identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
131BBD7A94379433FDEE577CFC3FFEE4.text	131BBD7A94379433FDEE577CFC3FFEE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia Turpaeva 1973	<div><p>Genus Hedgpethia Turpaeva, 1973</p><p>Type species</p><p>Colossendeis arcuata Loman, 1908 (by original designation).</p><p>Diagnosis of the genus (emended from Turpaeva 1973 and Takahashi et al. 2007)</p><p>Trunk four-segmented, lateral processes separated by twice their own diameter or less. Cephalon with pair of anterior projections on both sides of proboscis, on which palps articulate; anterior projection individualised, base segmented, apparently not movable. Proboscis massive, as long or longer than body, inflated around midpoint, constricted at base, tapering at tip. Abdomen small, ventralised and bent downward. Chelifores absent. Palps 9-articled. Ovigers 10-articled plus terminal claw. Legs slender. Propodus devoid of auxiliary claws.</p><p>Also includes Hedgpethia caudata Turpaeva, 1993 which does not match this diagnosis (see Discussion).</p><p>Remarks</p><p>Palps in species of Hedgpethia have been generally regarded as 10-articled. This number is due to the cephalic anterior projections, of which the base seems articled, is included in the count. This projection is likely homologous to that found in the closely related genus Colossendeis, which was demonstrated to be part of the cephalon, and not an article (Cano-Sánchez &amp; López-González 2016). The palps in Hedgpethia are therefore regarded as 9-articled in the present work.</p></div>	https://treatment.plazi.org/id/131BBD7A94379433FDEE577CFC3FFEE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94369433FD9F53FAFC89FBC3.text	131BBD7A94369433FD9F53FAFC89FBC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia articulata (Loman 1908)	<div><p>Hedgpethia articulata (Loman, 1908)</p><p>Fig. 2, Table 1</p><p>Colossendeis articulata Loman, 1908: 22–23, pl. 6 figs 66–73.</p><p>Colossendeis articulata – Losina-Losinsky &amp; Turpaeva 1958: 32 [key], tab. 4. — Stock 1958: tab. 1.</p><p>Rhopalorhynchus articulatum – Stock 1958: 116 [text], 118 [key], 136 [text], tab. 1; 1970: 9 [key].</p><p>Hedgpethia articulata – Turpaeva 1973: 185–186, fig. 2; 1993: 21 [text]. — Stock 1991a: 166 [text]. — Child 1994: 18 [text].</p><p>Type material (not examined)</p><p>Holotype</p><p>BANDA SEA • ♂; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.3&amp;materialsCitation.latitude=-5.4333334" title="Search Plazi for locations around (long 121.3/lat -5.4333334)">Gulf of Boni, west off Kabaena Island</a>; 5°26′ S, 121°18′ E; depth 1944 m; Siboga Expedition stn 210 a; NL, ZMA.PYC.1322.</p><p>Distribution and depth range</p><p>Besides the Gulf of Boni in Banda Sea (type locality), additional data on GBIF (2025) extends the distribution to the Gulf of Alaska and in the vicinities of the Commander Islands. The divergent collection localities may suggest the presence of more than one species, and this northern material should be reexamined side by side with the holotype. Minimal known depth range 1944–4200 m (1944 m only when excluding Alaska and Commander Islands material).</p></div>	https://treatment.plazi.org/id/131BBD7A94369433FD9F53FAFC89FBC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94369433FDE256E1FDD9F8FB.text	131BBD7A94369433FDE256E1FDD9F8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia atlantica (Stock 1970)	<div><p>Hedgpethia atlantica (Stock, 1970)</p><p>Fig. 2, Table 1</p><p>Rhopalorhynchus atlanticum Stock, 1970: 7–9, 9 [key], figs 8–13.</p><p>Rhopalorhynchus atlanticum – Rack 1971: 113. –– Arnaud 1987: 51–52. — Stock 1987: 508–509, 517 [text], fig. 12.</p><p>Hedgpethia atlantica – Stock 1991a: 166 [text]; 1991b: 138. — Turpaeva 1993: 22 [text]. — Munilla &amp; Soler-Membrives 2014: 218–220 [“ Hedgpthetia (sic) atlantica ”], fig. 120. — Soler-Membrives &amp; Munilla 2015: 11, 16 [text], tab. 2. — Staples 2019: 433 [text].</p><p>Type material (not examined)</p><p>Holotype</p><p>NORTHEASTERN ATLANTIC • ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-14.258333&amp;materialsCitation.latitude=36.678333" title="Search Plazi for locations around (long -14.258333/lat 36.678333)">Josephine Bank</a>; 36°40.7′ N, 14°15.5′ W; depth 211–241 m; 1 Jul. 1967; Meteor stn 120; ZMH-A50168.</p><p>Distribution and depth range</p><p>Eastern North Atlantic (West to Galicia, Portugal and Tanger), including Josephine Bank (type locality), and across the Gibraltar Strait east to the Île du Levant, Hyères (Mediterranean French coast). Minimal known depth range 100–1110 m.</p></div>	https://treatment.plazi.org/id/131BBD7A94369433FDE256E1FDD9F8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94319434FED15376FAA0FA6D.text	131BBD7A94319434FED15376FAA0FA6D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia bicornis (Turpaeva 1958)	<div><p>Hedgpethia bicornis (Turpaeva, in Losina-Losinsky &amp; Turpaeva, 1958)</p><p>Fig. 2, Table 1</p><p>Colossendeis bicornis Turpaeva in Losina-Losinsky &amp; Turpaeva, 1958: 27–29, 32 [key], 33 [text], fig. 3, tabs 3–5.</p><p>Colossendeis bicornis – Losina-Losinsky 1961: 109–110.</p><p>Rhopalorhynchus bicornis – Stock 1970: 9 [key].</p><p>Hedgpethia californica – Turpaeva 1973 [partim]: 185 [key], 186–189, tab. 2.</p><p>Hedgpethia californica bicornis – Turpaeva 1973: 187–189 [text], tab. 3; 2002: 1451 [text], 1451, 1452. — Dudnik &amp; Kremenetskaia (2025): fig. 6d, tab. 1.</p><p>Hedgpethia bicornis – Stock 1991a: 166 [text]. — Turpaeva 1993: 22 [text].</p><p>Type material (not examined)</p><p>Holotype</p><p>SEA OF OKHOTSK • sex unknown; northeastern Sea of Okhotsk; depth 1100 m; 3 Oct. 1952; R/V Vityaz cruise 12 stn 1780; IORAS, INV0002674.</p><p>Paratypes</p><p>SEA OF OKHOTSK • 2 specs; same data as for holotype; hosting institution unknown • 1 spec.; southern Sea of Okhotsk; 10 Aug 1949; R/V Vityaz; hosting institution unknown • 1 spec.; 1932; R/V Gagara stn 217; hosting institution unknown • 19 specs; 1932; R/V Gagara stn 247; hosting institution unknown • 1 spec.; 1932; R/V Gagara stn 250; hosting institution unknown • 1 spec.; 1932; R/V Gagara stn 261; hosting institution unknown • 2 specs; 1952; R/V Guidrolog; hosting institution unknown .</p><p>NORTHWESTERN PACIFIC • 2 specs; south of Southern Kuril Islands; 1949; R/V Toporok; hosting institution unknown .</p><p>Distribution and depth range</p><p>Sea of Okhotsk (type locality) and east to Kuril Islands. Minimal known depth range 303–1100 m.</p><p>Remarks</p><p>Stock (1991a) suggested this species to be a possible junior synonym to Hedgpethia californica (Hedgpeth, 1939), without explicit justification. However, the mediodorsal ornamentation of H. bicornis as well as the low tubercle on the anterior margin of its cephalon distinguish it from H. californica .</p></div>	https://treatment.plazi.org/id/131BBD7A94319434FED15376FAA0FA6D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94319435FE9B5473FB4CFC81.text	131BBD7A94319435FE9B5473FB4CFC81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia brevitarsis (Losina-Losinsky & Turpaeva 1958)	<div><p>Hedgpethia brevitarsis (Losina-Losinsky in Losina-Losinsky &amp; Turpaeva, 1958)</p><p>Fig. 2, Table 1</p><p>Colossendeis brevitarsis Losina-Losinsky in Losina-Losinsky &amp; Turpaeva, 1958: 26–27, 32 [key], 33 [text], fig. 2, tabs 2, 4–5.</p><p>Colossendeis brevitarsis – Losina-Losinsky 1961: 109.</p><p>Rhopalorhynchus brevitarsis – Stock 1970: 9 [key].</p><p>Hedgpethia brevitarsis (Losina-Losinsky &amp; Turpaeva, 1958) [sic]. — Nakamura &amp; Child 1983: 63, 63[key]; 1991: 62. — Child 1998: 60–61 [text].</p><p>Hedgpethia brevitarsis – Turpaeva 1973: 185 [key], tab. 2; 1993: 22 [text]; 2002: 1451. — Stock 1991a: 166 [text].</p><p>“ Hedgpethia borealis ” – Turpaeva 2002 [error in the English translation of the abstract].</p><p>Type material (not examined)</p><p>Holotype</p><p>SEA OF OKHOTSK • sex unknown; southern Sea of Okhotsk; depth unknown; 11 May 1953; R/V Vityaz; hosting institution unknown .</p><p>Paratypes</p><p>SEA OF OKHOTSK • 10 specs; depth unknown; 1932; R/V Gagara; hosting institution unknown .</p><p>Distribution and depth range</p><p>Sea of Okhotsk (type locality), South Kuril Islands and Eastern Honshu coast (Sagami Bay). Depth range 300– 780 m.</p><p>Remarks</p><p>The types should be at the IORAS, like the holotype of H. bicornis, but they were not listed in Dudnik &amp; Kremenetskaia (2025) nor were they retrieved after examination (Dudnik pers. com.).</p></div>	https://treatment.plazi.org/id/131BBD7A94319435FE9B5473FB4CFC81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94309435FD86511EFE3CF901.text	131BBD7A94309435FD86511EFE3CF901.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia californica (Hedgpeth 1939)	<div><p>Hedgpethia californica (Hedgpeth, 1939)</p><p>Fig. 2, Table 1</p><p>Colossendeis californica Hedgpeth, 1939: 59–60, pl. 1 figs a–e.</p><p>Colossendeis californica – Hilton 1943: 4. — Losina-Losinsky &amp; Turpaeva 1958: 32 [key], tab. 4. – Stock 1958: tab. 1.</p><p>Rhopalorhynchus californicum – Stock 1958: 116 [text], 118 [key], tab. 1; 1970: 9 [key].</p><p>Hedgpethia californica – Turpaeva 1973 [partim]: 185 [key], 186–189, tab. 2; 1993: 21–22 [text]; 2002: 1451. — Stock 1991a: 166 [text].</p><p>Hedgpethia californica californica – Turpaeva 1973: 186–189 [text], tab. 3; 2002: 1451 [text], 1452 [key].</p><p>Type material (not examined)</p><p>Holotype</p><p>NORTHEASTERN PACIFIC • ♂; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.41333&amp;materialsCitation.latitude=32.496666" title="Search Plazi for locations around (long -117.41333/lat 32.496666)">Off Tijuana</a>; 32°29.8′ N, 117°24.8′ W; depth 330 m; 2 Jul. 1916; Scripps Institution of Oceanography coll. no 1189; USNM 78022.</p><p>Paratype</p><p>NORTHEASTERN PACIFIC • ♂; same data as for holotype; USNM 78021 .</p><p>Distribution and depth range</p><p>Besides the type locality (off Tijuana), specimens were sampled in the Gulf of Alaska. Minimal known depth range 330– 560 m.</p></div>	https://treatment.plazi.org/id/131BBD7A94309435FD86511EFE3CF901	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94309436FDF9549EFE74FD75.text	131BBD7A94309436FDF9549EFE74FD75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia calva Arango 2009	<div><p>Hedgpethia calva Arango, 2009</p><p>Fig. 2, Table 1</p><p>Hedgpethia calva Arango, 2009: 5–7, figs 3, 8a.</p><p>Hedgpethia calva – Staples 2019: 433 [text].</p><p>Type material (not examined)</p><p>Holotype</p><p>GREAT AUSTRALIAN BIGHT • ♀; Southwestern Australia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.709&amp;materialsCitation.latitude=-35.281" title="Search Plazi for locations around (long 118.709/lat -35.281)">Bald Island</a>; 35°16′53.3″ S, 18°42′32.4″ E [sic; 118°42′32.4″ E] – 35°16′51.6″ S, 118°47′56.4″ NE; depth 976–980 m; 24. Nov. 2005; Australian Commonwealth Scientific and Industrial Research Organization (CSIRO) ‘Voyage of Discovery’ survey stn 036-008; WAM T92360.</p><p>Paratypes</p><p>GREAT AUSTRALIAN BIGHT • 2 ♂♂; same data as for holotype; NVM JM54989 .</p><p>Distribution and depth range</p><p>Reported only from Bald Island in the Great Australian Bight (type locality). Maximal known depth range 976– 980 m.</p></div>	https://treatment.plazi.org/id/131BBD7A94309436FDF9549EFE74FD75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94339436FDE4516AFE55FAD2.text	131BBD7A94339436FDE4516AFE55FAD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia caudata Turpaeva 1993	<div><p>Hedgpethia caudata Turpaeva, 1993</p><p>Fig. 2, Table 1</p><p>Hedgpethia caudata Turpaeva, 1993: 20–22, fig. 1.</p><p>Type material (not examined)</p><p>Holotype</p><p>SOUTHEASTERN PACIFIC • sex unknown; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.986664&amp;materialsCitation.latitude=-17.7" title="Search Plazi for locations around (long -78.986664/lat -17.7)">off Southern</a> Peru; 17°42′ S, 78°59.2′ W; 2710–3080 m; 20 Nov. 1968; R/V Akademik Kurtchatov stn 271; hosting institution unknown.</p><p>Distribution and depth range</p><p>Only known from the type locality (off Southern Peru); Maximal known depth range 2710–3080 m.</p><p>Remarks</p><p>The holotype was not retrieved; it was originally deposited at the IORAS as explicitly mentioned by Turpaeva (1993), but was not listed by Dudnik &amp; Kremenetskaia (2025) and was not retrieved there (Dudnik pers. com.).</p></div>	https://treatment.plazi.org/id/131BBD7A94339436FDE4516AFE55FAD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94339437FDE457D3FC9CFCC1.text	131BBD7A94339437FDE457D3FC9CFCC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia chitinosa (Hilton 1943)	<div><p>Hedgpethia chitinosa (Hilton, 1943)</p><p>Fig. 2, Table 1</p><p>Colossendeis chitinosa Hilton, 1943: 4 .</p><p>Colossendeis chitinosa – Hedgpeth 1949: 301, fig. 47e–h. — Stock 1954: 161, fig. 79; 1958: tab. 1. — Utinomi 1955: 34–35, fig. 20; 1959: 220; 1971: 337–338. — Losina-Losinsky &amp; Turpaeva 1958: 23–26, 32 [key], 33 [text], fig. 1, tabs 1, 4–5. — Child 1996: tab. 1.</p><p>Rhopalorhynchus chitinosum – Stock 1958: 116 [text], 118 [key], tab. 1; 1970: 9 [key].</p><p>Hedgpethia californica – Turpaeva 1973 [partim]: 185 [key], 186–189, tab. 2. — Child 1996: tab. 1.</p><p>Hedgpethia californica chitinosa – Turpaeva 1973: 186–189, tab. 3; 2002: 1451, 1451 [text], 1452 [key].</p><p>Hedgpethia chitinosa – Nakamura &amp; Child 1983: 63, 63 [key]; 1991: 63. — Nakamura 1987: 36–37, pl. 33. — Stock 1991a: 166 [text]. — Turpaeva 1993: 21 [text]. — Dudnik &amp; Kremenetskaia 2025: tab. 3.</p><p>Type material (not examined)</p><p>Holotype</p><p>BERING SEA • ♀; Aleutians West Census Area (Alaska), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-169.5&amp;materialsCitation.latitude=56.0333" title="Search Plazi for locations around (long -169.5/lat 56.0333)">Pribilof Islands, south of Saint George Island</a>; 56°1′59.88″ N, 169°30′ W; depth 0–221 m; 17 Jul. 1893; R/V Albatross, United States Fish Commission (USFC) stn 3500; USNM 81496.</p><p>Paratypes BERING SEA • 3 specs; Aleutians West Census Area (Alaska), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-170.35&amp;materialsCitation.latitude=58.383335" title="Search Plazi for locations around (long -170.35/lat 58.383335)">Pribilof Islands, north of Saint Paul Island</a>; 58°23′ N, 170°21′ W; depth 3939 m; 11 Aug. 1895; USFC stn 3603; hosting institution unknown .</p><p>Distribution and depth range</p><p>Recorded from the Aleutian Islands and the Bering Sea, including off Pribilof Islands (type locality), as well as around Japan (Sea of Japan, coasts of Honshu, Hokkaido and in the Tsushima Straits; Pacific coasts Honshu, Shikoku, Kyushu and down to southwest of Danjo Islands). Minimal known depth range 41–926 m? Hilton (1943) cites three specimens from 1771 fathoms (3939 m), and Dudnik (2025) records one specimen sampled at depths of 6410–6757 m. These four specimens should be reexamined for confirmation, and contamination should also be considered.</p></div>	https://treatment.plazi.org/id/131BBD7A94339437FDE457D3FC9CFCC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94329437FDE251DEFD78F962.text	131BBD7A94329437FDE251DEFD78F962.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia dampieri (Child 1975)	<div><p>Hedgpethia dampieri (Child, 1975)</p><p>Fig. 2, Table 1</p><p>Rhopalorhynchus dampieri Child, 1975: 8–10, fig. 4.</p><p>Hedgpethia dampieri – Stock 1991a: 166 [text]. — Turpaeva 1993: 22 [text]. — Child 1998: 60 [text]. — Staples 2007: 80 [list], 92, fig. 4a–d.</p><p>Type material (not examined)</p><p>Holotype</p><p>SOUTHEASTERN INDIAN OCEAN • ♂; Western Australia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.916664&amp;materialsCitation.latitude=-31.083334" title="Search Plazi for locations around (long 114.916664/lat -31.083334)">west of Lancelin Island</a>; 31°5′ S, 116°55′ E ” [reinterpreted as 114°55′ E]; depth 113–122 m; 5 Feb. 1964; CSIRO stn 46; WAM 70-3953.</p><p>Paratypes</p><p>SOUTHEASTERN INDIAN OCEAN • 1 ♀; Western Australia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.066666&amp;materialsCitation.latitude=-32.55" title="Search Plazi for locations around (long 115.066666/lat -32.55)">west of Mandurah</a>; 32°33′ S, 115°4′ E; depth 110 m; 23 Nov. 1970; R/V Diamantina stn 4; WAM 71-1789 • 1 ♀ gravid; Western Australia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.78333&amp;materialsCitation.latitude=-30.633333" title="Search Plazi for locations around (long 114.78333/lat -30.633333)">southwest of Jurien Bay</a>; 30°38′ S, 114°47′ E; depth 110 m; 9 Dec. 1970; R/V Diamantina stn 108; WAM 71-1791 • 1 ♀; Western Australia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.8&amp;materialsCitation.latitude=-32.958332" title="Search Plazi for locations around (long 114.8/lat -32.958332)">northwest of Bunbury</a>; 32°57.5′ S, 114°48′ E; depth 122–139 m; 15 Mar. 1972; R/V Diamantina cruise DM 1/72 stn 8; USNM 149237 .</p><p>Distribution and depth range</p><p>Southwestern coast of Australia (type locality west of Lancelin Island) and Great Australian Bight. Minimal known depth range 110– 130 m.</p></div>	https://treatment.plazi.org/id/131BBD7A94329437FDE251DEFD78F962	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94329438FDE25483FD4BFB04.text	131BBD7A94329438FDE25483FD4BFB04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia dofleini (Loman 1911)	<div><p>Hedgpethia dofleini (Loman, 1911)</p><p>Fig. 2, Table 1</p><p>Colossendeis dofleini Loman, 1911: 4–5, pl. 1 figs 9–13. Colossendeis dofleini – Fukui 1919: 98. — Ohshima 1936: 867. — Hedgpeth 1949: 300, fig. 47a–d. — Utinomi 1951: 167; 1955: 33; 1959: 220; 1971: 337. — Losina-Losinsky &amp; Turpaeva 1958: 32 [key], tab. 4. — Stock 1958: tab. 1.</p><p>Rhopalorhynchus dofleini – Stock 1958: 116 [text], 118 [key], tab. 1; 1970: 9 [key].</p><p>Hedgpethia dofleini – Turpaeva 1973: 184 [key], 185 [text], tab. 2; 1993: 21 [text]. — Nakamura &amp; Child 1991: 63. — Stock 1991a: 166 [text].</p><p>Type material (not examined)</p><p>Holotype</p><p>NORTHWESTERN PACIFIC • ♀; Eastern Honshu, Sagami Bay, Misaki; F. Doflein leg.; hosting institution unknown.</p><p>Other material examined</p><p>SOUTHWESTERN INDIAN OCEAN • 1 spec.; Mozambique Channel, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.033333&amp;materialsCitation.latitude=-12.498333" title="Search Plazi for locations around (long 45.033333/lat -12.498333)">north of Mayotte, northeast of Northern Reef</a>; 12°29′53.9988″ S, 45°1′59.9988″ E; depth 450 m; 30 Mar. 1977; BENTHEDI stn DS71; MNHN-IU-2009-3005 .</p><p>Distribution and depth range</p><p>West Kyushu, Eastern Coast of Japan from South Kyushu to Hokkaido and including Sagami Bay (type locality), east of Kurile Islands. GBIF (2025) records extend further this distribution to the Aleutian Islands and Alaska, and to Mayotte. This latter record is particularly distant from the known distribution. I could examine part of this material (specimen MNHN-IU-2009-3005), but did not identify any obvious morphological difference from Loman’s (1911) illustrations. However, these are insufficient in details and the original material should also be reexamined. Minimal known depth range 40–869 m? Some previous authors (starting from Utinomi 1971) extend this depth range up to 20 m but I have not found any corresponding record in the literature.</p></div>	https://treatment.plazi.org/id/131BBD7A94329438FDE25483FD4BFB04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A943D9438FDEC569AFAD7F8D4.text	131BBD7A943D9438FDEC569AFAD7F8D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia eleommata Child 1998	<div><p>Hedgpethia eleommata Child 1998</p><p>Fig. 2, Table 1</p><p>Hedgpethia eleommata Child, 1998: 60–62, fig. 7.</p><p>Hedgpethia eleommata – Staples 2007: 80 [list].</p><p>Type material (not examined)</p><p>Holotype</p><p>SOUTWESTERN PACIFIC • sex unknown; New Zealand, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=171.76167&amp;materialsCitation.latitude=45.965" title="Search Plazi for locations around (long 171.76167/lat 45.965)">southeast of South Island, off Dunedin</a>; 45°57.9′ N, 171°45.7′ E; depth 1400–1415 m; 23 Feb. 1995; New Zealand Oceanographic Institute (NZOI) stn W460; NIWA, H-695.</p><p>Paratypes</p><p>SOUTWESTERN PACIFIC • 4 specs; same details as holotype; NZOI, P-1142 • 3 specs; same data as for holotype .</p><p>Distribution and depth range</p><p>East off South Island in New Zealand (type locality). Minimal known depth range 1415–1442 m.</p></div>	https://treatment.plazi.org/id/131BBD7A943D9438FDEC569AFAD7F8D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A943C9439FE6C5376FB83FD0C.text	131BBD7A943C9439FE6C5376FB83FD0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia elongata Takahashi, Dick & Mawatari 2007	<div><p>Hedgpethia elongata Takahashi, Dick &amp; Mawatari, 2007</p><p>Fig. 2, Table 1</p><p>Hedgpethia elongata Takahashi et al., 2007: 70–72, fig. 4.</p><p>Type material (not examined)</p><p>Holotype</p><p>PHILIPPINE SEA • ♂; Okinawa, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.7&amp;materialsCitation.latitude=26.825" title="Search Plazi for locations around (long 127.7/lat 26.825)">northeast of Ie Island</a>; 26°49.50′ N, 127°42′ E; depth 199–202 m; 23 May 2003; TRV Toyoshio-maru; ZIHU 03171.</p><p>Distribution and depth range</p><p>Only known from Ie Island in Okinawa (type locality), depth range 199– 202 m.</p></div>	https://treatment.plazi.org/id/131BBD7A943C9439FE6C5376FB83FD0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A943C9439FD9B5092FC6FFABB.text	131BBD7A943C9439FD9B5092FC6FFABB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia filamentus Staples 2019	<div><p>Hedgpethia filamentus Staples, 2019</p><p>Fig. 2, Table 1</p><p>Hedgpethia filamentus Staples, 2019: 431–433, 435 [text], fig. 15, pl. 5e–f.</p><p>Type material (examined)</p><p>Holotype</p><p>SOUTHWESTERN INDIAN OCEAN • sex unknown; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=57.272335&amp;materialsCitation.latitude=-32.714333" title="Search Plazi for locations around (long 57.272335/lat -32.714333)">Atlantis bank</a>; 32°42.86′ S, 57°16.34′ E; depth 750 m; 14 Dec. 2011; R.R.S. James Cook voyage JC066 stn 8.29; NHMUK ZOO-2018.38.</p><p>Paratypes</p><p>SOUTHWESTERN INDIAN OCEAN • 4 specs; same data as for holotype; NHMUK ZOO-2018.39 • 1 spec.; same data as for holotype; NHMUK ZOO-2018.40 .</p><p>Distribution and depth range</p><p>Only known from Atlantis bank (type locality), depth 750 m.</p></div>	https://treatment.plazi.org/id/131BBD7A943C9439FD9B5092FC6FFABB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A943C9439FE595728FB7AF864.text	131BBD7A943C9439FE595728FB7AF864.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia magnirostris Arnaud & Child 1988	<div><p>Hedgpethia magnirostris Arnaud &amp; Child, 1988</p><p>Fig. 2, Table 1</p><p>Hedgpethia magnirostris Arnaud &amp; Child, 1988: 123 [list], 153–155, fig. 7.</p><p>Hedgpethia magnirostris – Stock 1991a: 166 [text].</p><p>Type material (not examined)</p><p>Holotype</p><p>SOUTHWESTERN INDIAN OCEAN • ♂; northeast of South Africa, off northern coast of KwaZulu-Natal (including former KwaZulu = Zululand in Arnaud &amp; Child 1988); 28°31.7′ S, 32°34′ E; depth 680 m; 24 May 1976; R/V Meiring Naudé stn SM103; SAM-18579.</p><p>Paratype</p><p>SOUTHWESTERN INDIAN OCEAN • 1 ♀; same data as for holotype; SAM-A19580 .</p><p>Distribution and depth range</p><p>Only known from the northern coast of KwaZulu-Natal (type locality), depth 680 m.</p></div>	https://treatment.plazi.org/id/131BBD7A943C9439FE595728FB7AF864	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A943F943AFDC55376FBFBFC3D.text	131BBD7A943F943AFDC55376FBFBFC3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia nasica Child 1994	<div><p>Hedgpethia nasica Child, 1994</p><p>Fig. 2, Table 1</p><p>Hedgpethia nasica Child, 1994: 16–18, fig. 5.</p><p>Type material (not examined)</p><p>Holotype</p><p>NORTHEASTERN PACIFIC • sex unknown; California, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.11667&amp;materialsCitation.latitude=34.75" title="Search Plazi for locations around (long -123.11667/lat 34.75)">west of Point Arguelo</a>; 34°45′ N, 123°7′ W; depth ca 4100 m; 24 Jun 1991; cruise Pulse VII stn 721M; USNM234637.</p><p>Paratype</p><p>NORTHEASTERN PACIFIC • 1 spec.; California, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.11667&amp;materialsCitation.latitude=34.816666" title="Search Plazi for locations around (long -123.11667/lat 34.816666)">west of Point Arguelo</a>; 34°49′ N, 123°7′ W; depth ca 4100 m; 25 Jun. 1989; cruise Pulse I stn 124; USNM234638 • 1 ♀; California, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.01667&amp;materialsCitation.latitude=34.716667" title="Search Plazi for locations around (long -123.01667/lat 34.716667)">west of Point Arguelo</a>; 34°43′ N, 123°1′ W; depth ca 4100 m; 23 Oct. 1990; cruise Pulse V stn 505M; USNM234639 • 1 sp.; California, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.05&amp;materialsCitation.latitude=34.7" title="Search Plazi for locations around (long -123.05/lat 34.7)">west of Point Arguelo</a>; 34°42′ N, 123°3′ W; depth ca 4100 m; 25 Oct. 1991; cruise Pulse X stn 1017M; USNM234640 .</p><p>Distribution and depth range</p><p>Only known from west of Point Arguelo (type locality), depth ca 4100 m.</p></div>	https://treatment.plazi.org/id/131BBD7A943F943AFDC55376FBFBFC3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A943F943AFDFC51A2FDAFF9F3.text	131BBD7A943F943AFDFC51A2FDAFF9F3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia shalei Staples 2019	<div><p>Hedgpethia shalei Staples, 2019</p><p>Fig. 2, Table 1</p><p>Hedgpethia shalei Staples, 2019: 433–435, fig. 16, pl. 5g –h.</p><p>Type material (examined)</p><p>Holotype</p><p>SOUTHWESTERN INDIAN OCEAN • sex unknown; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.454&amp;materialsCitation.latitude=-37.946583" title="Search Plazi for locations around (long 50.454/lat -37.946583)">Middle of What Seamount</a>; 37°56.795′ S, 50°27.240′ E [recovered from mission report]; depth 1414 m; 2 Dec. 2011; R.R.S. James Cook voyage JC066 stn 6.7; NHMUK ZOO-2018.41.</p><p>Paratype</p><p>SOUTHWESTERN INDIAN OCEAN • 1 spec.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=42.9095&amp;materialsCitation.latitude=-41.371834" title="Search Plazi for locations around (long 42.9095/lat -41.371834)">Coral Seamount</a>; 41°22.31′ S, 42°54.57′ E; depth 732 m; 20 Nov. 2011; R.R.S. James Cook voyage JC066 stn 4.38; NHMUK ZOO-2018.42 .</p><p>Distribution and depth range</p><p>Southwest Indian Ocean seamounts: Middle of What Seamount (type locality) and Coral Seamount. Known depth range 687–1414 m.</p></div>	https://treatment.plazi.org/id/131BBD7A943F943AFDFC51A2FDAFF9F3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A943F943BFE1F54F0FB87FEF0.text	131BBD7A943F943BFE1F54F0FB87FEF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia spinosa Takahashi, Kajihara & Mawatari 2012	<div><p>Hedgpethia spinosa Takahashi, Kajihara &amp; Mawatari, 2012 Fig. 2, Table 1</p><p>Hedgpethia spinosa Takahashi et al., 2012: 70–72, fig. 1.</p><p>Type material (not examined)</p><p>Holotype</p><p>PHILIPPINE SEA • ♂; Kagoshima Prefecture, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.634&amp;materialsCitation.latitude=30.148333" title="Search Plazi for locations around (long 130.634/lat 30.148333)">south of Yaku Island (= Yakushima Island)</a>; 30°8.9′ N, 130°38.04′ E; depth 197–207 m; 26 May 2005; TRV Toyoshio-maru; ZIHU 3335.</p><p>Distribution and depth range</p><p>Only known from south of Yaku Island (type locality), depth range 197– 207 m.</p></div>	https://treatment.plazi.org/id/131BBD7A943F943BFE1F54F0FB87FEF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A943E943BFDF853F1FE76FB56.text	131BBD7A943E943BFDF853F1FE76FB56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia tibialis Stock 1991	<div><p>Hedgpethia tibialis Stock, 1991</p><p>Fig. 2, Table 1</p><p>Hedgpethia tibialis Stock, 1991a: 166, figs 29–30.</p><p>Hedgpethia tibialis – Turpaeva 1993: 22 [text]. — Staples 2007: 80 [list]. — Sabroux et al. 2023a: 3 [text], fig. 2 [phylogeny], tab. 1.</p><p>Type material (examined)</p><p>Holotype</p><p>CORAL SEA • sex unknown; off New Caledonia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=163.36166&amp;materialsCitation.latitude=-18.88" title="Search Plazi for locations around (long 163.36166/lat -18.88)">Grand Passage</a>; 18°52.80′ S, 163°21.7′ E; depth 550 m; 19 Sep. 1985; MUSORSTOM 4 stn CP194; MNHN-IU-2007-4581 (formerly part of lot MNHNPy677).</p><p>Paratypes</p><p>CORAL SEA • 1 spec.; same data as for holotype; MNHN-IU-2011-5711 (formerly part of lot MNHNPy677) • 1 spec.; same data as for holotype; MNHN-IU-2011-5712 (formerly part of lot MNHN-Py677) • 1 spec.; same data as for holotype; MNHN-IU-2007-4582 (formerly part of lot MNHN-Py677) • 1 spec.; same data as for holotype; NL, ZMA.PYC.3334 • 1 spec.; southeast of New Caledonia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=167.28334&amp;materialsCitation.latitude=-22.928333" title="Search Plazi for locations around (long 167.28334/lat -22.928333)">southwest of Isle of Pins</a>; 22°55.7′ S, 167°17′ E; depth 520 m; 28 Sep. 1985; MUSORSTOM 4 stn CP215; MNHN-IU-2007-4634 (formerly MNHN-Py790) .</p><p>Distribution and depth range</p><p>Recorded from both northwest (type locality) and southeast extremities of New Caledonia. Known depth range 368– 550 m.</p></div>	https://treatment.plazi.org/id/131BBD7A943E943BFDF853F1FE76FB56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A943E943EFDC0574FFCB6F86A.text	131BBD7A943E943EFDC0574FFCB6F86A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia nosferatu Sabroux 2025	<div><p>Hedgpethia nosferatu sp. nov.</p><p>urn:lsid:zoobank.org:act: B4C8A512-51FE-4EBF-82A3-21E957D7BBFC</p><p>Figs 1–2, Table 1</p><p>Hedgpethia sp. – Sabroux et al. 2023a [phylogeny]: figs 2–3, tab. 1.</p><p>Etymology</p><p>In apposition. Named after Nosferatu, a popular vampire character appearing for the first time in ‘Nosferatu, eine Symphonie’ (1922), in reference to the somewhat bat-wing-like outline of the femorae.</p><p>Type material</p><p>Holotype</p><p>SOUTHWEST PACIFIC • ♀; south of New Caledonia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=167.65666&amp;materialsCitation.latitude=-22.94383" title="Search Plazi for locations around (long 167.65666/lat -22.94383)">south-southeast of Isle of Pines</a>; 22°56′37.7952″ S, 167°39′23.9832″ E; depth 310–403 m; 23 Aug. 2016; KANACONO stn DW4745; GenBank nos: OP985938 (mitochondrial genome), OQ065619 (18S); MNHN-IU-2016-6868.</p><p>Description (holotype, ♀ (MNHN-IU-2016-6868))</p><p>MEASUREMENTS (mm). Trunk + cephalon length 2.2; trunk width at second trunk segment (lateral processes included) 1.1; proboscis length 3.8; abdomen length 0.09; femur length 4.7; tibia 1 length 7.5; tibia 2 length 4.1; tarsus length 0.9; propodus length 0.8; main claw length 0.4.</p><p>Trunk fully segmented, compact. Trunk segments 1–3 posterior margin flaring and carrying one small dorsomedian pointed tubercle. Ocular tubercle medium-sized, pointed at tip, about 1.6 × as tall as base width, carrying four pigmented eyes. Lateral sense organs inconspicuous or absent. Palp insertion on cephalon at end of short anterolateral projection aside proboscis, base of which seemingly segmented. Oviger insertion ventral, just before first trunk segment lateral processes, individualised as short ventral projection. Lateral processes shorter than wide, without ornamentation, separated by less than own diameter.</p><p>Proboscis about 1.7 × as long as whole body. Base constricted, not as basal stalk. Proboscis inflating from base to mid-length with gentle constriction about proximalmost third. Distalmost half tapering, distalmost third almost pipette-like. Tip rounded.</p><p>Abdomen ventralised, not visible dorsally, directed diagonally downward, not reaching beyond lateral processes. Anus at tip.</p><p>Chelifore absent.</p><p>Palp 9-articled. Article 1 shortest, shorter than wide. Article 2 longest, curved. Article 3 about twice as long as wide. Article 4 long, slightly more than half as long as article 2. Article 5 short, about twice as long as wide, more than 5 × as short as article 4. Article 6 thrice as long as wide, 1.7 × as long as article 5. Article 7 about as long as article 6. Articles 8 and 9 about 0.9 × as long as article 6. Articles 2 to 4 sparsely setose, articles 5 to 9 very setose ventrally. Individualised medium size setae carried on dorsodistal margin of articles 2 and 3.</p><p>Oviger 10-articled plus terminal claw. Articles 1 and 3 about as long as wide. Article 2 slightly longer. Articles 4 and 6 longest, subequal. Article 5 short, about 5 × as small as article 4. Articles 7–10 forming strigilis, article 7 longest, articles 8 and 9 subequal, article 10 shortest. Strigilis spines spatulate, denticulated distally, distributed in fields. Distalmost ventral spine of article 10 curved, forming subchela with terminal claw. Terminal claw wedge-like, with ventral cutting edge.</p><p>Legs coxae 1–3 about as short as wide. Coxa 1 dorsally ornamented with dorsal spinules. Femur widest, with seven more or less elevated mediodorsal conical tubercles carrying one seta each, three of which on proximal half about half as high as femur width. Tibia 1 longest, 1.6 × as long as femur, carrying dorsally short setae, one slightly longer on distal margin. Tibia 2 slightly longer than half tibia 1, 0.9 × as long as femur, with one long seta dorsodistally and one strong spine ventrodistally. Tarsus about 0.2 × as long as tibia 2, with numerous ventral spines and more scarcely distributed dorsal spines. Propodus slightly shorter than tarsus, propodal sole carrying numerous spines. Main claw about half as long as propodus. Auxiliary claws absent.</p><p>Remarks</p><p>The femorae of H. nosferatu sp. nov. readily distinguish it from all other species in the genus. Tibia 1 is also particularly long for the genus, about 1.6 × as long as the femur. Most species have this ratio to 1.2 (e.g., H. bicornis, H. brevitarsis, H. chitinosa, H. shalei) or below. Hedgpethia eleommata has this ratio to 1.3, H. articulata 1.4, H. magnirostris 1.5, and H. tibialis 1.6 too. Hedgpethia nosferatu differs from H. magnirostris by its taller ocular tubercle; from H. articulata by the wider propodus and tarsus; from H. eleommata by the propodus / tarsus length ratio, and the shape of the proboscis; from H. tibialis, which is geographically close (New Caledonia), by the tarsus / propodus ratio.</p><p>Hedgpethia nosferatu is morphologically closest to H. chitinosa . Besides the above-mentioned characters, H. nosferatu differs from this species by the shape of the proboscis, and the presence of spines on the sole of the propodus. The main claw also tends to be longer relative to the propodus in H. chitinosa, although this character seems to present important variations (Nakamura, 1987).</p><p>Sea spider diversity in New Caledonia included so far 66 nominal species listed by Bamber (2007b). Among them, 36 appear so far as endemic (Bamber 2007b). Hedgpethia nosferatu is therefore the 67 th pycnogonid species recorded from New Caledonia, and potentially the 37 th endemic species. This is also the second species of Hedgpethia recorded from New Caledonia, after H. tibialis that was recorded north and south of the island (Stock 1991a). The new species holotype is likely a female given the wide opening of the gonopores ventrally on coxae 2.</p><p>Geographic and bathymetric distribution</p><p>There are two main geographic components in the distribution of Hedgpethia as recorded in the literature (Fig. 2):</p><p>A Northern Pacific distribution (Fig. 2B) represented by seven species ( H. bicornis, H. brevitarsis, H. californica, H. chitinosa, H. elongata, H. nasica, H. spinosa) distributed in an arch from Okinawa to California. Four of these species ( H. brevitarsis, H. californica, H. elongata, H. spinosa) were found between 0 and 1000 m (Table 1). The distribution of H. bicornis extends slightly deeper, down to 1100 m in the Okhotsk Sea (type locality; Dudnik 2025); it was found as shallow as 303 m in the Kuril Islands, and possibly as shallow as 64 m along the southeastern coast of Kamchatka (Losina-Losinsky 1961). Hedgpethia chitinosa was also mostly sampled within the 0–1000 m range, but two sampling events for this species are recorded to 3939 m (Bering Sea; Hilton 1943) and ca 6500 m (south off Commander Islands; Dudnik 2025). Hedgpethia nasica was found only once, at ca 4100 m deep off the California coast.</p><p>A Southern Indopacific distribution (Fig. 2A, C) represented by eight species ( H. calva, H. dampieri, H. eleommata, H. filamentus, H. magnirostris, H. nosferatu, H. shalei, H. tibialis) distributed from the eastern coast of South Africa to off New Caledonia and New Zealand, six of these species ( H. calva, H. dampieri, H. filamentus, H. magnirostris, H. nosferatu, H. tibialis) were sampled uniquely above 1000 m deep. Two species have their bathymetry extending even deeper: H. eleommata was sampled around 1400–1450 m southeast off New Zealand; H. shalei was sampled on the Southwest Indian Ridge at 687 and 1414 m deep. Hedgpethia filamentus and H. shalei are the two species in this genus presently known from seamounts (Atlantis Bank and Middle of What Seamount; Staples 2019).</p><p>Hedgpethia articulata and H. dofleini spread between these two geographic groups: H. articulata has the widest know bathymetric distribution and was sampled in Indonesia (ca 2000 m deep), the Bering Sea (ca 4000 m deep) and the Gulf of Alaska (3620 m deep); H. dofleini was mostly sampled off both sides of Japan (from Hokkaido to Kyushu; ca 40–900 m deep) and in the Aleutian Islands (ca 80-650 m deep), but one specimen recorded on GBIF (2025) was sampled off Mayotte in the Comoro Islands (450 m deep).</p><p>Outside these distributions, H. caudata is the only species known off the Pacific coast of South America (Peru; approximatively 3000 m deep), and H. atlantica is the sole species recorded in the Northeast Atlantic from around 200 m (Josephine Bank; Stock 1970) to below 1000 m deep (Galicia Bank; Stock 1991b); in the Gibraltar Strait (135 m deep; Stock 1987), and in the Western Mediterranean (French Mediterranean coast at a depth of 100 m; Arnaud 1987).</p></div>	https://treatment.plazi.org/id/131BBD7A943E943EFDC0574FFCB6F86A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
131BBD7A94259422FF1857D4FA90FE25.text	131BBD7A94259422FF1857D4FA90FE25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedgpethia Turpaeva 1973	<div><p>Identification key to worldwide species of Hedgpethia Turpaeva, 1973</p><p>Below is an identification key for all Hedgpethia, the backbone of which is modified from the key of Stock (1970) for the species of the “ articulatum -group” Rhopalorhynchus [ = articulatum section in Stock 1958], that were later assigned to Hedgpethia .</p><p>1. Ocular tubercle rudimentary. Eyes absent ........................................................................................ 2</p><p>– Ocular tubercle and eyes well developed .......................................................................................... 3</p><p>2. Tibia 1 longer than tibia 2 or femur. Abdomen a small ventral knob ... H. articulata (Loman, 1908)</p><p>– Tibia 1 shorter than tibia 2 or femur. Abdomen about twice as long as wide .. H. nasica Child, 1994</p><p>3. Ventral margin of tarsus and propodus without spines ..................................................................... 4</p><p>– Ventral margin of tarsus and propodus with a closely set row of spines .......................................... 7</p><p>4. Ocular tubercle a tall cone, at least thrice as high as base width. Oviger’s article 10 with a large spine opposing the terminal claw, forming a subchelate structure ........... H. californica (Hedgpeth, 1939)</p><p>– Ocular tubercle not higher than twice as base width. No enlarged oviger spine, an no subchelate structure on the distal oviger articles ................................................................................................ 5</p><p>5. Tarsus slightly longer than propodus ........................................................ H. dofleini (Loman, 1911)</p><p>– Tarsus shorter than propodus ............................................................................................................ 6</p><p>6. Ocular tubercle conical. Legs articles smooth. Proboscis without narrowing of its inflated part around proximalmost third .......................................................................................... H. calva Arango, 2009</p><p>– Ocular tubercle rounded with tapering tip, legs spinous. Proboscis inflated section narrowed at proximalmost third .................................................................................... H. dampieri (Child, 1975)</p><p>7. Tarsus distinctly shorter than propodus ............................................................................................ 8</p><p>– Tarsus as long as to distinctly longer than propodus ...................................................................... 14</p><p>8. Main claw about as long as propodus ...................................................... H. eleommata Child, 1998</p><p>– Main claw less than half as long as the propodus ............................................................................. 9</p><p>9. Proboscis cylindrical or tubular in shape, abdomen horizontal……………. H. caudata Turpaeva, 1993</p><p>– Proboscis inflating medially, constricted at base, tapering distally ................................................ 10</p><p>10. Proximal constriction of the proboscis developed as a long stalk. Ocular tubercle about 4 × as high as base width ............................................................................................. H. atlantica (Stock, 1970)</p><p>– Proximal constriction of the proboscis not developed as a stalk. Ocular tubercle no more than thrice as tall as base width .........................................................................................................................11</p><p>11. Posterior margin of trunk segments 1–3 carrying one rounded dorsomedian tubercle each ............... ............................................................................................... H. brevitarsis (Losina-Losinsky, 1958)</p><p>– Posterior margin of trunk segments 1–3 flaring, without dorsomedian tubercle ............................ 12</p><p>12. Proboscis flask-shaped, almost pipette-like distally; ocular tubercle a low cone, about as long as base width ............................................................................................................... H. tibialis Stock, 1991</p><p>– Proboscis pyriform, gently tapering from its widest point to the tip; ocular tubercle a cone, about twice as tall as base width ............................................................................................................... 13</p><p>13. Space separating lateral processes 2–4 wider than their own diameter, proximal constriction of the proboscis forming a short stalk with parallel sides ................................ H. filamentus Staples, 2019</p><p>– Space separating lateral processes 2–4 smaller than their own diameter, proximal constriction not forming a stalk ............................................................................................... H. shalei Staples, 2019</p><p>14. Anterior margin of cephalic segment with two tubercles ............................................................... 15</p><p>– Anterior margin of cephalic segment smooth ................................................................................. 17</p><p>15. Main claw half as long as propodus ................... H. spinosa Takahashi, Kajihara &amp; Mawatari, 2012</p><p>– Main claw nearly as long as propodus ............................................................................................ 16</p><p>16. Posterior margin of trunk segments 1–3 carrying one pointy dorsomedian tubercle each ................. .............................................................................................................. H. bicornis (Turpaeva, 1958)</p><p>– Posterior margin of trunk segments 1–3 flaring, without dorsomedian tubercle ................................ .............................................................................................. H. magnirostris Arnaud &amp; Child, 1988</p><p>17. Proximal constriction of the proboscis as a long stalk, about as long as 2 nd trunk segment ............... ................................................................................. H. elongata Takahashi, Dick &amp; Mawatari, 2007</p><p>– Proximal constriction of the proboscis as a very short stalk or no stalk at all ................................ 18</p><p>18. Femur dorsal surface smooth except for short setae, tibia 1 about 1.2 × as long as femur ................ ................................................................................................................. H. chitinosa (Hilton, 1943)</p><p>– Femur dorsal surface ornamented with conical tubercles, height of the tallest about half of femur diameter; tibia 1 about 1.6 × as long as femur .................................................. H. nosferatu sp. nov.</p></div>	https://treatment.plazi.org/id/131BBD7A94259422FF1857D4FA90FE25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sabroux, Romain	Sabroux, Romain (2025): The genus Hedgpethia Turpaeva, 1973 (Pycnogonida: Colossendeidae): a review and a new species. European Journal of Taxonomy 1026: 147-170, DOI: 10.5852/ejt.2025.1026.3111, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3111/13875
