identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
002A9E7D23C651B8A397282EACAFF600.text	002A9E7D23C651B8A397282EACAFF600.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillinus Casey 1918	<div><p>Genus Anillinus Casey, 1918</p><p>Anillinus Casey, 1918: 167. Type species: Anillus (Anillinus) carolinae Casey, 1918, by original designation.</p><p>Micranillodes Jeannel, 1963a: 57. Synonymy established by Bousquet (2012: 699) and confirmed by Sokolov et al (2014: 83). Type species: Micranillodes depressus Jeannel, 1963a, by original designation.</p><p>Troglanillus Jeannel, 1963b: 147. Synonymy established by Barr (1995: 240). Type species: Troglanillus valentinei Jeannel, 1963b, by original designation.</p></div>	https://treatment.plazi.org/id/002A9E7D23C651B8A397282EACAFF600	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sokolov, Igor M.	Sokolov, Igor M. (2021): Two new species of the genus Anillinus Casey (Coleoptera, Carabidae, Anillini) from the southern United States. ZooKeys 1016: 63-76, DOI: http://dx.doi.org/10.3897/zookeys.1016.61397, URL: http://dx.doi.org/10.3897/zookeys.1016.61397
4F0B7AC4707B51B19461F7B6FB837A01.text	4F0B7AC4707B51B19461F7B6FB837A01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillinus felicianus Sokolov 2021	<div><p>Anillinus felicianus sp. nov. Figs 1, 2, 5</p><p>Type material.</p><p>Holotype male (NMNH), dissected, labeled: \ USA-LA: West Feliciana Par., ~4mi SW Jackson at 30.794°N, 91.254°W, mixed pine-hardwood forest, soil washing/berlese Sokolov I.M. 19-25 Apr 2018 \. Paratype: one female, labeled as holotype (NMNH); three males and three females labeled: \USA, LA, W. Feliciana Par. Feliciana Preserve Natural Area, Orange Trail, 30.792649°N, 91.253382°W, 29 Oct. 2015 \ Soil washing in hardwood forest B.E. Owens and C.E. Carlton \(LSAM).</p><p>Etymology.</p><p>The name of this species is a Latinized adjective based on the name of Feliciana Preserve, in which this species occurs. Feliciana Preserve is a privately owned nature reserve created by several professors of the Louisiana State University (principal developer Dr. Dorothy Prowell) and located in the Tunica Hills area of southeastern Louisiana.</p><p>Type locality.</p><p>USA, Louisiana, West Feliciana Parish, Tunica Hills, 4 mi SW of Jackson.</p><p>Recognition.</p><p>Adults of A. felicianus can be distinguished from those of other subterranean members of Anillinus by the combination of smooth pronotum and completely microsculptured head. Males of A. felicianus can be also distinguished from those of other congeners by the structure of the median lobe.</p><p>Description.</p><p>Moderate-sized for genus (ABL 1.59-1.68 mm, mean 1.64 ± 0.064 mm, n = 2).</p><p>Habitus: Body form (Fig. 1A) moderately convex, subparallel, elongate (WE/ABL 0.36 ± 0.001), head moderately large in comparison to pronotum (WH/WPm 0.77 ± 0.008), pronotum of moderate width in comparison to elytra (WPm/WE 0.82 ± 0.006).</p><p>Integument: Body color brunneo-rufous, appendages testaceous. Microsculpture (Fig. 1B-D) present across all head and elytra, where it is represented by isodiametric polygonal sculpticells; and absent from disc of pronotum. Body surface shiny, surface sparsely and finely punctate, covered with sparse, yellowish, short setae. Vestiture of elytra (Fig. 1D) short (~0.3 length of discal setae). Elytral chaetotaxy typical for Anillinus, umbilicate series of type A (sensu Jeannel 1963a and Giachino and Vailati 2011): nine setae arranged in three groups, subhumeral (3+1), middle (2) and subapical (1+2), with the last two (8th and 9th) pores of umbilicate series “geminate”, much closer to each other than 7th pore is to 8th; in subapical group the 8th pore is the longest.</p><p>Prothorax: Pronotum (Fig. 1C) moderately convex, of moderate size (LP/LE 0.41 ± 0.010) and moderately transverse (WPm/LP 1.26 ± 0.009), with lateral margins almost rectilinearly and moderately constricted posteriorly (WPm/WPp 1.26 ± 0.009). Anterior angles indistinct, posterior angles slightly obtuse (105-110°). Width between posterior angles equals width between anterior angles (WPa/WPp 0.97 ± 0.001). Basal margin slightly concave in middle.</p><p>Scutellum: Externally visible, triangular, with pointed apex.</p><p>Elytra: Slightly convex, of average length (LE/ABL 0.58 ± 0.012) and width (WE/LE 0.63 ± 0.015) for genus, with traces of 4-5 striae. Humeri distinct, rounded, in outline forming obtuse angle with longitudinal axis of body. Lateral margins subparallel in middle, slightly convergent at basal fourth, evenly rounded to apex at apical third, with shallow subapical sinuation. Basal margination distinct.</p><p>Legs: Protarsi of male with moderately dilated tarsomere 1. Profemora moderately swollen. Metafemora unmodified.</p><p>Male genitalia: Median lobe (Fig. 2A) of aedeagus anopic, slightly arcuate and moderately twisted. Shaft dilated in apical two-thirds, with moderately elongate apex, slightly tapered to rounded tip. Ventral margin of median lobe straight, not enlarged, with few poriferous canals, curved downward close to basal orifice. Endophallus with dorsal copulatory sclerites fused to form slightly curved dorsal blade-like structure and straight, apically-pointed ventral plate of moderate length. Spines and scaled membranous folds of endophallus absent. Left paramere (Fig. 2B) of shape common in genus, paramere apex with four setae getting longer toward apex. Right paramere (Fig. 2C) of moderate length, bearing three long setae of approximately same length as paramere.</p><p>Female genitalia: Spermatheca (Fig. 2E) unsclerotized, shaped as a question mark, with sharply dilated bean-like distal part. Length of spermathecal gland shorter than length of spermatheca. Spermathecal duct uncoiled. Gonocoxites and laterotergite as in Fig. 2D. Gonocoxite 2 falciform, more than 2 times longer than wide basally, with acute ensiferous setae. Laterotergite with 7-8 setae.</p><p>Geographic distribution.</p><p>This species is known only from the type locality in the Tunica Hills area of West Feliciana Parish, Louisiana (Fig. 5, green circle).</p><p>Habitat.</p><p>All specimens of this species were collected from loess soil samples using soil washing techniques (Southwood and Henderson 2000). These samples were taken under forest canopy on the top of a hill between two gullies with temporal creeks. Anillinus felicianus is a true endogean species and has never been found in litter samples.</p><p>Relationships.</p><p>The species belongs to group V of endogean species (sensu Sokolov et al. 2004), characterized by a combination of a completely microsculptured head and smooth disc of pronotum. Within this group, A. felicianus is most closely related to the endogean A. sinuaticollis Jeannel (Jeannel 1963b), differing from the latter by a smooth - not microsculptured - base of the pronotum, a bigger size, and details of the pronotum shape and body proportions. The range of A. sinuaticollis is confined to Roane County of Tennessee (Fig. 5, green area with vertical line pattern); thus, its range lies about 500 miles north of the type locality of A. felicianus .</p></div>	https://treatment.plazi.org/id/4F0B7AC4707B51B19461F7B6FB837A01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sokolov, Igor M.	Sokolov, Igor M. (2021): Two new species of the genus Anillinus Casey (Coleoptera, Carabidae, Anillini) from the southern United States. ZooKeys 1016: 63-76, DOI: http://dx.doi.org/10.3897/zookeys.1016.61397, URL: http://dx.doi.org/10.3897/zookeys.1016.61397
588729DA5AFB535A9D11D17A06E306A6.text	588729DA5AFB535A9D11D17A06E306A6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillinus relictus Sokolov 2021	<div><p>Anillinus relictus sp. nov. Figs 3, 4A-C, 5</p><p>Type material.</p><p>Holotype, one male (CMNH), dissected, labeled: \ ALABAMA: Blount Co., Tidwell Hollow Nature Trail east of Oneonta. T. N. King April 1 1972 \ 4/1/72 o [handwritten] \ THOMAS C. BARR COLLECTION 2011 Acc. No. 38,014 \. Paratype, one female, labeled as holotype (CMNH).</p><p>Etymology.</p><p>The specific epithet is a Latin adjective, Anillinus relictus (from Latin: abandoned, forsaken), in the masculine form, and refers to the geographical isolation of this species from its morphologically closest congeners, as it is believed to be the only remaining eastern representative of an ancestral group once more widespread.</p><p>Type locality.</p><p>USA, Alabama, Blount County, the Oneonta area.</p><p>Recognition.</p><p>Adults of A. relictus can be distinguished from those of other members of eastern Anillinus by the combination of the large size, completely microsculptured head and pronotum, and, especially, by the long elytral vestiture equals to 0.5-0.7 of length of discal elytral setae.</p><p>Description.</p><p>Large-sized for genus (ABL 2.29-2.42 mm, mean 2.36 ± 0.092 mm, n = 2).</p><p>Habitus: Body form (Fig. 3A) moderately convex, ovoid (WE/ABL 0.39 ± 0.012), head of average proportions for genus (WH/WPm 0.71 ± 0.023), pronotum moderately narrow in comparison to elytra (WPm/WE 0.77 ± 0.007).</p><p>Integument: Body color piceo-brunneus, appendages testaceous. Microsculpture (Fig. 3B-D) present across all head, pronotum, and elytra, where it is represented by isodiametric polygonal sculpticells. Body surface shiny, surface sparsely and finely punctate, covered with moderately dense, yellowish, long setae. Vestiture of elytra (Fig. 3D) long (0.5-0.7 length of discal setae). Elytral chaetotaxy typical for Anillinus, umbilicate series of type A (sensu Jeannel 1963a and Giachino and Vailati 2011).</p><p>Prothorax: Pronotum (Fig. 3C) moderately convex, of moderate size (LP/LE 0.40 ± 0.003) and moderately transverse (WPm/LP 1.26 ± 0.028), with lateral margins almost rectilinearly and slightly constricted posteriorly (WPm/WPp 1.19 ± 0.013). Anterior angles slightly prominent, posterior angles nearly rectangular (95-100°). Width between posterior angles much greater than between anterior angles (WPa/WPp 0.87 ± 0.038). Basal margin slightly concave in middle.</p><p>Scutellum: Externally visible, triangular, with rounded apex.</p><p>Elytra: Narrowly depressed along suture, of average length (LE/ABL 0.59 ± 0.005) and width (WE/LE 0.66 ± 0.026) for genus, with traces of 6-7 striae. Humeri distinct, rounded, in outline forming obtuse angle with longitudinal axis of body. Lateral margins subparallel in middle, slightly convergent at basal fifth, evenly rounded to apex at apical fourth, with shallow subapical sinuation. Basal margination distinct.</p><p>Legs: Protarsi of male with moderately dilated tarsomere 1. Profemora moderately swollen. Metafemora unmodified.</p><p>Male genitalia: Median lobe of aedeagus (Fig. 4A) anopic, slightly arcuate and slightly twisted. Basal orifice comparatively short for the genus. Shaft with long subparallel basal part, slightly dilating in apical third. Apical part with enlarged apex in form of rounded parallelogram. Dorsal margin slightly convex and strongly sclerotized at middle. Ventral margin curved near middle, where it is suddenly enlarged right before the apex. Endophallus with dorsal sclerite in form of a semicircular filament-like structure with short basal prolongations. Ventral sclerite located at apical orifice, in form of golf gap wedge plate. Dorsal scaly membranous field present at middle of dorsal sclerite. Enlarged apical area of median lobe with a dark spine-like structure (Fig. 4A, ss). Left paramere (Fig. 4B) modified, with long, subparallel apical half of moderate width with one seta at angulate tip, basally with strong concave keel (bk), and thick basal processes of different length. Right paramere (Fig. 4C) of moderate length, with eight setiferous pores bearing only three long setae (several others might be broken), which are shorter than length of paramere.</p><p>Female genitalia: Spermatheca not investigated. Ovipositor sclerites standard for genus with falciform gonocoxite 2 bearing two ensiform setae. Laterotergite with 8-9 setae.</p><p>Geographic distribution.</p><p>This species is known only from the type locality in Blount County, Alabama (Fig. 5, red circle).</p><p>Habitat.</p><p>The label does not contain any habitat information. Presumably, this species is not a cavernicolous species.</p><p>Relationships.</p><p>Based on the structure of the median lobe, A. relictus is a sister species to the endogean A. sinuatus (Jeannel) (Jeannel 1963a). The latter species is known to occur in Bexar County, Texas, where it was documented by a small series of three specimens extracted from the soil during surveys in peach orchards (Jeannel 1963a; Sokolov et al. 2014). The range of A. sinuatus (Fig. 5, black cross) is situated approximately 770 miles southwest of the type locality of A. relictus .</p></div>	https://treatment.plazi.org/id/588729DA5AFB535A9D11D17A06E306A6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sokolov, Igor M.	Sokolov, Igor M. (2021): Two new species of the genus Anillinus Casey (Coleoptera, Carabidae, Anillini) from the southern United States. ZooKeys 1016: 63-76, DOI: http://dx.doi.org/10.3897/zookeys.1016.61397, URL: http://dx.doi.org/10.3897/zookeys.1016.61397
