taxonID	type	description	language	source
1269837C3D07FFC92B13FAE73BBFF96E.taxon	diagnosis	DIAGNOSIS (modified from Engel, 2000): ♀: Labral distal process broadly triangular; lateral teeth absent; labral basal elevation absent. Mandible simple, large, bladelike (Figs. 2, 12, 32, 35, 45). Prementum greatly elongate, narrowed; galeal apex acute; galeal comb absent; galeal base extending to stipital base. Hypostomal ridge carinate, becoming weakly lamellate posteriorly, particularly at posterior angle; anterior angle rounded. Length of malar space less than basal mandibular width. Clypeal apex relatively straight (in Megaloptina and Megommation the clypeal apex is deeply concave, a putative synapomorphy uniting these groups and tentatively suggesting that Cleptommation may not be derived from within the former taxa). Epistomal sulcus orthogonal. Ventral apex of facial medial line forming small tubercle in basal part of supraclypeal area. Ocelli not greatly enlarged; ocellar furrow absent. Vertex not expanded or ridged posterior to ocelli. Preoccipital ridge rounded. Pronotal dorsal ridge rounded; lateral angle not produced, obtuse; lateral ridge rounded. Mesoscutal anterior border rounded; mesoscutal lip rounded. Tegula oval (not notched). Probasitarsal brush absent; metatibial scopa absent (Figs. 2, 43); metabasitibial plate absent; inner metatibial spur serrate. Forewing marginal cell with acute apex; hind wing distal hamuli with irregular pattern of spacing. ♂: Labrum without distal process; basal area not notched. Mandible simple. Ventral apex of facial medial line forming small tubercle in basal part of supraclypeal area. Antenna extending back to mesoscutellum or metanotum; first flagellomere as long as second flagellomere. Inner metatibial spur serrate. Dense patch of plumose setae surrounding propodeal spiracle. Metasoma slightly elongate; apical margins of metasomal sterna II – III with slight median projection, that of sternum III more prominent than sternum II; apical margins sterna IV – V unmodified; apical margin of sternum VI emarginate; apical margins of sternum VII – VIII with median process; proctiger with anal filaments. Gonobasal bridge narrow; dorsal lobes strong; basal process of gonostylus absent; parapenial lobe present; dorsal process of gonostylus partially membranous; prong present on ventral surface of penis valve. INCLUDED SPECIES: Aside from the type species, C. minutum, three additional species are recognized herein: C. bulivyense n. sp., C. tupaqi n. sp., and C. chibchani n. sp.	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D07FFC92B13FAE73BBFF96E.taxon	distribution	DISTRIBUTION: Presently, individuals of the various species are recorded from Costa Rica and Panama in Central America, from Andean Ecuador and Bolivia, across Amazonian Peru and Brazil, and in French Guiana, Guyana, and Suriname along the Guiana Shield. With further collecting it is assured that specimens, and perhaps even further species, will be found in at least Colombia, Andean Peru, and Venezuela.	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D07FFC92B13FAE73BBFF96E.taxon	biology_ecology	BIOLOGY: As alluded to above, no biological information exists for this small group of cleptoparasitic bees. Presumed hosts are among Megaloptina, which occur in the same areas as species of Cleptommation. It would be revealing to discover not only the hosts, but the mode of parasitism and immature stages. As nothing really is known about the biology of Megaloptina, such an investigation would be fruitful beyond that of the just the cleptoparasites.	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D05FFC32BD4FB14394BFAF7.taxon	description	(Figs. 1 – 12)	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D05FFC32BD4FB14394BFAF7.taxon	diagnosis	DIAGNOSIS: Cleptommation bulivyense is most similar to C. tupaqi, particularly in the structure of the male terminalia and coloration of the mesosoma. The Bolivian species can be distinguished from C. tupaqi by the brown to dark brown metasoma (Figs. 1 – 4, 10, 11) and the male genitalia (Figs. 7 – 9), and from others in the genus by the form of the hidden sterna (Fig. 6) and the coloration of the mesosoma (Figs. 1 – 4, 10, 11).	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D05FFC32BD4FB14394BFAF7.taxon	description	DESCRIPTION: ♂: Total body length 8.44 mm (8.00 – 9.13 mm); forewing length 7.33 mm (6.13 – 7.33 mm). Head length 1.93 mm (1.80 – 2.00 mm), width 2.27 mm (2.23 – 2.33 mm; upper interorbital distance 1.17 mm (1.17 – 1.20 mm); lower interorbital distance 0.83 mm (0.80 – 0.93 mm). Intertegular distance 1.50 mm (1.47 – 1.50 mm). Forewing basal vein nearly confluent with cu-a, only slightly set distally; 1 rs-m confluent with 1 m-cu; 2 rs-m distad 2 m-cu by 10 times vein width, 2 rs-m relatively straight; first submarginal cell longer than combined lengths of second and third submarginal cells; second submarginal narrow, anterior border of second submarginal cell along Rs less than length of third submarginal cell along same vein; anterior border of third submarginal cell about three-quarters length of posterior border; hind wing with distal hamuli arranged 3 - 1 - 1 - 3. Terminalia as in figures 6 – 9. Labrum imbricate. Clypeus imbricate with faint, coarse punctures over yellow surface, otherwise smooth with smaller, faint punctures separated by a puncture width or less over basal brown integument; supraclypeal area smooth with small punctures separated by a puncture width or less laterally; remainder of face with small punctures separated by a puncture width or less, progressively becoming more dense on upper face with punctures separated by less than a puncture width; punctures become minute and more widely-spaced by ocellar area, separated by 1 – 2 times a puncture width in ocellocular area and 2 – 4 times medially on vertex; gena with punctures like those on vertex; postgena impunctate. Pronotum imbricate with scattered, minute punctures. Mesoscutum smooth with minute punctures separated by 1 – 2.5 times a puncture width, punctures fainter and sparser along anteromedial border around medial line and integument faintly imbricate; mesoscutellum as on disc of mesoscutum except punctures separated by 1 – 3 times a puncture width; metanotum smooth, weakly nodulose near setal bases and with punctures separated by 1.5 – 3 times a puncture width. Preëpisternal area imbricate; mesepisternum with small punctures separated by a puncture width or less, punctures becoming fainter and sparser ventrally and posteriorly; hypoepimeral area impunctate; metepisternum faintly imbricate and impunctate; lateral and posterior surfaces of propodeum weakly imbricate and impunctate; basal dorsal-facing area of propodeum finely imbricate and impunctate. First metasomal tergum smooth, shining, and impunctate; remaining terga finely imbricate; sterna finely imbricate. Mandible yellow except reddish at apex; labrum yellow; clypeus yellow except basal margins brown with strong metallic brassy green highlights; supraclypeal area entirely brown with strong metallic brassy green highlights. Scape yellow but frequently with slightly brownish patch at upper apical margin; pedicel and flagellum brown. Face, vertex, and gena brown with strong metallic brassy green highlights; postgena brown with faint metallic highlights. Labiomaxillary complex yellow. Pronotum and propleura yellow, sometimes with brownish patches with metallic green highlights on dorsal-facing part of pronotum bordering mesoscutum; remainder of mesosoma brown with strong metallic brassy green highlights, highlights sometimes weaker on disc of mesoscutum, mesoscutellum, metanotum, and dorsal-facing surface of propodeum to varying degrees; preëpisternal area frequently with yellowish area and lower posterior corner of mesepisternum often with patch of yellow; lateral surface of propodeum with large surface of yellow; tegula brown without metallic highlights. Wing membranes hyaline and slightly tinged like parchment; veins amber colored except Sc + R slightly darker. Legs entirely yellow. Metasoma entirely brown (Figs. 1, 3, 4), sometimes with patch of yellow in anterior portion of first tergum and corresponding sternum. Pubescence generally yellow; face with particularly prominent short, subappressed, plumose setae which become more sparse toward ocellar area (Fig. 5); vertex with scattered erect, long setae; gena with plumose setae as those on face although not as dense; postgena with sparse, simple, long, erect setae. Mesoscutum with scattered, fine, simple or little-branched, subappressed, yellow setae; with short plumose setae as those on face along lateral borders; mesoscutellum similar to mesoscutum, posteriorly with more elongate and more erect setae; metanotum similar to mesoscutellum although erect setae much more numerous and over entire surface; pleura with scattered yellow setae, becoming longer ventrally; propodeum with setae as on pleura except with dense patches of moderately long, plumose setae around propodeal spiracle. First metasomal tergum with sparse setae over much of surface except fine, suberect, yellow setae becoming more numerous, albeit still widely scattered, in posterior half; remainder of metasomal terga with fine, short, suberect to subappressed setae scattered over surface, becoming progressively more numerous and slightly longer on more apical terga; sterna with scattered, fine, suberect yellow setae, those of apicolateral corners of sterna II – IV typically longer and somewhat more sinuate. ♀: As described for the male except as follows: Total body length 8.53 – 9.13 mm; forewing length 7.27 – 7.53 mm. Head length 2.20 mm, width 2.67 – 2.80 mm; upper interorbital distance 1.37 – 1.53 mm; lower interorbital distance 1.30 – 1.43 mm. Intertegular distance 1.57 – 1.77 mm. Punctation generally as described for male, although mesepisternal punctures smaller, fainter, and more sparse than those of male, thereby giving mesepisternum a more smooth and shining appearance. Pedicel and first flagellomere yellowish brown; remainder of flagellum brown. Metasoma entirely brown (Figs. 10, 11) except basal area of first tergum lighter and areas of yellow basally on first and second sterna. Protibia with minute patch of slightly fuscous setae on outer apex; mesotibia with patch of dark brown to black setae on outer apex (Fig. 10), most setae distinctly curved apically; metatibia with patch of dark brown to black setae on outer apex, patch extending proximally as a thin row of setae to cover about apical one-third of metatibia (Fig. 10), most setae distinctly curved or sinuate apically. Metasomal sterna with scattered, fine, short to long, largely-erect, yellow setae in apical half to third.	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D05FFC32BD4FB14394BFAF7.taxon	materials_examined	HOLOTYPE: ♂, BOLIVIA: Cochabamba: Cochabamba, 67.5 km NE, Est. Biol. Valle del Sajita, Univ. de San Simon, 300 m, 17 º 6 ’ 33 ’’ S, 64 º 47 ’ 52 ’’ W, 9 – 13 Feb 1999, F. Genier, ex: flight intercept trap (SEMC). PARATYPES: 2 ♂♂, 1 ♀, BOLIVIA: Cochabamba: Cochabamba, 67.5 km NE, Est. Biol. Valle del Sajita, Univ. de San Simon, 300 m, 17 º 6 ’ 33 ’’ S, 64 º 47 ’ 52 ’’ W, 9 – 13 Feb 1999, F. Genier, ex: flight intercept trap (SEMC); 1 ♂, Cochabamba: Cochabamba, 67.5 km NE, Est. Biol. Valle del Sajita, Univ. de San Simon, 300 m, 17 º 6 ’ 33 ’’ S, 64 º 47 ’ 52 ’’ W, 9 – 13 Feb 1999, R. Hanley, ex: flight intercept trap (SEMC); 1 ♀, Cochabamba, Villa Tunari, Hotel El Puenta, 357 m, 15 – 27 - xii- 2005 [15 – 27 December 2005], S. & J. Peck (SEMC).	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D05FFC32BD4FB14394BFAF7.taxon	etymology	ETYMOLOGY: The specific epithet is based on the Quechua spelling of Bolivia (Bulivya Mamallaqta = Estado Plurinacional de Bolivia).	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D0EFFC02BCCFA643984FAD7.taxon	description	(Figs. 13 – 19)	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D0EFFC02BCCFA643984FAD7.taxon	diagnosis	DIAGNOSIS: The new species is most similar to C. bulivyense in overall coloration of the head and mesosoma and form of the hidden sterna (i. e., the more broad and blunt apical process of sternum VII: cf. figures 6 and 16) (vide supra). It differs from that species in the metasomal terga largely yellowish with brown apical margins (Figs. 13, 15) and male genitalia (Figs. 17 – 19).	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D0EFFC02BCCFA643984FAD7.taxon	description	DESCRIPTION: ♂: Total body length 8.67 mm; forewing length 6.40 mm. Head length 1.70 mm, width 2.07 mm; upper interorbital distance 1.10 mm; lower interorbital distance 0.83 mm. Intertegular distance 1.27 mm. Forewing basal vein distad cu-a by about one vein width; 1 rs-m confluent with 1 m-cu; 2 rs-m distad 2 m-cu by 6 times vein width, 2 rs-m relatively straight; first submarginal cell longer than combined lengths of second and third submarginal cells; second submarginal narrow, anterior border of second submarginal cell along Rs much less than length of third submarginal cell along same vein; anterior border of third submarginal cell less than three-quarters length of posterior border; hind wing with distal hamuli arranged 4 - 1 - 2. Terminalia as in figures 16 – 19. Labrum imbricate. Clypeus imbricate with faint, coarse punctures over yellow surface, otherwise smooth with smaller, faint punctures separated by a puncture width or less over basal brown integument; supraclypeal area smooth with small punctures separated by a puncture width or less laterally; remainder of face with small punctures separated by a puncture width or less, progressively becoming more dense on upper face with punctures separated by less than a puncture width; punctures then become minute and more widely-spaced by ocellar area, separated by 1 – 2.5 times a puncture width in ocellocular area and 2 – 5 times medially on vertex; gena with punctures like those on vertex; postgena impunctate. Pronotum imbricate with scattered, minute punctures. Mesoscutum smooth with minute punctures separated by 1 – 2.5 times a puncture width, punctures fainter and sparser along anteromedial border around medial line and integument in that area faintly imbricate; mesoscutellum as on disc of mesoscutum except punctures separated by 1 – 3 times a puncture width; metanotum smooth, weakly nodulose near setal bases and with punctures separated by 1 – 3 times a puncture width. Preëpisternal area imbricate; mesepisternum with small punctures separated by 0.5 – 2 times a puncture width, punctures becoming fainter and sparser ventrally and posteriorly; hypoepimeral area impunctate; metepisternum faintly imbricate and impunctate; lateral and posterior surfaces of propodeum weakly imbricate and impunctate; basal dorsal-facing area of propodeum finely imbricate and impunctate. First metasomal tergum smooth, shining, and impunctate; terga II and III finely and faintly imbricate, remaining terga finely imbricate but more noticeably so than on terga II and III; sterna finely imbricate. Mandible yellow except reddish at apex; labrum yellow; clypeus yellow except basal margins brown with strong metallic brassy green highlights (Fig. 14); supraclypeal area entirely brown with strong metallic brassy green highlights. Scape entirely yellow (Figs. 13 – 15); pedicel and first flagellomere yellowish brown, remainder of flagellum brown. Face, vertex, and gena brown with strong metallic brassy green highlights; postgena brown with faint metallic highlights. Labiomaxillary complex yellow. Pronotum and propleura yellow; remainder of mesosoma brown with strong metallic brassy green highlights, highlights weaker on disc of mesoscutum and dorsal-facing surface of propodeum; lower posterior corner of mesepisternum with a patch of yellow; lowermost portion of metepisternum yellow; lateral surface of propodeum yellow; tegula translucent yellow without metallic highlights. Wing membranes hyaline and clear; veins yellowish brown except Sc + R slightly darker and pterostigma distinctly yellow. Legs entirely yellow. Metasoma light yellow brown, with apical margins of terga brown. Pubescence generally yellow; face with particularly prominent short, subappressed, plumose setae which become more sparse toward ocellar area; vertex with scattered erect, long setae; gena with plumose setae as those on face although not as dense; postgena with sparse, simple, long, erect setae. Mesoscutum with scattered, fine, simple or little-branched, subappressed, yellow setae; with short plumose setae as those on face along lateral borders; mesoscutellum similar to mesoscutum, posteriorly with more elongate and more erect setae; metanotum similar to mesoscutellum although erect setae much more numerous and over entire surface; pleura with scattered yellow setae, becoming longer ventrally; propodeum with setae as on pleura except with dense patches of moderately long, plumose setae around propodeal spiracle. First metasomal tergum with sparse setae over much of surface except fine, suberect, yellow setae becoming more numerous, albeit still widely scattered, in posterior half; remainder of metasomal terga with fine, short, suberect to subappressed setae scattered over surface, becoming progressively more numerous and slightly longer on more apical terga; sterna with scattered, fine, suberect yellow setae, those of apicolateral corners of sterna II – V typically longer. ♀: Unknown.	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D0EFFC02BCCFA643984FAD7.taxon	materials_examined	HOLOTYPE: ♂, ECUADOR: Pich. [Pichincha], Tinalandia, Santa Domingo, 14 June 1976, S. & J. Peck (SEMC).	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D0EFFC02BCCFA643984FAD7.taxon	etymology	ETYMOLOGY: The specific epithet recognizes Incan leader Túpac Inca Yupanqui (ca. 1453? – 1493) (Quechua: Tupaq Inka Yupanki), tenth Sapa Inca, who expanded the empire to encompass regions as far North as much of modern Ecuador.	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D0DFFDA2B3CFA443BA8F9B7.taxon	description	(Figs. 20 – 35)	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D0DFFDA2B3CFA443BA8F9B7.taxon	diagnosis	DIAGNOSIS: Cleptommation minutum is most similar to C. chibchani, described below (vide infra), both having a similar form of the hidden sterna (cf. figures 23 and 39) and both have similar head and mesosomal coloration [e. g., the scape and pedicel yellow (Figs. 22, 38, 45), rather than the pedicel much darker than the scape (Figs. 5, 12)]. The former species can be separated readily from the latter by the almost entirely brown metasoma (e. g., Figs. 20, 27, 30, 31, 33), the lack of a strong metallic green coloration to the mesepisternum (e. g., Figs. 20, 27, 30, 33; although this sometimes occurs in females of C. chibchani), the more fuscous setae of the metasomal terga, and the male genitalia (Figs. 24 – 26).	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D0DFFDA2B3CFA443BA8F9B7.taxon	description	DESCRIPTION: ♂: Total body length 8.47 mm (8.07 – 8.67 mm); forewing length 6.53 mm (6.20 – 6.53 mm). Head length 1.97 mm (1.83 – 1.97 mm), width 2.37 mm (2.27 – 2.37 mm); upper interorbital distance 1.33 mm (1.17 – 1.33 mm); lower interorbital distance 0.97 mm (0.83 – 0.97 mm). Intertegular distance 1.37 mm (1.37 – 1.43 mm). Forewing basal vein confluent with cu-a to distad cu-a by vein width; 1 rs-m confluent with 1 m-cu; 2 rs-m distad 2 m-cu by 6 – 8 times vein width, 2 rs-m relatively straight; first submarginal cell longer than combined lengths of second and third submarginal cells; second submarginal narrow, anterior border of second submarginal cell along Rs less than length of third submarginal cell along same vein; anterior border of third submarginal cell less than three-quarters length of posterior border; hind wing with distal hamuli arranged 3 - 1 - 1 - 2 or 3 - 1 - 2. Terminalia as in figures 23 – 26. Labrum imbricate. Clypeus imbricate with faint, coarse punctures; supraclypeal area smooth with small punctures separated by a puncture width or less laterally; remainder of face with small punctures separated by a puncture width or less, progressively becoming more dense on upper face with punctures virtually contiguous; punctures become minute and more widely-spaced by ocellar area, separated by 1.5 – 2 times a puncture width in ocellocular area and 2 – 5 times medially on vertex; gena with punctures like those on vertex; postgena impunctate. Pronotum imbricate with scattered, minute punctures. Mesoscutum smooth with minute punctures separated by 1 – 2 times a puncture width, punctures fainter and sparser medially, particularly on anteromedial border around medial line and integument there faintly imbricate; mesoscutellum similar to disc of mesoscutum except punctures separated by 1 – 3 times a puncture width; metanotum smooth, weakly nodulose near setal bases and with punctures separated by 1 – 3 times a puncture width. Preëpisternal area imbricate with scattered punctures; mesepisternum with small punctures separated by 0.5 – 2 times a puncture width, punctures becoming fainter and sparser ventrally and posteriorly; hypoepimeral area impunctate; metepisternum faintly imbricate and impunctate; lateral and posterior surfaces of propodeum weakly imbricate and impunctate; basal dorsal-facing area of propodeum finely imbricate and impunctate. First metasomal tergum smooth, shining, and impunctate; remaining terga finely imbricate; sterna finely imbricate. Mandible yellow except reddish at apex; labrum yellow; clypeus entirely yellow (Fig. 22); supraclypeal area entirely brown with strong metallic brassy green highlights. Scape and pedicel yellow; flagellum brown. Face, vertex, and gena brown with strong metallic brassy green highlights; postgena light brown to yellow, sometimes with faint metallic highlights. Labiomaxillary complex yellow. Pronotum and propleura yellow; mesoscutum brown with metallic brassy green highlights (sometimes highlights are quite weak); remainder of mesosoma yellow except sometimes metanotum as on mesoscutum; tegula translucent yellow without metallic highlights. Wing membranes hyaline and clear to slightly tinged; veins light brown to amber colored except Sc + R slightly darker. Legs entirely yellow. Metasoma entirely brown except first tergum basally yellow with brown apically (Fig. 21) to entirely yellow or yellowish brown (yellow areas more reddish brown in much older, faded specimens), basal quarter to third of second tergum yellow, and first and second sterna yellow to yellowish brown. Pubescence generally yellow; face with particularly prominent short, subappressed, plumose setae and becoming more sparse toward ocellar area; vertex with scattered erect, long setae; gena with plumose setae as those on face although not as dense; postgena with sparse, simple, long, erect setae. Mesoscutum with scattered, fine, simple or little-branched, subappressed, yellow setae; with short plumose setae as those on face along lateral borders; mesoscutellum similar to mesoscutum, posteriorly with more elongate and more erect setae; metanotum similar to mesoscutellum although erect setae much more numerous and over entire surface; pleura with scattered yellow setae, becoming longer ventrally; propodeum with setae as on pleura except with dense patches of moderately long, plumose setae around propodeal spiracle. First metasomal tergum with sparse setae over much of surface except fine, suberect, yellow setae becoming more numerous, albeit still widely scattered, in posterior half; remainder of metasomal terga with fuscous (rather than yellow), fine, short, suberect to subappressed setae scattered over surface, becoming progressively more numerous and slightly longer on more apical terga; sterna with scattered, fine, suberect yellow setae, those of apicolateral corners of sterna II – IV typically longer and somewhat more sinuate. ♀: As described for the male except as follows: Total body length 8.40 – 9.67 mm; forewing length 6.67 – 7.27 mm. Head length 1.87 – 2.13 mm, width 2.43 – 2.67 mm; upper interorbital distance 1.27 – 1.37 mm; lower interorbital distance 1.20 – 1.30 mm. Intertegular distance 1.40 – 1.63 mm. Punctation generally as described for male, although mesepisternal punctures fainter and more sparse. Clypeus sometimes with small areas of brown near basal margin (e. g., Fig. 35), otherwise entirely yellow. Metasoma as in male but sternum II with apical third brown and apical margins of terga II – V sometimes more dark brown (Figs. 30, 31). Protibia with minute patch of slightly fuscous setae on outer apex; mesotibia with patch of dark brown to black setae on outer apex, setae curved apically or sinuate; metatibia with patch of dark brown to black setae on outer apex, patch extending proximally as a thin row of setae over apical one-third of metatibia, most setae distinctly curved or sinuate apically. LECTOTYPE (here designated): ♂, (Figs. 27 – 29), BRAZIL: Estado do Amazonas, Tefé, 25.9.1904 [25 September 1904], Ducke (ZMHB). ADDITIONAL MATERIAL: Amazonian Basin: 1 ♂, PERU: Tambopata Prov., 15 km NE Pto. Maldonaldo, 200 m, 13 June 1989, R. A. Leschen, # 34 ex: malaise trap (SEMC); 1 ♀, Tambopata Prov., 15 km NE Pto. Maldonaldo, 200 m, 28 June 1989, J. Ashe, R. Leschen, # 308 ex: flight intercept trap (SEMC); 1 ♂, PERU: Tambopata Reserve, 270 m, 22 – 30 May 1995, S. Cameron and J. Whitfield (UADE). Guiana Shield: 1 ♀, SURINAME: Para: Carolina Creek, 11 km SE Zanderij Airport, 30 m, 5 º 23 ’ 36 ’’ N, 55 º 9 ’ 29 ’’ W, 19 – 20 Jun [June] 1999, Z. H. Falin, A. Gangadin, ex: flight intercept trap (SEMC); 1 ♂, FRENCH GUIANA: Wanaboo (near Nason), Marowijne River, 40 m, 4 º 43 ’ 35 ’’ N, 54 º 26 ’ 36 ’’ W, 31 May- 5 Jun [June] 1999, Z. H. Falin, A. B. DeDijn, ex: flight intercept trap (SEMC); 1 ♂, Saül, 7 km N, Les Eaux Claires, 220 m, 3 º 39 ’ 46 ’’ N, 53 º 13 ’ 19 ’’ W, 31 May- 3 Jun [June] 1997, J. Ashe, R. Brooks, ex: flight intercept trap (SEMC); 1 ♂, GUYANA: Region 8, Iwokrama Forest, 1 km W Kurupukari, Iwokrama Field Stn., 60 m, 4 º 40 ’ 19 ’’ N, 58 º 41 ’ 4 ’’ W, 20 – 25 May 2001, R. Brooks, Z. Falin, ex: flight intercept trap (SEMC).	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D0DFFDA2B3CFA443BA8F9B7.taxon	discussion	COMMENTS: For now I have kept together those specimens coming from the Amazon Basin and Guiana Shield as they are remarkably similar, including the male genitalia. Nonetheless, there is noticeable color variation and it is possible that the individuals from the Amazon Basin (here represented by material from northern Brazil, including the holotype, and eastern Amazonian Peru) are specifically distinct and would constitute the ‘ true’ C. minutum. Should the accumulation of larger series and data from other sources (e. g., genetic data, hosts) suggest those in the Guiana Shield are truly different, then they would constitute a fifth and undescribed species for the genus. I have segregated the data from the small series of specimens into their respective geographic types (vide supra). I hope that the conservative position I’ve taken for the moment will be useful for future workers and stimulate some to more aggressively seeking these bees in the wild and to the discovery of their biology. Previously I and colleagues considered the male we were sent from Berlin as the holotype (Engel et al., 1997). This was clearly an error as Friese (1926) mentions more than one specimen. Accordingly, I hereby designate a lectotype in order to stabilize the application of the name.	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D17FFDD2BD9F9A43E7DFCF7.taxon	description	(Figs. 36 – 45)	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D17FFDD2BD9F9A43E7DFCF7.taxon	diagnosis	DIAGNOSIS: Refer to diagnosis for C. minutum (vide supra).	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D17FFDD2BD9F9A43E7DFCF7.taxon	description	DESCRIPTION: ♂: Total body length 8.13 mm (8.00 – 10.00 mm); forewing length 6.33 mm (5.73 – 7.00 mm). Head length 1.93 mm (1.67 – 2.03 mm), width 2.07 mm (2.00 – 2.43 mm); upper interorbital distance 1.03 mm (1.00 – 1.07 mm); lower interorbital distance 0.97 mm (0.83 – 1.00 mm). Intertegular distance 1.47 mm (1.40 – 1.70 mm). Forewing basal vein distad cu-a by vein width; 1 rs-m confluent with 1 m-cu; 2 rs-m distad 2 m-cu by 10 times vein width, 2 rs-m relatively straight; first submarginal cell longer than combined lengths of second and third submarginal cells; second submarginal narrow, anterior border of second submarginal cell along Rs less than length of third submarginal cell along same vein; anterior border of third submarginal cell about one-half to three-quarters length of posterior border; hind wing with distal hamuli arranged 3 - 1 - 3. Terminalia as in figures 39 – 42. Labrum imbricate. Clypeus imbricate with faint, coarse punctures; supraclypeal area smooth with small punctures separated by a puncture width or less laterally; remainder of face with small punctures separated by a puncture width or less, progressively becoming more dense on upper face; punctures minute and more widely-spaced by ocellar area, separated by 1 – 2 times a puncture width in ocellocular area and 2 – 4 times medially on vertex; gena with punctures like those on vertex; postgena impunctate. Pronotum imbricate with scattered, minute punctures. Mesoscutum smooth with minute punctures separated by 1 – 2 times a puncture width, punctures fainter and sparser on anteromedial border around medial line and integument there faintly imbricate; mesoscutellum as on disc of mesoscutum except punctures separated by 1 – 3 times a puncture width; metanotum smooth, weakly nodulose near setal bases and with punctures separated by 1 – 2.5 times a puncture width. Preëpisternal area imbricate with scattered punctures; mesepisternum with small punctures separated by 0.5 – 2 times a puncture width, punctures becoming fainter and sparser ventrally and posteriorly; hypoepimeral area impunctate; metepisternum faintly imbricate and impunctate; lateral and posterior surfaces of propodeum weakly imbricate and impunctate; basal dorsal-facing area of propodeum finely imbricate and impunctate. First metasomal tergum smooth, shining, and impunctate; remaining terga finely imbricate; sterna finely imbricate. Mandible yellow except reddish at apex; labrum yellow; clypeus entirely yellow, sometimes with basal margins brown with metallic green highlights; supraclypeal area entirely brown with strong metallic brassy green highlights. Scape and pedicel yellow (Fig. 38); flagellum brown. Face, vertex, and gena brown with strong metallic brassy green highlights; postgena light brown to yellow, sometimes with faint metallic highlights. Labiomaxillary complex yellow. Pronotum and propleura yellow; mesoscutum brown with strong metallic brassy green highlights, sometimes with highlights weaker on disc; mesoscutellum yellow, sometimes with weak metallic green highlights; metanotum often similar to mesoscutellum, sometimes slightly darker; propodeum yellow, basal area often with weak metallic green highlights; preëpisternum light brown with metallic green highlights above, yellow below; mesepisternum brown with strong metallic brassy green highlights except yellow in lower posterior section; upper part of metepisternum often brown with metallic green highlights, remainder yellow; tegula translucent yellow without metallic highlights. Wing membranes hyaline and clear to slightly tinged as parchment; veins amber colored except Sc + R slightly darker. Legs entirely yellow. Metasoma entirely yellow except apical margins of terga brown (Figs. 36, 37). Pubescence generally yellow; face with particularly prominent short, subappressed, plumose setae and becoming more sparse toward ocellar area; vertex with scattered erect, long setae; gena with plumose setae as those on face although not as dense; postgena with sparse, simple, long, erect setae. Mesoscutum with scattered, fine, simple or little-branched, subappressed, yellow setae; with short plumose setae as those on face along lateral borders; mesoscutellum as on mesoscutum, posteriorly with more elongate and more erect setae; metanotum as on mesoscutellum although erect setae much more numerous and over entire surface; pleura with scattered yellow setae, becoming longer ventrally; propodeum with setae as on pleura except with dense patches of moderately long, plumose setae around propodeal spiracle. First metasomal tergum with sparse setae over much of surface except fine, suberect, yellow setae becoming more numerous, albeit still widely scattered, in posterior half; remainder of metasomal terga with fine, short, suberect to subappressed setae scattered over surface, becoming progressively more numerous and slightly longer on more apical terga; sterna with scattered, fine, suberect yellow setae, those of apicolateral corners of sterna II – IV typically longer and somewhat more sinuate. ♀: As described for the male except as follows: Total body length 8.20 – 10.13 mm; forewing length 6.67 – 7.33 mm. Head length 2.07 – 2.27 mm, width 2.50 – 2.77 mm; upper interorbital distance 1.40 – 1.50 mm; lower interorbital distance 1.37 – 1.50 mm. Intertegular distance 1.50 – 1.77 mm. Punctation generally as described for male, although mesepisternal punctures distinctly fainter and more sparse. Mespisternum sometimes entirely yellow with metallic green highlights. Mesotibia with patch of dark brown to black setae on outer apex, most setae distinctly curved apically; metatibia with patch of dark brown to black setae on outer apex (Fig. 43), patch extending proximally as a thin row of setae to cover about apical one-quarter of metatibia, most setae distinctly curved or sinuate apically. Metasomal sterna with scattered, fine, short to long, largely-erect, yellow setae in apical half to third.	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D17FFDD2BD9F9A43E7DFCF7.taxon	materials_examined	HOLOTYPE: ♂, PANAMA: Colón, 6.1 km on Pipeline Rd., nr. Gamboa, 40 m, 9 º 6 ’ N, 79 º 45 ’ W, 27 – 29. V. 1995 [27 – 29 May 1995], J. Ashe, ex: flight intercept trap (SEMC). PARATYPES: 1 ♀, PANAMA: Canal Zone, Barro Colorado Is., 27. iv. 63 [27 April 1963], C. W. & M. E. Rettenmeyer / taken in Malaise trap (SEMC); 2 ♀♀, PANAMA: Panamá, Barro Colorado Island, 9 º 11 ’ N, 79 º 51 ’ W, 3 July 1994, D. Banks, ex: flight intercept trap (SEMC); 1 ♀, PANAMA: Panamá, Barro Colorado Island, 9 º 11 ’ N, 79 º 51 ’ W, 8 July 1994, D. Banks, ex: flight intercept trap (SEMC); 2 ♀♀, PANAMA: Panamá, Barro Colorado Island, 9 º 11 ’ N, 79 º 51 ’ W, 22 July 1994, D. Banks, ex: flight intercept trap (SEMC); 1 ♂, PANAMA: Panamá, Barro Colorado Island, 9 º 11 ’ N, 79 º 51 ’ W, 1 Aug 1994, D. Banks, ex: flight intercept trap (SEMC); 1 ♂, 1 ♀, PANAMA: Panamá, Barro Colorado Island, 9 º 11 ’ N, 79 º 51 ’ W, 6 Aug 1994, D. Banks, ex: flight intercept trap (SEMC); 1 ♂, PANAMA: Coclé, 7.2 km NE El Copé, 730 m, 8 º 37 ’ N, 80 º 35 ’ W, 20. v – 7. vi. 1995, J. Ashe, R. Brooks, flt. intercept trap (SEMC); 1 ♂, PANAMA: Darién: Cana Biological Station, Serrania de Pirre, 1450 m, 7 º 45 ’ 18 ’’ N, 77 º 41 ’ 6 ’’ W, 7 – 9 Jun 1996, J. Ashe, R. Brooks, ex: flight intercept trap (SEMC); 1 ♂, PANAMA: Darién: Cana Biological Station, 550 m, 7 º 45 ’ 18 ’’ N, 77 º 41 ’ 6 ’’ W, 7 – 9 Jun 1996, J. Ashe, R. Brooks, ex: flight intercept trap (SEMC); 1 ♂, PANAMA: Darién: Cana Biological Station, 530 m, 7 º 45 ’ 18 ’’ N, 77 º 41 ’ 6 ’’ W, 3 – 7 Jun 1996, J. Ashe, R. Brooks, ex: flight intercept trap (SEMC); 2 ♂♂, PANAMA: Colon: 15 km N. jct. Escobal & Pina Rds., ca. 30 m, 2 Jun 1996, J. Ashe, R. Brooks, ex: flight intercept trap (SEMC); 1 ♀, PANAMA: Panama: Barro Colorado Island, 40 m, 9 º 11 ’ 0 ’’ N, 79 º 51 ’ 0 ’’ W, 17 – 23 Jun 2000, S. Chatzimanolis, ex: flight intercept trap (SEMC); 1 ♀, PANAMA: Panama: Barro Colorado Island, 40 m, 9 º 11 ’ 0 ’’ N, 79 º 51 ’ 0 ’’ W, 18 – 22 Jun 2000, S. Chatzimanolis, ex: flight intercept trap (SEMC); 2 ♀♀, PANAMA: Panama: Barro Colorado Island, 40 m, 9 º 11 ’ 0 ’’ N, 79 º 51 ’ 0 ’’ W, 22 – 25 Jun 2000, S. Chatzimanolis, ex: flight intercept trap (SEMC); 1 ♂, PANAMA: Panama: Barro Colorado Island, 40 m, 9 º 11 ’ 0 ’’ N, 79 º 51 ’ 0 ’’ W, 23 – 27 Jun 2000, S. Chatzimanolis, ex: flight intercept trap (SEMC); 1 ♂, PANAMA: Panama: Barro Colorado Island, 40 m, 9 º 11 ’ 0 ’’ N, 79 º 51 ’ 0 ’’ W, 25 – 30 Jun 2000, S. Chatzimanolis, ex: flight intercept trap (SEMC); 1 ♀, PANAMA: Panama: Barro Colorado Island, 40 m, 9 º 11 ’ 0 ’’ N, 79 º 51 ’ 0 ’’ W, 31 Jul – 4 Aug 2000, S. Chatzimanolis, ex: flight intercept trap (SEMC); 1 ♂, COSTA RICA: Alajuela: Bijagua, 20 km S. Upala, 18 – 26 March 1991, F. D. Parker (EMUS); 1 ♀, COSTA RICA: Alajuela: Bijagua, 20 km S. Upala, 16 February 1991, F. D. Parker (EMUS); 1 ♀, COSTA RICA: Heredia: Chilamate, 75 m, December 1989 – March 1990, Hanson and Godoy (EMUS); 1 ♂, COSTA RICA: Limón: 16 km W. Guápiles, 400 m, April 1989, P. Hanson (EMUS); 1 ♀, COSTA RICA: Puntarenas: Golfo Dulce, 24 km W. Piedras Blancas, 200 m, January 1992, Hanson (EMUS); 1 ♀, COSTA RICA: Puntarenas: Golfo Dulce, 24 km W. Piedras Blancas, 200 m, March 1992, Hanson (EMUS); 1 ♀, COSTA RICA: Puntarenas: Golfo Dulce, 24 km W. Piedras Blancas, 200 m, April-May 1992, Hanson (EMUS); 1 ♀, COSTA RICA: Puntarenas: Golfo Dulce, 24 km W. Piedras Blancas, 200 m, February-March 1993, Hanson (EMUS).	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D17FFDD2BD9F9A43E7DFCF7.taxon	etymology	ETYMOLOGY: The specific epithet honors the Chibchan-speaking peoples (the Chibchan language family, including Waimí, Borũca, Talamanca, Pech, Kuna, Chibcha, Arwako, and many others) who inhabited much of pre-Columbian Panama and Costa Rica (as well as southern Nicaragua and portions of Honduras and Colombia).	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
1269837C3D10FFDC2B8DFC673ED1FE77.taxon	discussion	Known distributions are summarized in the couplets but are not meant as diagnostic for identification. Females are unknown for C. tupaqi.	en	Engel, Michael S. (2013): Revision of the cleptoparasitic bee genus Cleptommation (Hymenoptera: Halictidae). Journal of Melittology 2013 (22): 1-26, DOI: 10.17161/jom.v0i22.4608, URL: https://doi.org/10.17161/jom.v0i22.4608
