) and male of T. karitiana (HW 17 mm) is more than would be expected taking as reference the differences in size between males and females of the six species for which both sexes are known.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCEFFFCFF04FD3BFE0CF8F5.taxon	etymology	Etymology: The name refers to the Macuxi indian people, who are settled in the region where the holotype was collected.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCEFFFCFF04FD3BFE0CF8F5.taxon	materials_examined	Type specimens: Holotype (3): BRAZIL, Roraima State, Maracá Ecological Station, Maracá Island (3 º 25´N, 61 º 36´W), 24 - II- 1988. P. A. R. Machado leg (within forest, away from water) (ABMM).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCEFFFCFF04FD3BFE0CF8F5.taxon	description	Description. Male holotype. Head. Labium yellowish white. Labrum and anteclypeus yellow; genae and base of mandibles grey; postclypeus, antefrons, and top of head brown; postocular lobe yellowish ochre. A subtriangular blue postocular spot adjacent to eye. Rear of head yellowish white. Thorax. Prothorax: Pronotum yellow ochre with bluish markings on anterior lobe. Propleuron yellowish with greyish areas. Pterothorax: mesopleuron and metepisternum yellow ochre with a narrow light blue stripe at middle part of mesepisternum and at anterior part of metepisternum. Metepimeron yellowish white. Legs yellowish white, wings hyaline, venation brown. Pterostigma yellow. Px in FW 11, in HW 10. R 3 originating between Px 5 and 6 in both wings. Petiolation distal to Ac by a distance equal or twice as long as length of Ac in both wings. Abdomen. S 1 – 2 dorsally yellow ochre, laterally light blue, ventrally yellowish white with a dark brown terminal line. S 3 – 7 light brown dorsally, laterally yellow with a dark brown distal ring. S 8 – 10, cercus, and paraproct light brown. Structural characters. Hind prothoracic lobe with well-developed lateral lobes (Fig. 45). Median lobe semicircular and two-lipped; ventral lip more produced than dorsal lip. Mesepisternal tubercles (Figs 45, 46) subcylindrical, high (1.51 mm), directed dorsally, with bases well separated from mid-dorsal carina. Cercus about 1 / 2 shorter than paraproct (Figs. 69 – 70), ventral process in lateral view tapering into a chitinized tip directed latero-posteriorly (Fig. 71). Penis as described for genus (as in Fig. 89). Dimensions (mm). Abdomen 21.9; HW 15.35	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCEFFFCFF04FD3BFE0CF8F5.taxon	discussion	Remarks. T. macuxi has high mesepisternal tubercles with bases well separated from mid-dorsal carina, long paraprocts, and hind prothoracic lobe with well-developed lateral lobes. This combination makes it unique within the genus belonging to none of the two species groups recognized here.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFCEFFFCFF04FD3BFE0CF8F5.taxon	description	Female. Unknown.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC1FFF1FF04FF70FE6DFD92.taxon	distribution	Distribution. Brazil, Venezuela, and Suriname.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC1FFF1FF04FF70FE6DFD92.taxon	discussion	Remarks. T. mammilaris was described by Calvert (1909) based on a complete male and parts of a second male from Chapada in Mato Grosso State, Brazil, and a single female from Rio de Janeiro. The description is good with simple illustrations of male appendages and thorax and female hind prothoracic lobe. Recently R. W. Garrison studied the complete male syntype from CMNH and provided good illustrations of the main structural characters which were reproduced in Lencioni (2006) and are also included here. De Marmels (1992) recorded a male and a female of T. mammilaris from Venezuela and his identification of the male was confirmed by R. W. Garrison by direct comparison with the complete male syntype. De Marmels (1992) illustrated hind prothoracic lobe of the female from Venezuela, which is completely different from the one illustrated by Calvert (1909, Fig. 117) for the syntype female from Rio de Janeiro. In view of this, De Marmels (1992) concluded that the true female of T. mammilaris is that from Venezuela. R. W. Garrison has examined Calvert´s syntype female labeled as L. mammilaris and has identified it as belonging to Leptobasis sensu stricto. In order to clarify application of the name, I designate the complete syntype male from Brazil, Chapada, H. H. Smith leg. (CMNH) illustrated by Calvert (1909, Figs. 112, 113, 115, 116) and by R. W. Garrison (Figs 24, 78 – 80, 91) as lectotype. The description of Calvert (1909) does not mention the presence of pale postocular spots and R. W. Garrison (in litt.) confirmed that there are lacking in the lectotype. The male from Venezuela differs from the lectotype by having pale postocular spots that are present also in females, meaning that this character is variable in T. mammilaris.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC3FFF7FF04FD61FE21F87D.taxon	etymology	Etymology: The name refers to the Tirio indian people, who helped the author’s collecting activities during his one- month visit to their village.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC3FFF7FF04FD61FE21F87D.taxon	materials_examined	Type specimens: Holotype (3) and allotype (Ƥ): BRAZIL, Pará State, Tumucumaque Indian Park, two days walk east from the Tirió Indian village (temporary pool in savannah II- 1963), Machado & Acewa leg. (ABMM). Paratype (1 3) same data as holotype (MNRJ).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC3FFF7FF04FD61FE21F87D.taxon	description	Description. Male holotype. Head. Labium yellowish white. Genae and base of mandibles dark brown. Labrum, anteclypeus, postclypeus, and antefrons black. Top of head black. Postocular spot and area behind eyes orange brown. Rear of head yellowish white. Thorax. Prothorax. Pronotum brownish yellow. Propleuron greyish blue. Pterothorax with bluish grey or silver stripes as follows: one on the mesepisternum adjacent to the humeral suture, another on metepisternum continuing on to mesinfrepisternum and a third one on metepimeron. Legs yellowish, wings hyaline, venation brown, pterostigma light brown surrounded with yellow. Px in FW 11, in HW 9 – 10 R 3 in FW originating near Px 5, in HW near 4 .. Petiolation distal to Ac by a distance equal (50 %) or ½ as long (50 %) as Ac in both wings. Abdomen. S 1 – 2 dorsally brown laterally bluish gray. S 3 – 7 dorsally brown ventro-laterally brownish yellow. S 8 – 10, cercus, and paraproct brownish orange. Structural characters. Hind prothoracic lobe with well-developed lateral lobes (Fig. 41); median lobe projected caudally, subtriangular, with borders two-lipped. Mesepisternal tubercles low (0.64 mm), in dorsal view rounded, with bases well separated from mid-dorsal carina (Fig. 41). Cercus subquadrangular in dorsal view (Fig. 81); ventral process in posterior view with medial margin straight (Fig. 83), in lateral view shaped like an isosceles triangle tapering into a narrow, straight tip (Fig. 82). Paraproct subequal to cercus. Penis as described for genus, but filiform process of internal fold extends beyond lateral border of penis (Fig. 92). Dimensions (mm). Abdomen 26.77 – 27.50 (mean 26.91); HW 14.8 – 15.73 (mean 15.32). Description. Female holotype Head. Labium yellowish. Labrum brown surrounded by yellow. Base of mandibles and genae yellow; anteclypeus and postclypeus black; antefrons and top of head dominantly brownish orange, with a black band adjoining the eye dilated posteriorly to form a large subtriangular black area connected with that of the opposite side by a narrow black stripe at occipital bar. A rounded blue postocular spot within black area. A transverse orange stripe at postocular area connecting with that of the opposite side through occipital bar. Antennae brown, the two basal segments black. Thorax. Color including legs and wings, as in the male. Px in FW 12, in HW 11; R 3 in FW originating near Px 6, in HW near 5. Petiolation distal to Ac by a distance equal to length in both wings. Abdomen. S 1 dorsally orange, laterally bluish grey, S 2 and proximal 1 / 3 of S 3 dorsally brown, laterally bluish grey. Distal part of S 3, and S 4 – 8 dorsally brown, ventro-laterally brownish yellow. S 9 – 10 brownish yellow. Structural characters. Hind prothoracic lobe with a semicircular and depressed median lobe separated from lateral lobes by two deep concavities (Fig. 51). Hind margin of mesostigmal plate curved anteriorly and then laterally forming a C-shaped structure (Fig. 51). Mesepisternal tubercles lacking. Dimensions (mm). Abdomen 30.71; HW 18.82.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC3FFF7FF04FD61FE21F87D.taxon	discussion	Remarks. T. tirio belongs in the mammilaris species group together with T. geijskesi, T. karitiana, T. mammilaris, and T. yanomami. Male is very close to T. mammilaris, but female is more similar to T. yanomami. Differences are given in the keys. Specimens were collected at temporary pools in the savannah. The ecological significance of this habitat is discussed under Ecological Considerations. Characters Tuberculobasis Leptobasis Metaleptobasis Denticulobasis Frons Rounded Rounded Angulated Rounded Pale postocular Present 97.7 %, absent Present 53.2 %, Absent 100 % Present 100 % spots * 2.23 % absent 46.8 % n = 25 n = 7 n = 45 n = 32 Median lobe of Distinguishable and Poorly Mostly distinguishable Distinguishable and hind prothoracic projected caudally distinguishable and but not projected projectedcaudally lobe in males not projected caudally caudally Mesepisternal Present in males and Absent Present in males and Present in males and processes most females females females Type of Mostly horns in males, Horns in males, mesepisternal Tubercles No processes tubercles in females tubercles in females processes Supplementary Well-developed but tooth of tarsal claws Well-developed small Absent Well-developed Petiolation in At or slightly distal: At or slightly Distal by ½ to twice Distal by ½ to twice relation to Ac ** 11.74 %, distal by ½ to distal: 100 % length of Ac (100 %) length of Ac l: 100 % twice length of Ac: n = 133 n = 90 n = 28 88.25 % n = 315 Pair of lateral chitinized spines on Absent Present Absent Absent distal segment of penis Penis internal fold Present Absent Absent Present Filiform process of penis internal fold Present Absent Absent Present Length of cercus in relation to S 10 *** Much shorter Much shorter Subequal Subequal Ventral process of Present Present Absent Absent cercus Membranous depression at dorsal Absent Absent Absent Present surface of cercus Vulvar spine **** Present 98.2 %, absent Present 73.3 %, Absent 100 % Replaced by a 0.98 % absent 26.7 % n = 20 denticulated vulvar n = 57 n = 30 plate Ovipositor Not surpassing tip of Surpassing tip of Not surpassing tip of Not surpassing tip of cercus cercus cercus cercus Denticles of ventral Subtriangular, high and Subtriangular, low Subrectagular, high and border of ovipositor directed ventro- and directed directed ventro- lateral valve in medially. Associated posteriorly. Not studied with SEM medially. Associated S. E. M. seta long and fine tipped Associated seta seta short with apex long and fine truncated tipped * n = number of specimens herein described, those from the literature plus 30 specimens of T. costalimai, 32 of L. vacillans, and 25 specimens of 10 species of Metaleptobasis. ** n = number of wings of species here described plus 80 wings of T. costalimai, 30 of L. vacillans and 90 of 10 species of Metaleptobasis. *** excluding distal lobe. **** n = number of female specimens of species here described, those from literature plus 40 females of T. costalimai and 30 of L. vacillans.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC7FFF4FF04FBB8FA32FC36.taxon	etymology	Etymology: Named after Edward Bruce Williamson, whose odonatological expeditions to Colombia and Venezuela yielded this species and an enormous number of valuable odonata specimens.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC7FFF4FF04FBB8FA32FC36.taxon	materials_examined	Type specimens: Holotype (3) and allotype (Ƥ): COLOMBIA, Fundación, State of Magdalena, 11 - I- 1917, J. H. & E. B. Williamson leg. (UMMZ). Paratypes: 3 3 and 5 Ƥ: same data as holotype (three paratypes were collected along a stream in a shaded area); 1 3, 2 Ƥ: El Banco, State of Magdalena, 21 - I- 1917, J. H. & E. B. Williamson leg.; 1 3, 2 Ƥ: VENEZUELA, Zulia State, Encontrados, 20 - IV- 1920, J. M., E. B. Williamson & D. H. Ditzleri leg; 1 3: San Felipe, 20 - III- 1920, J. M., E. B. Williamson & D. H. Ditzleri leg .. 3 3 and 7 Ƥ paratypes (UMMZ), 1 3 and 1 Ƥ paratypes (MNRJ); 1 3 and 1 Ƥ (A. B. M. M), 1 3 and 1 Ƥ (R. W. G.).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC7FFF4FF04FBB8FA32FC36.taxon	description	Description. Male holotype. Head. Labium yellowish white. Labrum yellow; genae grey, base of mandibles, anteclypeus, postclypeus, antefrons, and top of head brown. Postocular lobes light orange with grey postocular spots. Rear of head yellowish white. Thorax. Prothorax: Anterior lobe grey, median and posterior lobes brown. Propleuron yellowish brown. Pterothorax (as Fig. 101 b) brown with greyish stripes on mesepisternum and metepisternum. Legs yellowish brown, wings hyaline, venation and pterostigma brown. Px in FW 11 (10 %), 12 (90 %); in HW 9 (30 %), 10 (40 %), 11 (30 %). R 3 in FW arising near Px 5 (20 %) and 6 (80 %), in HW at Px 4 (10 %), 5 (80 %), 6 (10 %). Petiolation distal to Ac by a distance equal (60 %) or ½ (40 %) of its length in FW, equal (30 %) or ½ (70 %) in HW. Abdomen. S 1 – 2 brownish yellow with a lateral grey stripe. S 2 with a square yellow dorsal spot at distal 1 / 4. S 3 – 5 dorsally yellowish brown, laterally yellowish with brown distal rings; S 6 brown, S 7 – 10 yellowish brown. Cercus brownish yellow with dorso-lateral border dark brown; paraproct brownish. Structural characters. Hind lobe of prothorax (Figs 33, 34) with median lobe semicircular and projected caudally; lateral lobes very small. Borders of median lobe two-lipped; dorsal lip upraised. Mesepisternal tubercles high (1.58 mm) in dorsal view strongly divergent (Fig. 33), conical, directed antero-laterally, with bases adjacent to mid-dorsal carina (Fig. 33). Cercus 1 / 2 shorter than paraproct (Figs. 72 – 73), in posterior view (Fig. 74) with a dorso-lateral semilunar cavity limited by a denticulated dark brown border. Penis (Fig. 93) as described for the genus. Dimensions (mm). Abdomen 32.0 – 34.4 (mean 32.6); HW 17.5 – 19.0 (mean 18.2). Description. Female allotype. Color of head, thorax, and abdomen as described for male. Px in FW 12 (30 %), 13 (60 %), 14 (10 %), in HW 10 (40 %), 11 (30 %), 12 (30 %). R 3 in FW originating near Px 6 (90 %) or 5 (10 %) in HW near Px 5 (80 %) or 6 (20 %). Petiolation in FW distal to Ac by a distance equal (14.2 %) or ½ (71.5 %) of its own length, in HW slightly distal (14.2 %), equal (14.2 %) or ½ (71.4 %) its own length. Structural characters. Hind lobe of prothorax (Fig. 52) slightly concave at middle with no definition of lateral lobes. Mesepisternal tubercles well-developed but low, connected with anterior margin of mesostigmal plate (Fig. 52). Cercus conical (Fig. 98). Ovipositor (excluding styli) not surpassing level of cercus tip (Fig. 98). Dimensions (mm). Abdomen 31.0 – 32.1 (mean 31.5); HW 18.9 – 20.4 (mean 19.8).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC7FFF4FF04FBB8FA32FC36.taxon	discussion	Remarks. By having male mesepisternal tubercles high with base adjacent to mid-dorsal carina, and male paraproct much longer than cercus, T. williamsoni belongs in the costalimai species group, together with T. arara, T. costalimai, T. guarani, and T. inversa. It can be separated from them by characters given in the keys.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC6FFCBFF04F970FC5EFF59.taxon	description	Specimens examined. 1 (Ƥ) BRAZIL, Amazonas State, Manaus, INPA, 19 - VI- 1977, Joselita Santos leg. (ABMM).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC6FFCBFF04F970FC5EFF59.taxon	discussion	Remarks. This species is remarkable by the size and shape of its mesepisternal tubercles (Fig. 54). Whereas in all known females of Tuberculobasis tubercles are absent or small, in this species they are as high as those of males of the costalimai group. By having a curved plate connecting tubercles with mesostigmal plate it resembles males of T. inversa. Hind prothoracic lobe is also quite different from that of other known species, but median lobe is single-lipped as in all females of Tuberculobasis. It has postocular spots and it is most probably a new species, but I refrain from naming it pending the possibility that it might be the female of one of the six species of Tuberculobasis of which only the male is known.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC6FFF4FF04FC0DFE25F998.taxon	distribution	Distribution. Venezuela, Amazonas Federal Territory and Brazil, State of Pará.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFC6FFF4FF04FC0DFE25F998.taxon	discussion	Remarks. By having well-developed lateral lobes, low mesepisternal tubercles, and paraprocts about as long as cerci, T. yanomami belongs in the mammilaris species group, together with T. geijskesi, T. karitiana, T. mammilaris, and T. tirio. It can be separated from these species by characters given in the keys. Male of T. yanomami is close to T. mammilaris, but female is closer to T. tirio. The three type specimens described by De Marmels (1992) were collected in the forest on sun-hit bushes, away from water. This is the first record of this species from Brazil.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFF9FFCBFF04FEBEFC35FD54.taxon	description	Specimens examined. 1 Ƥ, PERU, Loreto Department, Exploration Lodge, 50 miles, NE of Iquitos on Amazon River at junction of the Yanomano River (3 º 21´59 " S, 72 º 47´56 " W), 27 - VIII- 1991, S. W. Dunkle leg (SWD).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFF9FFCBFF04FEBEFC35FD54.taxon	discussion	Remarks. Median lobe of hind prothoracic lobe (Fig. 55) similar to that of female of T. williamsoni (Fig. 52), but mesepisternal tubercles different and not connected with mesostigmal plate as in T. williamsoni (Fig. 52). Tubercle of Tuberculobasis sp. 2 is unique by having the shape of a ball with a basal ring (Fig. 55). This species also has top of head orange and it is most probably new. However, since it could represent the female one of the six species with unknown females, I am not naming it.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFF9FFCAFF04F959FC4AFED5.taxon	description	Specimens examined. 1 Ƥ, SURINAME, Boshhinis, 19 - IV- 1943, D. K. Geijskes leg. (MNRJ).	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFF9FFCAFF04F959FC4AFED5.taxon	discussion	Remarks. So far two species of Tuberculobasis have been reported from Suriname: T. mammilaris by Belle (2002) and T. geijskesi described here. Median lobe of hind prothoracic lobe of T. sp. 3 (Fig. 56) is identical to that of the female of T. mammilaris (Fig. 50), and mesostigmal plate is also similar. However, an important difference is the presence of mesepisternal tubercles in T. sp. 3 and their absence in the female of T. mammilaris. It is unlikely that T. sp. 3 represents the female of T. geijskesi in spite of the fact that these two species were collected in Suriname. Tuberculobasis geijskesi belongs in the mammilaris species group together with T. tirio, T. mammilaris, and T. yanomami. In all of them, females have no mesepisternal tubercles, and the same most probably occurs with the so far unknown female of T. geijskesi. Thus, T. sp. 3 is most likely a new species. Since, however, it might represent the female of one of the six species of Tuberculobasis known only from the male, I refrain from naming it here.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
1312774BFFF8FFCAFF04FE2AFC0AFCBE.taxon	description	According to O'Grady & May (2003), the taxonomic characters commonly used for subfamily identification in Coenagrionidae are continuously distributed and subfamilies should not be recognized within Coenagrionidae until well-supported subdivisions are demonstrated. In view of this, I refrain from ascribing Denticulobasis and Tuberculobasis to any of the extant subfamilies. Denticulobasis and Tuberculobasis are close to Leptobasis and key out to this genus in Von Ellenrieder & Garrison (2008). Both genera share with Metaleptobasis the presence of mesepisternal processes that occur also in some species of Argia (Garrison, 1994). Table 2 shows the characters that separate these four genera.	en	Machado, Angelo B. M. (2009): Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa 2108: 1-36, DOI: 10.5281/zenodo.187806
