identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
34466794E3B35AA996FACD46E195FF5F.text	34466794E3B35AA996FACD46E195FF5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes akitai Wewalka 1997	<div><p>Microdytes akitai Wewalka, 1997</p><p>Fig. 18</p><p>Microdytes akitai Wewalka, 1997: 17; Wewalka 2011: 37; Nilsson and Hájek 2023: 211.</p><p>Type locality.</p><p>Laos, Vientiane Province, Mt. Phou Khao Khouay.</p><p>Material examined.</p><p>Laos: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.160835&amp;materialsCitation.latitude=18.386168" title="Search Plazi for locations around (long 104.160835/lat 18.386168)">Bolikhamsay Province</a>. 22 exs., Nam Khading National Bio-Diversity Conservation Area, Tad Paloy campsite, 18°23.17'N, 104°09.65'E, alt. 300 m, forest stream, 8.-11.VII.2010 &amp; 24.-28.V.2011, NHMB Basel Laos 2010 &amp; 2011 Expeditions, M. Brancucci, M. Geiser, D. Hauk, A. Phantala &amp; E. Vongphachan leg. (CGW, CRO, NMB, NMP) (Fig. 18) .</p><p>Comments.</p><p>Specimens from Bolikhamsay have more distinct elytral markings and darker head color than in the specimens from the type locality but no other differences have been observed, in particular, in the male genitalia.</p><p>Distribution.</p><p>Laos: Vientiane and Bolikhamsay (first record) provinces.</p></div>	https://treatment.plazi.org/id/34466794E3B35AA996FACD46E195FF5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
1ADE86AE29DD50F28720770561AC9E3B.text	1ADE86AE29DD50F28720770561AC9E3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes balkei Wewalka 1997	<div><p>Microdytes balkei Wewalka, 1997</p><p>Figs 17, 19, 39D</p><p>Microdytes balkei Wewalka, 1997: 18; Wewalka 2011: 29; Nilsson and Hájek 2023: 211.</p><p>Type locality.</p><p>Thailand, Rayong Province, Khao Chamao NP.</p><p>Material examined.</p><p>Thailand: Mukdahan Province. 1♂, Phu Pha Thoep N.P., small pools (23), 1.I.2000, Mazzoldi leg. (CGW); Ubon Ratchathani Province. 1♂, 1♀, Nam Yuen District, Dom Pradit St. 240 (alt. 260 m), 22.V.2021, R. Okada leg. (CRO, THNHM) ; Saraburi Province. 1♀, Kaeng Khoi District, Ched Khot St. 122 (alt. 140 m), 30.V.2020, R. Okada leg. (CRO) ; Chonburi Province. 2♂♂, 1♀, Ban Bueng District, Khlong Kiu St. 261 (alt. 200 m), 6.XI.2021, R. Okada leg. (CGW, CRO) (Fig. 17) ; 2♂♂, 1♀, Si Racha District, Bang Phra St. 269 (alt. 90 m), 26.XII.2021, R. Okada leg. (CRO) ; Rayong Province. 4♂♂, Wang Chan District, Pa Yup Nai St 299 (alt. 200 m), 17.VII.2022, R. Okada leg. (CRO, THNHM) .</p><p>Laos: Savannakhet Province. 203 exs., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.63333&amp;materialsCitation.latitude=17.0" title="Search Plazi for locations around (long 105.63333/lat 17.0)">Phou Xang He</a> NBCA, ca. 5 km SW Ban Pa Phaknau, 17°00'N, 105°38'E, alt. 250-400 m, 31.V.-6.VI. 2011, NHMB Basel , Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala &amp; E. Vongphachan leg. (CGW, CRO, NMB, NMP); Sekong Province. 3♀♀, ca. 51 km N Sekong (river) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.66333&amp;materialsCitation.latitude=15.826667" title="Search Plazi for locations around (long 106.66333/lat 15.826667)">Ho Chi Minh</a> trail, 15°49.6'N, 106°39.8'E, alt. 410 m, 14.-15.V.2010, J. Hájek leg. (NMP) .</p><p>Cambodia: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.11667&amp;materialsCitation.latitude=11.566667" title="Search Plazi for locations around (long 103.11667/lat 11.566667)">Koh Hong Province</a>. 12 exs., 20 km SW Koh Hong, Tatai River, 11°34'N, 103°07'E, alt. 50-300 m, 3.-19.V.2005, E. Jendek &amp; O. Šauša leg. (CGW, NMB) .</p><p>Comments.</p><p>Since the figure of the median lobe of this species was not depicted clearly in Wewalka (1997), we present a new figure of the aedeagus including parameres (Fig. 17).</p><p>Distribution.</p><p>Thailand: Mukdahan, Rayong, Trat, Ubon Ratchathani (first record), Saraburi (first record), and Chonburi (first record) provinces, Khao Yai NP [Nakhon Ratchatsima or Nakhon Nayok Province]; Laos (first record): Savannakhet and Sekong provinces; Cambodia (first record): Koh Hong Province.</p></div>	https://treatment.plazi.org/id/1ADE86AE29DD50F28720770561AC9E3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0E028AD633F156FC96DC5E7C696DE758.text	0E028AD633F156FC96DC5E7C696DE758.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes dimorphus Wewalka 1997	<div><p>Microdytes dimorphus Wewalka, 1997</p><p>Fig. 20</p><p>Microdytes dimorphus Wewalka, 1997: 22; Wewalka 2011: 37; Nilsson and Hájek 2023: 211.</p><p>Type locality.</p><p>Thailand, Nakhon Ratchasima or Nakhon Nayok Province, Khao Yai NP.</p><p>Material examined.</p><p>Thailand: Chonburi Province. 1♂, Si Racha District, Bang Phra St. 269 (alt. 90 m), 26.XII.2020, R. Okada leg. (CRO) (Fig. 20) .</p><p>Comments.</p><p>Microdytes dimorphus was described based on a single male specimen from Khao Yai National Park and no additional records are known so far. Therefore, this is the second record of this species. Wewalka (1997) suggested M. dimorphus is more closely related to M. menopausis Wewalka, 1997 (Fig. 27) but differs by having coarser punctures on the elytra similar in size, a punctured abdomen, a more produced clypeus, and larger second segment of antennae. The specimen examined in this study corresponds with M. dimorphus due to the status of punctures on the elytra and abdomen, although the characteristics of clypeus and antenna are intermediate between M. dimorphus and M. menopausis . This species can be distinguished from M. menopausis in having a more slender body shape and indistinct elytral markings at both shoulders and post-median part (Fig. 20).</p><p>Distribution.</p><p>Thailand: Khao Yai NP [Nakhon Ratchasima or Nakhon Nayok Province] and Chonburi (first record) Province.</p></div>	https://treatment.plazi.org/id/0E028AD633F156FC96DC5E7C696DE758	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E3744614B5C25BF8985606CE116951EF.text	E3744614B5C25BF8985606CE116951EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes eliasi Wewalka & Okada 2023	<div><p>Microdytes eliasi Wewalka &amp; Okada sp. nov.</p><p>Figs 2, 7, 12, 42C, 43C</p><p>Type locality.</p><p>Cambodia, Koh Hong Province, 20 km SW Koh Hong, Tatai River, 11°34'N, 103°07'E, alt. 50-300 m.</p><p>Type material.</p><p>Holotype (NMB): ♂ "Cambodia SW, Tatai riv. 20 km SE Koh Hong 11°34'N, 103°07'E, 3.-19.v.2005, 50-300 m, E. Jendek, O. Šauša leg." [printed white label] \ "HOLOTYPUS Microdytes eliasi sp. nov. Wewalka &amp; Okada 2022" [printed red label]. Paratypes: (47 exs.): 7♂♂, 14♀♀, with same data as the holotype (CGW, CRO, NMB, NMP); 2♂♂, Thailand, Saraburi Province, Kaeng Khoi District, Ched Khot St. 122 (alt. 140 m), 14°28'26"N, 101°09'59"E, 30.V.2020, R. Okada leg. (CRO, THNHM); 4♂♂, 4♀♀, Thailand, Sa Kaeo Province, Mueang Sa Kaeo District, Ban Kaeng St. 277 (alt. 80 m), 13°58'47"N, 102°11'29"E, 30.I.2022, R. Okada leg. (CGW, CRO, THNHM); 2♂♂, 3♀♀, Thailand, Chonburi Province, Bo Thong District, That Thong St. 107 (alt. 100 m), 13°15'01"N, 101°22'34"E, 14.III.2020, R. Okada leg. (CGW, CRO, THNHM); 2♂♂, 2♀♀, same locality St. 298 (alt. 100 m), 13.VII.2022, R. Okada leg. (CGW, CRO); 1♀, same province, Si Racha District, Bang Phra St. 269 (alt. 90 m), 13°14'39"N, 101°02'20"E, 26.XII.2021, R. Okada leg. (CRO); 3♂♂, 2♀♀, Thailand, Chanthaburi Province, Khaeng Hong Maeo District, Kaeng Hong waterfall St. 257 (alt. 160 m), 13°02'57"N, 101°45'35"E, 26.VI.2021, R. Okada leg. (CGW, CRO, THNHM); 1♀, same district, Kha riv. St. 258 (alt. 90 m), 12°57'38"N, 101°46'30"E, 26.VI.2021, R. Okada leg. (CRO). All paratypes are provided with printed red paratype labels.</p><p>Diagnosis.</p><p>Microdytes eliasi sp. nov. very closely resembles M. maculatus (Motschulsky, 1860) (Figs 25, 41) in size and coloration but differs from this species by the more regularly oval habitus, the laterally expanded median lobe at apex and constricted tips of paramere (Figs 12, 42C, 43C) (see also comments in M. maculatus). It is also similar to M. feryi Wewalka, 2011 (Fig. 21) in habitus and coloration, but it is smaller and can be distinguished by male genitalia [compared with a male paratype from Myanmar, "Tenasserim, Birma Coll. V. Helfer National Museum Prague" (CGW)].</p><p>Description.</p><p>Measurements. TL = 1.64-1.85 mm (1.76 mm), TL-H = 1.49-1.65 mm (1.55 mm), MW = 1.12-1.25 mm (1.24 mm), TL/MW = 1.33-1.44 (1.42). Body regularly oval, moderately convex (Fig. 2).</p><p>Coloration. Head reddish brown. Pronotum reddish brown, narrowly dark brown along anterior and posterior margins. Elytron dark brown with yellowish brown markings forming a distinct transverse band near base not reaching suture connected along lateral margin with a post-median transverse lateral band, with a small post-median spot near suture, and a triangular spot near apex (Figs 2, 7). Ventral surface of head, prothorax and elytral epipleuron yellowish brown; thorax and abdominal ventrites reddish brown to dark brown. Legs, antennae and palpi yellowish brown.</p><p>Sculpture and structure. Head finely, sparsely, and relatively regularly punctured; anterior half to two-thirds finely microreticulate; clypeus not bordered. Pronotum quite regularly, sparsely, and fairly strongly punctured, with coarser punctures along posterior margin; without microreticulation; lateral margins finely bordered, regularly rounded. Elytron quite regularly, moderately densely and fairly strongly punctured, progressively finer and sparser towards lateral margin; without longitudinal rows of stronger punctures; highly polished and shining; without microreticulation. Ventral surface: metacoxae and metasternum strongly but sparsely punctured, abdomen finely and sparsely punctured; without microreticulation.</p><p>Male. The two parts of the median lobe expanded laterally at apex in ventral aspect (Figs 12A, 42A); slightly curved in lateral aspect (Fig. 12B). The tips of parameres twisted in lateral aspect (Fig. 12C, D); constricted in ventral aspect (Fig. 43C).</p><p>Female. Without secondary sexual characters. Sclerotized spermatheca not found.</p><p>Variation.</p><p>Variation of markings is shown in Fig. 7.</p><p>Etymology.</p><p>This species is dedicated to Elias Bramböck, Vienna, Austria. The species epithet is a name in the genitive singular.</p><p>Habitat.</p><p>In Thailand, this species was collected in small streams at low altitude lower than 200 m (Fig. 45).</p><p>Distribution.</p><p>Thailand: Saraburi, Sa Kaeo, Chonburi, and Chanthaburi provinces; Cambodia: Koh Hong Province.</p></div>	https://treatment.plazi.org/id/E3744614B5C25BF8985606CE116951EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
31E418F324E952669D9F2B64F15A6BC2.text	31E418F324E952669D9F2B64F15A6BC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes franzi Wewalka & Wang 1998	<div><p>Microdytes franzi Wewalka &amp; Wang, 1998</p><p>Fig. 22</p><p>Microdytes franzi Wewalka &amp; Wang, 1998: 65; Wewalka 2011: 38; Nilsson and Hájek 2023: 211.</p><p>Type locality.</p><p>Laos, Vientiane Province, Mt. Phou Khao Khouay.</p><p>Material examined.</p><p>Laos: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.160835&amp;materialsCitation.latitude=18.386168" title="Search Plazi for locations around (long 104.160835/lat 18.386168)">Bolikhamsay Province</a>. 20 exs., Nam Khading National Bio-Diversity Conservation Area, Tad Paloy campsite, 18°23.17'N, 104°09.65'E, alt. 300 m, forest stream, 8.-11.VII.2010 &amp; 24.-28.V.2011, NHMB Basel Laos 2010 &amp; 2011 Expeditions, M. Brancucci, M. Geiser, D. Hauk, A. Phantala &amp; E. Vongphachan leg. (CGW, CRO, NMB, NMP) (Fig. 22) .</p><p>Distribution.</p><p>Laos: Vientiane and Bolikhamsay (first record) provinces.</p></div>	https://treatment.plazi.org/id/31E418F324E952669D9F2B64F15A6BC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
7F29A223BC0C56BEB0FC38B35495A93B.text	7F29A223BC0C56BEB0FC38B35495A93B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes gabrielae Wewalka 1997	<div><p>Microdytes gabrielae Wewalka, 1997</p><p>Fig. 23</p><p>Microdytes gabrielae Wewalka, 1997: 24; Wewalka 2011: 29; Nilsson and Hájek 2023: 211.</p><p>Type locality.</p><p>Thailand, Phetchabun Province, Huai Nam Phang.</p><p>Material examined.</p><p>Thailand: Phetchabun Province. 1♂, Phu Hin Rongkla NP, small stream (7), 25.XII.1999, Mazzoldi leg. (CGW) (Fig. 23); 4♂♂, 3♀♀, Lom Kao District, Ban Noen St. 294 (alt. 1620 m), 18.VI.2022, R. Okada leg. (CGW, CRO, THNHM) ; Phitsanulok Province. 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.00967&amp;materialsCitation.latitude=16.996973" title="Search Plazi for locations around (long 101.00967/lat 16.996973)">Nakhon Thai Distr.</a>, Phu Hin Rong Kla NP, in waterfall, 16°59'49.1"N, 101°00'34.8"E, 7.III.2016, A. Damaška leg. (NMP) .</p><p>Distribution.</p><p>Thailand: Phetchabun and Phitsanulok provinces.</p></div>	https://treatment.plazi.org/id/7F29A223BC0C56BEB0FC38B35495A93B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
5BA38C25DE725916ACB8998D206BB050.text	5BA38C25DE725916ACB8998D206BB050.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes heineri Wewalka 2011	<div><p>Microdytes heineri Wewalka, 2011</p><p>Figs 24, 39A</p><p>Microdytes heineri Wewalka, 2011: 23; Nilsson and Hájek 2023: 211.</p><p>Type locality.</p><p>China, Yunnan Province, Simao Prefecture.</p><p>Material examined.</p><p>Thailand: Chiang Mai Province. 8 exs., Mae Taeng District, Pa Pae St. 49 (alt. 1000 m), 15.VI.2019, R. Okada leg. (CGW, CRO, THNHM) ; 1♂, same district, Kuet Chang St. 170 (alt. 520 m), 16.VIII.2020, R. Okada leg. (CRO) (Fig. 24) .</p><p>Comments.</p><p>Male specimens have distinct setae on the metacoxae (Fig. 39A).</p><p>Distribution.</p><p>Thailand: Nan and Chiang Mai (first record) provinces; Laos: Luang Namtha and Luang Prabang provinces; China: Yunnan Province.</p></div>	https://treatment.plazi.org/id/5BA38C25DE725916ACB8998D206BB050	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0A7211761CF1522295AA7A35CFED94ED.text	0A7211761CF1522295AA7A35CFED94ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes jeenthongi Okada & Wewalka 2023	<div><p>Microdytes jeenthongi Okada &amp; Wewalka sp. nov.</p><p>Figs 3, 8, 13, 39E</p><p>Type locality.</p><p>Thailand, Chiang Mai Province, Mae Chaem District, Tha Pha, 18°30'46"N, 98°25'25"E, alt. 720 m.</p><p>Type material.</p><p>Holotype (THNHM): ♂, "THAI: Chiang Mai Mae Chaem Dist., Tha Pha St. 166 (alt. 720 m) 15.VIII.2020" [printed white label] \ "HOLOTYPE Microdytes jeenthongi sp. nov. Okada &amp; Wewalka 2022 [printed red label]". Paratypes (3 exs.): 2♂♂, same data as the holotype (CGW, NMW); 1♂, same locality, 4.VII.2020, R Okada leg (CRO). All paratypes are provided with printed red paratype labels.</p><p>Diagnosis.</p><p>Microdytes jeenthongi sp. nov. resembles M. shunichii Satô, 1995 (Fig. 36) and M. zetteli Wewalka, 1997 (Fig. 37) in habitus, coloration and very finely punctured metasternum and metacoxae. From M. shunichii it differs by the reddish brown pronotum and very finely and sparsely punctured elytra. From M. zetteli it can be distinguished by larger size, less finely and sparsely punctured head.</p><p>Description.</p><p>Measurements. TL = 1.77-1.82 mm (1.79 mm), TL-H = 1.23-1.26 mm (1.26 mm), MW = 1.18-1.23 mm (1.23 mm), TL/MW = 1.46-1.50 (1.49). Body regularly oval, moderately convex (Fig. 3).</p><p>Coloration. Head reddish brown. Pronotum reddish brown, along the anterior margin narrowly and along the posterior margin more widely dark brown, especially medially. Elytron dark brown with a distinct transverse band at the base not reaching suture, continuing along the margin to a post-median transverse band and a triangular spot near the apex, and a small round post-median spot near suture (Figs 3, 8). Ventral surface of head and prothorax yellowish brown; thorax, abdominal ventrites and elytral epipleuron reddish brown to dark brown. Legs, antennae and palpi yellowish brown.</p><p>Sculpture and structure. Head finely and sparsely, slightly irregularly punctured; anterior forth finely microreticulate; clypeus not bordered. Pronotum punctured moderately irregularly, finely, and sparsely, very coarsely along posterior margin; without microreticulation; lateral margins finely bordered, regularly rounded. Elytron very sparsely and finely punctured, progressively finer and sparser towards lateral margin; one longitudinal row of punctures distinct; highly polished and shining; without microreticulation. Ventral surface: metasternum and metacoxae very finely, sparsely, and irregularly punctured; abdomen almost without punctures; without microreticulation.</p><p>Male. The two parts of the median lobe expanded and pointed at apex forming the unique arrowhead shape in ventral aspect (Fig. 13A); almost straight and slightly expanded downwards at apex in lateral aspect (Fig. 13B). Parameres moderate triangular in lateral aspect (Figs 13C, D).</p><p>Female. Unknown.</p><p>Variation.</p><p>Variation of markings is shown in Fig. 8.</p><p>Etymology.</p><p>This species is dedicated to Tadsanai Jeenthong, collection manager of National Science Museum, Thailand. The species epithet is a name in the genitive singular.</p><p>Habitat.</p><p>This species was collected in a small, shallow stream with gravel bottom flowing through a small valley. In this stream, specimens were collected from a restricted point where tree roots were exposed along the river bottom (Fig. 46).</p><p>Distribution.</p><p>Thailand: Chiang Mai Province.</p></div>	https://treatment.plazi.org/id/0A7211761CF1522295AA7A35CFED94ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E88287CB88D056EA9854CBECE0AAD8BB.text	E88287CB88D056EA9854CBECE0AAD8BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes maculatus (Motschulsky 1860)	<div><p>Microdytes maculatus (Motschulsky, 1860)</p><p>Figs 25, 39C, 41</p><p>Hydrovatus maculatus Motschulsky, 1860: 42; Sharp 1882: 814, 973; Régimbart 1899: 231; Zaitzev 1915: 293; Zimmermann 1920: 34.</p><p>Desmopachria maculatus: Gschwendtner 1936: 367; Balfour-Browne 1946: 106 (uncertain assignment).</p><p>Microdytes maculatus: Vazirani 1969: 301; Vazirani 1977: 24; Satô 1981: 68; Wewalka 1997: 27; Biström et al. 1997: 75; Miller and Wewalka 2010: 35; Wewalka 2011: 29; Nilsson and Hájek 2023: 211.</p><p>Type locality.</p><p>"Ind or", "Dohen [a place with this name exists in Pakistan, Jammu and Kashmir Province, but it is very unlikely that this is the type locality]".</p><p>Material examined.</p><p>Thailand: Chiang Mai Province. 6♂♂, 1♀, Chom Thong District, Ban Luang St. 165 (alt. 360 m), 15.VIII.2020, R. Okada leg. (CGW, CRO) (Fig. 25) ; 9 exs., Mae Chaem District, Tha Pha St. 166 (alt. 720 m), 15.VIII.2020, R. Okada leg. (CGW, CRO) ; 1♂, 1♀, Chiang Dao District, Ping Khong St. 226 (alt. 430 m), 20.III.2021, R. Okada leg. (CRO) ; Chiang Rai Province. 3♀♀, Wiang Kaen District, Muang Yai St. 70 (alt. 470 m), 2.VIII.2019, R. Okada leg. (CRO) ; Lampang Province. 5♂♂, 9♀♀, Thoen District, Mae Pa St. 197 (alt. 200 m), 15.XI.2020, R. Okada leg. (CGW, CRO) ; Mae Hong Son Province. 3♂♂, Pai District, Mueang Paeng St. 93 (alt. 760 m), 1.XII.2019, R. Okada leg. (CRO) ; 3♂♂, 2♀♀, Pha Bong, ca. 12 km S Mae Hong Son, 12.XI.1995, H. Zettel leg (CGW, NMW); Phrae Province. 1♂, 15 km E Phrae, Mae Khaem, 16.XI.1995, H. Zettel leg. (NMW) ; Uttaradit Province. 2♂♂, 1♀, Nam Pat District, Saen To St. 184 (alt. 210 m), 13.IX.2020, R. Okada leg. (CRO) ; Phetchabun Province. 1♀, Huai Nam Phang, S Ban Nam Nao, 25.XI.1995, H. Zettel leg. (NMW) ; Bueng Kan Province. 2♂♂, Seka District, Ban Tong St. 206 (alt. 190 m), 27.XII.2020, R. Okada leg. (CGW, CRO) ; Nakhon Phanom Province. 12 exs, Bang Phaeng District, Phai Lom St. 203 (alt. 200 m), 27.XII.2020, R. Okada leg. (CGW, CRO) ; Sakon Nakhon Province. 1♀, Charoen Sin District, Huai Yang St. 208 (alt. 320 m), 29.XII.2020, R. Okada leg. (CRO) ; 2♂♂, 2♀♀, Tao Ngoi District, Nan Tan St. 210 (alt. 310 m), 29.XII.2020, R. Okada leg. (CGW, CRO) ; Ubon Ratchathani Province. 1♂, 1♀, Si Mueang Mai District, Nam Thaeng St. 221 (alt. 190 m), 26.II.2021, R. Okada leg. (CRO) ; 1♂, 1♀, Nam Yuen District, Dom Pradit St. 224 (alt. 190 m), 27.II.2021, R. Okada leg. (CRO) ; Kanchanaburi Province. 1♂, Si Sawat District, Na Suan St. 28 (alt. 310 m), 12.I.2019, R. Okada leg. (CRO) .</p><p>Laos: Luang Prabang Province. 4 exs., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.96667&amp;materialsCitation.latitude=19.583334" title="Search Plazi for locations around (long 101.96667/lat 19.583334)">Thong Khan</a>, 19°35'N, 101°58'E, alt. ca. 750 m, 11-12.V.2002, V. Kubáň leg. (CGW, NMB) .</p><p>Comments.</p><p>The lectotype of Microdytes maculatus (Fig. 41) has a rhomboid-like habitus similar to that of most specimens from Myanmar, northern Thailand, and China, while most specimens from northeastern Thailand and Laos have a more regularly rounded habitus, but this is not a discriminant character.</p><p>There is also a geographic variation in the shapes of the median lobe and the apical part of the parameres. In specimens from Myanmar, northern Thailand, and China, the two parts of the median lobe have obtuse apical medial angles (Fig. 42A), and the parameres have narrow tips (Fig. 43A), while in those of northeastern Thailand and western Laos the two parts of the median lobe have almost right apical medial angles (Fig. 42B) and the parameres have rounded tips (Fig. 43B). Because some specimens show an intermediate status and no other distinguishing character can be found, they are treated as geographic variations belonging to the same species. Molecular biological studies are required to establish if they are different species within a complex.</p><p>Distribution.</p><p>India: southern Andaman Islands; Myanmar: Chin and Shan states, Saganing Division; Thailand: Chiang Mai, Chiang Rai (first record), Lampang (first record), Lamphun, Mae Hong Son, Uttaradit (first record), Phetchabun (first record), Bueng Kan (first record), Mukdahan, Nakhon Phanom (first record), Sakon Nakhon (first record), Ubon Ratchathani (first record), and Kantchanaburi provinces, Khao Yai NP [Nakhon Ratchasima or Nakhon Nayok Province]; Laos: Vientiane, Luang Prabang (first record) and Khammouan provinces; China: Yunnan Province.</p></div>	https://treatment.plazi.org/id/E88287CB88D056EA9854CBECE0AAD8BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
F6E632D1B8EB551B9F043CBB656211DC.text	F6E632D1B8EB551B9F043CBB656211DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes mariannae Wewalka 1997	<div><p>Microdytes mariannae Wewalka, 1997</p><p>Fig. 26</p><p>Microdytes mariannae Wewalka, 1997: 28; Wewalka 2011: 30; Nilsson and Hájek 2023: 211.</p><p>Type locality.</p><p>Thailand, Phetchabun Province, Nam Nao.</p><p>Material examined.</p><p>Thailand: Phetchabun Province. 8♂♂, 10♀♀, Lom Kao District, Ban Noen St. 187 (alt. 1620 m), 20.IX.2020, R. Okada leg. (CGW, CRO, THNHM) (Fig. 26) ; 2♂♂, 2♀♀, same locality, St. 294, 18.VI.2022, R. Okada Leg. (CRO, THNHM) .</p><p>Distribution.</p><p>Thailand: Phetchabun and Loei provinces.</p></div>	https://treatment.plazi.org/id/F6E632D1B8EB551B9F043CBB656211DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
B2456B3F39C75B3982D2EEF6964E097D.text	B2456B3F39C75B3982D2EEF6964E097D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes maximiliani Wewalka & Okada 2023	<div><p>Microdytes maximiliani Wewalka &amp; Okada sp. nov.</p><p>Figs 4, 9, 14, 38</p><p>Type locality.</p><p>Laos, Louang Namtha Province, 10 km E Muang Sing, 21°09-10'N, 101°13-15'E, alt. 750-1400 m.</p><p>Type material.</p><p>Holotype (NMB): ♂, "Laos, Louang Namtha prov., 10 km E Muang Sing, 750-1400 m, Ban Oudonsinh B. Nam Det \ B. Nam Mai, 21°09-10'N \ 101°13-15'E, 14.-20.V.2011" \ "NHMB Basel Laos 2011 Expedition D. Hauk &amp; M. Geiser" [printed white labels] \ "HOLOTYPUS Microdytes maximiliani sp. nov. Wewalka &amp; Okada 2022" [printed red label]. Paratypes (4 exs.): 2♂♂, 1♀, with same data as the holotype (CGW, NMB, NMP); 1♀, China: Yunnan, Xishuangbanna, ca. 10 km NW Menglun, ca. 700-800 m, 7.XI.1999, M. Jäch et al. (CWBS 360) (NMW). All paratypes are provided with printed red paratype labels.</p><p>Diagnosis.</p><p>Microdytes maximiliani sp. nov. resembles M. satoi Wewalka, 1997 in coloration of pronotum and elytra but differs from this species by darker head, larger size, a distinct impression on lateral side in anterior third of elytron (Fig. 38) and male genitalia. It is similar to M. paoloi Wewalka, 2011 in size and coloration of pronotum but can be distinguished by elytral markings, the impression on lateral side of elytron and male genitalia. The shape of median lobe of aedeagus of M. maximiliani is very similar to that of M. schoenmanni Wewalka, 1997, but this species is much smaller and differs in coloration, punctation and a missing impression on lateral side of elytron.</p><p>Description.</p><p>Measurements. TL = 1.60-2.00 mm (1.86 mm), TL-h = 1.40-1.80 mm (1.61 mm), MW = 1.30-1.35 mm (1.30 mm), TL/MW = 1.23-1.43 (1.43). Body broad and regularly oval, quite convex (Figs 4, 38).</p><p>Coloration. Head dark reddish brown, darker on vertex and along eyes. Pronotum dark brown. Elytron dark brown with yellowish brown markings forming a distinct transverse band near base not reaching suture and lateral margin, a post-median lateral spot often connected with a triangular spot near apex (Figs 4, 9). Elytral epipleuron and ventral surface dark brown to black. Legs, antennae and palpi reddish brown.</p><p>Sculpture and structure. Head finely and sparsely punctured, stronger punctures along eyes and on vertex; with fine microreticulation missing on vertex; clypeus not bordered. Pronotum irregularly punctured with sparse fine punctures and strong punctures concentrated along posterior margin and sparsely on disc; without microreticulation; lateral margins finely bordered, scarcely rounded. Elytron quite regularly, moderately densely and fairly strongly punctured, with traces of two longitudinal rows of punctures; highly polished and shining; without microreticulation; with a distinct impression on lateral side in anterior third (Fig. 38). Ventral surface: metacoxae, metasternum and abdomen moderately strongly but very sparsely punctured; without microreticulation.</p><p>Male. Median lobe of aedeagus in ventral aspect slightly tapered from base to apex, expanded before apex, apex truncate (Fig. 14A); in lateral aspect straight, tapered in apical ninth, apex bent downwards (Fig. 14B). Parameres narrow triangular in lateral aspect (Fig. 14C, D).</p><p>Female. Without secondary sexual characters. Sclerotized spermatheca not found.</p><p>Variation.</p><p>Variation of markings is shown in Fig. 9.</p><p>Etymology.</p><p>This species is dedicated to Maximilian Bramböck, Vienna, Austria. The species epithet is a name in the genitive singular.</p><p>Habitat.</p><p>The species was collected in Yunnan in a 1-2-m wide stream flowing through dense primary forest ( Jäch and Ji 2003).</p><p>Distribution.</p><p>Laos: Louang Namtha Province; China: Yunnan Province.</p></div>	https://treatment.plazi.org/id/B2456B3F39C75B3982D2EEF6964E097D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
ABD36E1BAF395CFE9DCC61C1A970DD2C.text	ABD36E1BAF395CFE9DCC61C1A970DD2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes menopausis Wewalka 1997	<div><p>Microdytes menopausis Wewalka, 1997</p><p>Fig. 27</p><p>Microdytes menopausis Wewalka, 1997: 29; Wewalka 2011: 30; Nilsson and Hájek 2023: 211.</p><p>Type locality.</p><p>Thailand, Khao Yai NP.</p><p>Material examined.</p><p>Thailand: Nakhon Phanom Province. 1♂, 2♀♀, Bang Phaeng District, Phai Lom St. 203 (alt. 200 m), 27.XII.2020, R. Okada leg. (CGW, CRO) ; Sakon Nakhon Province. 1♀, Tao Ngoi District, Nan Tan St. 210 (alt. 310 m), 29.XII.2020, R. Okada leg. (CRO) (Fig. 27) ; Ubon Ratchathani Province. 1♀, Nam Yuen District, Dom Pradit St. 224 (alt. 190 m), 27.II.2021, R. Okada leg. (CGW) ; 1♂, Si Mueang Mai District, Nam Thaeng St. 241 (alt. 190 m), 23.V.2021, R. Okada leg. (THNHM) .</p><p>Laos: Khammouan Province. 1♂, Nakai-Nam Theun NPA, Ban Navang env., 17°57-59'N, 105°13-16'E, alt. 600-750 m, 18.-21.V.2012, NHMB Basel , Laos 2012 Expedition, M. Brancucci, M. Geiser, K. Phanthavong &amp; S. Xayalath leg. (NMB); Savannakhet Province. 3 exs., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.63333&amp;materialsCitation.latitude=17.0" title="Search Plazi for locations around (long 105.63333/lat 17.0)">Phou Xang He</a> NBCA, ca. 5 km SW Ban Pa Phaknau, 17°00'N, 105°38'E, alt. 250-400 m, 31.V.-6.VI. 2011, NHMB Basel , Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala &amp; E. Vongphachan leg. (CGW, NMB) .</p><p>Distribution.</p><p>Thailand: Trat, Nakhon Phanom (first record), Sakon Nakhon (first record), and Ubon Ratchathani (first record) provinces, Khao Yai NP [Nakhon Ratchasima or Nakhon Nayok Province]; Laos: Khammouan, Savannakhet (first record) and Sekong provinces.</p></div>	https://treatment.plazi.org/id/ABD36E1BAF395CFE9DCC61C1A970DD2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
A442A1623EAA5C269C5E43F6FE525BFC.text	A442A1623EAA5C269C5E43F6FE525BFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes paoloi Wewalka 2011	<div><p>Microdytes paoloi Wewalka, 2011</p><p>Fig. 28</p><p>Microdytes paoloi Wewalka, 2011: 24; Nilsson and Hájek 2023: 211.</p><p>Type locality.</p><p>Thailand, Phetchabun Province, Phu Hin Rongkla NP.</p><p>Material examined.</p><p>Thailand: Phetchabun Province. 1♂, paratypus, Phu Hin Rongkla NP, small stream, alt. 1250 m (4), 25.XII.1999, Mazzoldi leg. (CMW) (Fig. 28) .</p><p>Distribution.</p><p>Thailand: Phetchabun and Loei provinces.</p></div>	https://treatment.plazi.org/id/A442A1623EAA5C269C5E43F6FE525BFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
556A83E1850053D881226F8B74C93A32.text	556A83E1850053D881226F8B74C93A32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes pasiricus (Csiki 1938)	<div><p>Microdytes pasiricus (Csiki, 1938)</p><p>Fig. 29</p><p>Hydrovatus pasiricus Csiki, 1938: 126 (including var. Hydrovatus pasiricus simplicor and var. Hydrovatus pasiricus unicolor).</p><p>Microdytes pasiricus: Biström et al. 1997: 75; Wewalka 1997: 30; Wewalka 2011: 30; Freitag et. al. 2016: 185; Nilsson and Hájek 2023: 212.</p><p>Type locality.</p><p>Indonesia, Central Java, Sarangan near Lake Pasir.</p><p>Material examined.</p><p>Thailand: Chiang Rai Province. 3♂♂, Wiang Kaen District, Muang Yai St. 71 (alt. 440 m), 11.VIII.2019, R. Okada leg. (CGW, CRO) (Fig. 29) ; Chiang Mai Province. 2♀♀, Mae Chaem District, Tha Pha St. 166 (alt. 720 m), 15.VIII.2020, R. Okada leg. (CRO, THNHM) .</p><p>Distribution.</p><p>Thailand: Chiang Rai (first record), Chiang Mai (first record) and Phetchabun provinces; Philippines: Busuanga, Luzon, Mindoro, Palawan; Singapore; Indonesia: Java.</p></div>	https://treatment.plazi.org/id/556A83E1850053D881226F8B74C93A32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
197593CBCB385EBCBE29E151501763E4.text	197593CBCB385EBCBE29E151501763E4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes pederzanii Wewalka 2011	<div><p>Microdytes pederzanii Wewalka, 2011</p><p>Fig. 30</p><p>Microdytes pederzanii Wewalka, 2011: 25; Nilsson and Hájek 2023: 212.</p><p>Type locality.</p><p>Thailand, Phetchabun Province, Phu Hin Rongkla NP.</p><p>Material examined.</p><p>Thailand: Phetchabun Province. 2♀♀, Lom Kao District, Ban Noen St. 187 (alt. 1620 m), 20.IX.2020, R. Okada leg. (CRO) ; Uttaradit Province. 1♂, 1♀, Nam Pad District, Phu Soi Dao NP, pine forest, 12.VII.2020, T. Jeenthong leg. (THNHM) (Fig. 30) .</p><p>Distribution.</p><p>Thailand: Phetchabun and Uttaradit (first record) provinces.</p></div>	https://treatment.plazi.org/id/197593CBCB385EBCBE29E151501763E4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0A4BB3A407CE548D9DAED53A6D2C6516.text	0A4BB3A407CE548D9DAED53A6D2C6516.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes rocchii Wewalka 2011	<div><p>Microdytes rocchii Wewalka, 2011</p><p>Fig. 31</p><p>Microdytes rocchii Wewalka, 2011: 26; Nilsson and Hájek 2023: 212.</p><p>Type locality.</p><p>Laos, Khammuan Province, Ban Khoun Ngeun.</p><p>Material examined.</p><p>Laos: Khammouan Province. 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.63333&amp;materialsCitation.latitude=17.0" title="Search Plazi for locations around (long 105.63333/lat 17.0)">Ban</a> Khon Ngeun, 18°07'N, 104°29'E, alt. ca 200 m, 19.-31.V.2001, Pacholátko leg. (CGW) (Fig. 31); Savannakhet Province. 1♂, Phou Xang He NBCA, ca. 5 km SW Ban Pa Phaknau, 17°00'N, 105°38'E, alt. 250-400 m, 31.V.-6.VI. 2011, NHMB Basel , Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala &amp; E. Vongphachan leg. (NMB) .</p><p>Distribution.</p><p>Laos: Khammuan and Savannakhet (first record) provinces.</p></div>	https://treatment.plazi.org/id/0A4BB3A407CE548D9DAED53A6D2C6516	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
C58B04105D2657D09BCA3D874C15E5E9.text	C58B04105D2657D09BCA3D874C15E5E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes schoedli Wewalka 1997	<div><p>Microdytes schoedli Wewalka, 1997</p><p>Figs 32, 39B</p><p>Microdytes schoedli Wewalka, 1997: 33; Wewalka 2011: 31; Nilsson and Hájek 2023: 212.</p><p>Type locality.</p><p>Thailand, Phetchabun Province, Nam Nao NP.</p><p>Material examined.</p><p>Thailand: Chiang Mai Province. 2♀♀, Chiang Dao District, Ping Khong St. 226 (alt. 430 m), 20.III.2021, R. Okada leg. (CRO) ; Lampang Province. 1♂, 1♀, Thoen District, Mae Pa St. 197 (alt. 200 m), 15.XI.2020, R. Okada leg. (CRO) ; Mae Hong Son Province. 1♂, Pai District, Thung Yao St. 50 (alt. 610 m), 15.VI.2019, R. Okada leg. (CRO) ; 2♂♂, 2♀♀, Muang Mae Hong Son District, Pha Bong St. 250 (alt. 480 m), 13.VI.2021, R. Okada leg. (CGW, CRO) (Fig. 32) ; Phetchabun Province. 1♂, 2♀♀, Lom Sak District, Nam Chun St. 296 (alt. 200 m), 19.VI.2022, R. Okada leg. (CRO) ; Kanchanaburi Province. 2♂♂, 1♀, Si Sawat District, Khao Chot St. 27 (alt. 420 m), 12.I.2019, R. Okada leg. (CRO) ; 1♀, same district, Na Suan St. 28 (alt. 310 m), 12.I.2019, R. Okada leg. (CRO) .</p><p>Laos: Khammouan Province. 1♂, 2♀♀, Nakai-NamTheun NPA, Ban Navang env., 17°57-59'N, 105°13-16'E, alt. 600-750 m, 18.-21.V.2012, NHMB Basel , Laos 2012 Expedition, M. Brancucci, M. Geiser, K. Phanthavong &amp; S. Xayalath leg. (NMB); Savannakhet Province. 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.63333&amp;materialsCitation.latitude=17.0" title="Search Plazi for locations around (long 105.63333/lat 17.0)">Phou Xang He</a> NBCA, ca. 5 km SW Ban Pa Phaknau, 17°00'N, 105°38'E, alt. 250-400 m, 31.V.-6.VI. 2011, NHMB Basel , Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala &amp; E. Vongphachan leg. (NMB) .</p><p>Distribution.</p><p>Thailand: Chiang Mai, Lampang (first record), Mae Hong Son (first record), Phetchabun, Tak, Mukdahan, and Kanchanaburi (first record) provinces; Laos: Khammouan (first record), Savannakhet (first record) and Sekong provinces.</p></div>	https://treatment.plazi.org/id/C58B04105D2657D09BCA3D874C15E5E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E191489899C25EDA81A2D993849DADE4.text	E191489899C25EDA81A2D993849DADE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes schoenmanni Wewalka 1997	<div><p>Microdytes schoenmanni Wewalka, 1997</p><p>Fig. 33</p><p>Microdytes schoenmanni Wewalka, 1997: 34; Miller and Wewalka 2010: 35; Wewalka 2011: 31; Nilsson and Hájek 2023: 212.</p><p>Type locality.</p><p>Thailand, Trat Province, Koh Chang Island, Klong Prao.</p><p>Material examined.</p><p>Myanmar: 1♂, Shan State, N Aungban, halfway between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.541466&amp;materialsCitation.latitude=20.971184" title="Search Plazi for locations around (long 96.541466/lat 20.971184)">Pindaya</a> and Ye’ngan, 20°58.271'N, 96°32.488'E, ca. alt. 1241 m, stream, 10.VI.2004, Shaverdo leg. (CGW) (Fig. 33) .</p><p>Distribution.</p><p>India: Darjeeling; Nepal; Myanmar: Chin and Shan states, Saganing and Tanintharyi divisions; Thailand: Trat Province, Khao Yai NP [Nakhon Ratchasima or Nakhon Nayok Province]; Laos: Attapeu Province; China: Yunnan Province.</p></div>	https://treatment.plazi.org/id/E191489899C25EDA81A2D993849DADE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
B70D2C12B4DC50A0B553DCC9BD80B24A.text	B70D2C12B4DC50A0B553DCC9BD80B24A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes schwendingeri Wewalka 1997	<div><p>Microdytes schwendingeri Wewalka, 1997</p><p>Figs 34, 40B</p><p>Microdytes schwendingeri Wewalka, 1997: 36; Wewalka 2011: 37; Nilsson and Hájek 2023: 212.</p><p>Type locality.</p><p>Thailand, Sakon Nakhon Province, Phu Pan NP.</p><p>Material examined.</p><p>Thailand: Nakhon Phanom Province. 1♂, 1♀, Bang Phaeng District, Phai Lom St. 203 (alt. 200 m), 27.XII.2020, R. Okada leg. (CRO, THNHM) (Fig. 34) .</p><p>Laos: Savannakhet Province, 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.63333&amp;materialsCitation.latitude=17.0" title="Search Plazi for locations around (long 105.63333/lat 17.0)">Phou Xang He</a> NBCA, ca. 5 km SW Ban Pa Phaknau, 17°00'N, 105°38'E, alt. 250-400 m, 31.V.-6.VI. 2011, NHMB Basel , Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala &amp; E. Vongphachan leg. (NMB); Champasak Province. 2♂♂, 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.64667&amp;materialsCitation.latitude=15.135" title="Search Plazi for locations around (long 106.64667/lat 15.135)">Bolavens Plateau</a>, waterfall ca. 2 km E Tad Katamtok, 15°08.1'N, 106°38.8'E, alt. 415 m, 10.-12.V.2010, J. Hájek leg. (CGW, NMP) .</p><p>Comments.</p><p>Some specimens from Laos have less distinct elytral markings but no other differences from typical specimens have been observed.</p><p>Distribution.</p><p>Thailand: Nakhon Phanom (first record) and Sakon Nakhon provinces; Laos (first record): Savannakhet and Champasak provinces.</p></div>	https://treatment.plazi.org/id/B70D2C12B4DC50A0B553DCC9BD80B24A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
1D47CF0C5B8854CDAE37D1797CB021E7.text	1D47CF0C5B8854CDAE37D1797CB021E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes sekaensis Okada & Wewalka 2023	<div><p>Microdytes sekaensis Okada &amp; Wewalka sp. nov.</p><p>Figs 5, 10, 15</p><p>Type locality.</p><p>Thailand, Bueng Kan Province, Seka District, Ban Tong, 18°08'16"N, 103°59'35"E, alt. 190 m.</p><p>Type material.</p><p>Holotype (THNHM): ♂, "THAI: Bueng Kan Seka Dist., Ban Tong St. 206 (alt. 190 m) 28.XII.2020 R. Okada leg. \ HOLOTYPE Microdytes sekaensis sp. nov. Okada &amp; Wewalka 2022" [printed red label]. Paratypes (37 exs.): 1♂, with same data as the holotype (CGW); 36 exs., Laos, Bolikhamsay Province, Nam Kading NPA, Tad Paloy campsite, 18°21-23'N, 104°09'E, alt. 250-400 m, 24.-28.V.2011, NHMB Basel Laos 2011 Expedition, M. Geiser, D. Hauk, A. Phantala &amp; E. Vongphachan leg. (CGW, CRO, NMB, NMP). All paratypes are provided with printed red paratype labels.</p><p>Diagnosis.</p><p>Microdytes sekaensis sp. nov. resembles M. hainanensis Wewalka, 1997 and M. schwendingeri Wewalka, 1997 (Figs 34, 40B) in habitus and size but differs by elytral markings and clypeus not bordered. Size and coloration of M. sekaensis sp. nov. is also similar to those of M. gabrielae Wewalka, 1997 (Fig. 23), but it is distinguishable from the latter by the more rounded-oval body and finely and sparsely punctured ventral surface.</p><p>Description.</p><p>Measurements. TL = 1.34-1.36 mm (1.36 mm), TL-h = 0.92-0.93 mm (0.93 mm), MW = 0.99-1.01 mm (1.01 mm), TL/MW = 1.35-1.36 (1.35). Body broadly oval, strongly convex (Fig. 5).</p><p>Coloration. Head yellowish brown. Pronotum yellowish brown, along posterior margin narrowly dark brown. Elytron reddish brown to dark brown with two yellowish brown spots: one basal spot not reaching base, one anterior lateral spot continuing along the lateral margin to a post-median lateral spot and a transverse spot near apex, also with a round median spot near suture (Figs 5, 10). Ventral surface predominantly yellowish brown. Legs, antennae and palpi yellowish brown.</p><p>Sculpture and structure. Head very finely and sparsely, slightly irregularly punctured; entirely and distinctly microreticulate; clypeus not bordered. Pronotum quite irregularly, finely, and sparsely punctured on the disc, additionally with coarser punctures at medial part of posterior margin; without microreticulation; lateral margins finely bordered, regularly rounded. Elytron fairly finely, regularly, and moderately densely punctured, progressively finer and sparser towards lateral margin; two longitudinal rows of punctures moderately distinct; highly polished and shining; without microreticulation. Ventral surface: metacoxae and metasternum finely and sparsely punctured, epipleura and abdomen almost without punctures; without microreticulation.</p><p>Male. Median lobe of aedeagus in ventral aspect slightly tapered from base to apex and narrowly constricted before apex, apex spoon shaped (Fig. 15A); in lateral aspect slightly curved with an arrowhead shaped apical expansion (Fig. 15B). Paramere narrowly trapezoid in lateral aspect (Fig. 15C, D).</p><p>Female. Without secondary sexual characters. Sclerotized spermatheca not found.</p><p>Variation.</p><p>Variation of markings is shown in Fig. 10.</p><p>Etymology.</p><p>This species is named after the district name of the type locality.</p><p>Habitat.</p><p>This species was collected in the remaining water pools in a dried-up stream which had a sandy bottom with rich leaf deposits (Fig. 47).</p><p>Distribution.</p><p>Thailand: Bueng Kan Province; Laos: Bolikhamsay Province.</p></div>	https://treatment.plazi.org/id/1D47CF0C5B8854CDAE37D1797CB021E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
FAAD1C24D633582CAA7DE3F55B9F8B09.text	FAAD1C24D633582CAA7DE3F55B9F8B09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes shepardi Wewalka 1997	<div><p>Microdytes shepardi Wewalka, 1997</p><p>Fig. 35</p><p>Microdytes shepardi Wewalka, 1997: 37; Wewalka 2011: 33; Nilsson and Hájek 2023: 212.</p><p>Type locality.</p><p>Thailand, Phetchabun Province, Nam Nao NP.</p><p>Material examined.</p><p>Thailand: Chiang Mai Province. 1♂, 1♀, Chiang Dao District, Ping Khong St. 226 (alt. 430 m), 20.III.2021, R. Okada leg. (CRO) ; Chiang Rai Province. 1♂, Wiang Kaen District, Muang Yai St. 71 (alt. 440 m), 2.VIII.2019, R. Okada leg. (CRO) ; Uthai Thani Province. 5♂♂, 2♀♀, Ban Rai District, Kaen Makrut St. 128 (alt. 440 m), 20.VI.2020, R. Okada leg. (CGW, CRO, THNHM) (Fig. 35) .</p><p>Laos: 1♂, Luang Prabang Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.96667&amp;materialsCitation.latitude=19.583334" title="Search Plazi for locations around (long 101.96667/lat 19.583334)">Thong Khan</a>, 19°35'N, 101°58'E, alt. ca. 750 m, 11.-12.V.2002, V. Kubáň leg. (NMB) .</p><p>Distribution.</p><p>Thailand: Chiang Mai, Chiang Rai (first record), Mae Hong Son, Phetchabun, Uthai Thani (first record), Chaiyaphum and Sakon Nakhon provinces; Laos: Luang Prabang Province; China: Yunnan Province.</p></div>	https://treatment.plazi.org/id/FAAD1C24D633582CAA7DE3F55B9F8B09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
44C6F2709CAB5C19B4AC75C7B962567D.text	44C6F2709CAB5C19B4AC75C7B962567D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes shunichii Sato 1995	<div><p>Microdytes shunichii Sato, 1995</p><p>Fig. 36</p><p>Microdytes shunichii Satô, 1995: 313; Wewalka 1997: 38; Wewalka 2011: 33; Nilsson and Hájek 2023: 212.</p><p>Microdytes holzmanni Wewalka &amp; Wang, 1998: 66.</p><p>Microdytes holzmannorum: Nilsson 2007: 51 (as unjustified emendation of Microdytes holzmanni).</p><p>Type locality.</p><p>Vietnam, Vinh Phuc Province, Mt. Tam Dao.</p><p>Material examined.</p><p>Thailand: Phetchabun Province. 1♀, Lom Kao District, Ban Noen St. 187 (alt. 1620 m), 20.IX.2020, R. Okada leg. (CRO) ; 3♂♂, 2♀♀, same locality, St. 294, 18.VI.2022, R. Okada leg. (CGW, CRO, THNHM) .</p><p>Laos: Luang Namtha Province. 62 exs., 10 km E Muang Sing, Ban Oudomsinh / B. Nam Det / B. Nam Mai, 21°09-10'N, 101°13-15'E, 750-1400 m, 14.-20.V.2011, NHMB Basel, Laos 2011 Expedition, D. Hauk &amp; M. Geiser (CGW, CRO, NMB, NMP) (Fig. 36); Luang Prabang Province. 1♂, 5 km W Ban Song Cha, 20°33-34'N, 102°14'E, 1200 m, 24.-30. IV.1999, V. Kubáň leg. (NMB); Khammouan Province. 2♂♂, Nakai-Nam Theun NPA, Ban Navang env., 17°57-59'N, 105°13-16'E, alt. 600-750 m, 18.-21.V.2012, NHMB Basel , Laos 2012 Expedition, M. Brancucci, M. Geiser, K. Phanthavong &amp; S. Xayalath leg. (NMB); Savannakhet Province. 1♀, Phou Xang He NBCA, ca. 5 km SW Ban Pa Phaknau, 17°00'N, 105°38'E, alt. 250-400 m, 31.V. - 6.VI. 2011, NHMB <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.63333&amp;materialsCitation.latitude=17.0" title="Search Plazi for locations around (long 105.63333/lat 17.0)">Basel</a>, Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala &amp; E. Vongphachan leg. (CGW, NMB, NMP); Attapeu Province. 1♂, 1♀, Dong Amphan National Bio-Diversity Conservation Area, Nong Fa (crater lake) env., 15°05.9'N, 107°25.6'E, alt. ca. 1160 m, 30.IV.-6.V.2010, J. Hájek leg. (NMP); 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.42667&amp;materialsCitation.latitude=15.098333" title="Search Plazi for locations around (long 107.42667/lat 15.098333)">Thong Kai Ohk</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.42667&amp;materialsCitation.latitude=15.098333" title="Search Plazi for locations around (long 107.42667/lat 15.098333)">Ban Kachung</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.42667&amp;materialsCitation.latitude=15.098333" title="Search Plazi for locations around (long 107.42667/lat 15.098333)">Mai</a>) env., 15°01-02'N, 107°26-27'E, 1200-1450 m, 10.-24.VI.2011, NHMB <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.42667&amp;materialsCitation.latitude=15.098333" title="Search Plazi for locations around (long 107.42667/lat 15.098333)">Basel</a> Laos 2011 Expedition, M. Geiser, D. Hauk, A. Phantala &amp; E. Vongphachan leg. (NMB) .</p><p>Distribution.</p><p>Thailand: Nan, Chiang Mai, and Phetchabun (first record) provinces; Laos: Phongsali (first record), Luang Namtha (first record), Oudomxai, Luang Prabang (first record), Vientiane, Khammouan, Savannakhet (first record), and Attapeu (first record) provinces; China: Yunnan Province, Hong Kong; Vietnam: Vinh Phuc Province.</p></div>	https://treatment.plazi.org/id/44C6F2709CAB5C19B4AC75C7B962567D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E631B55B5C055D5EBBEF1A3FD015833B.text	E631B55B5C055D5EBBEF1A3FD015833B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes ubonensis Okada & Wewalka 2023	<div><p>Microdytes ubonensis Okada &amp; Wewalka sp. nov.</p><p>Figs 6, 11, 16, 40A</p><p>Type locality.</p><p>Thailand, Ubon Ratchathani Province, Si Mueang Mai Dist., Nam Thaeng, 15°32'19"N, 105°24'08"E, alt. 200 m.</p><p>Type material.</p><p>Holotype (THNHM): ♂, "THAI: Ubon Ratchathani Si Mueang Mai Dist., Nam Thaeng St. 242 (alt. 200 m) 23.V.2021 R. Okada leg." [printed white label] / HOLOTYPE Microdytes ubonensis sp. nov. Okada &amp; Wewalka 2022" [printed red label]. Paratypes (52 exs.): 3♂♂, 1♀, with same data as the holotype (CGW, CRO); 4♀♀, Thailand, Ubon Ratchatani Province, Nam Yuen Dist., Dom Pradit St. 221 (alt. 190 m), 14.402°N, 104.215°E, 26.II.2021 R Okada leg. (THNHM); 4♂♂, 4♀♀, Thailand, Ubon Ratchathani Province, Nam Yuen District, Dom Pradit St. 240 (alt. 260 m), 14°23'22"N, 105°09'25"E, 22.V.2021, R. Okada leg. (CRO); 18 exs., Laos, Savannakhet Province, Phou Xang He NBCA, ca. 5 km SW Ban Pa Phaknau, 17°00'N, 105°38'E, alt. 250-400 m, 31.V.-6.VI. 2011, NHMB Basel, Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala &amp; E. Vongphachan leg. (CGW, NMB, NMP); 1♂, Laos, Champasak Province, Bolavens Plateau, waterfall ca. 2 km E Tad Katamtok, 15°08.1'N, 106°38.8'E, alt. 415 m, 10.-12.V.2010, J. Hájek leg. (NMP); 15 exs., Laos, Champasak Province, Bolavens Plateau, ca. 1 km S Ban Lak 40 [vill.] Tad Yueang waterfall, 15°10.8'N, 106°08.3'E, alt. 900-970 m, 28.IV.2010, J. Hájek leg. (CGW, CRO, NMP); 1♂, Laos, Vientiane Province, Phou Khao Khouay NBCA ca. 46 km N Vientiane (waterfall), 18°22.4'N, 102°42.4'E, alt. 270 m, 18.V.2010, J. Hájek leg. (NMP); 1♀, Laos, Vientiane Province, 90 km E Vientiane, Phou Khao Khouay NP, Nam Leuk, 1.-8.VI.1996, H. Schillhammer leg (NMW). All paratypes are provided with printed red paratype labels.</p><p>Diagnosis.</p><p>Microdytes ubonensis sp. nov. is similar to M. boukali Wewalka, 1997 and M. lotteae Wewalka, 1998 in habitus, size, and pronotal coloration but differs from these species by having yellowish head and the elytral markings with distinct post-median spot near suture and finely and sparsely punctured ventral surface. M. ubonensis sp. nov. also resembles M. huangyongensis Bian et al., 2015 in habitus and coloration but it differs in size and male genitalia.</p><p>Description.</p><p>Measurements. TL = 1.25-1.40 mm (1.37 mm), TL-h = 1.08-1.19 mm (1.18 mm), MW = 0.87-0.95 mm (0.95 mm), TL/MW = 1.42-1.47 (1.44). Body regularly oval, moderately convex (Fig. 6).</p><p>Coloration. Head yellowish brown. Pronotum dark brown, indistinctly yellowish brown at lateral sides. Elytron dark brown with yellowish brown markings forming a distinct transverse band at base not reaching suture, with a post-median lateral spot and a triangular spot near apex and often with a distinct longitudinal post-median spot near suture (Figs 6, 11). Ventral surface of head and prothorax yellowish brown; thorax, abdominal ventrites and elytral epipleuron reddish brown to dark brown. Legs, antennae and palpi yellowish brown.</p><p>Sculpture and structure. Head finely, sparsely, and relatively regularly punctured; almost entirely microreticulate; clypeus not bordered (Fig. 40A). Pronotum quite regularly, sparsely, and fairly strongly punctured, coarser punctures along the posterior margin; without microreticulation; lateral margins very finely bordered, regularly rounded. Elytron moderately regularly and densely punctured, progressively finer and sparser towards lateral margin; longitudinal rows of stronger punctures not distinct; highly polished and shining; without microreticulation. Ventral surface: metacoxae and metasternum finely and sparsely punctured, abdomen without punctures; without microreticulation.</p><p>Male. Median lobe of aedeagus in ventral aspect equally tapered from base to apex, with a small swelling at apex (Fig. 16A); in lateral aspect tapered from base to apex, with a small ventral hook at apex (Fig. 16B). Paramere narrow triangular in lateral aspect (Fig. 16C, D).</p><p>Female. Without secondary sexual characters. Sclerotized spermatheca not found.</p><p>Variation.</p><p>Variation of markings is shown in Fig. 11.</p><p>Etymology.</p><p>This species is named after the popular provincial name of the type locality.</p><p>Habitat.</p><p>This species was collected in a variety of habitats. At the type locality, specimens were collected from a small stream on a more or less deforested bedrock (Fig. 48). At other localities, specimens were collected in small pit holes near small forest streams with rich leaf deposits.</p><p>Distribution.</p><p>Thailand: Ubon Ratchathani Province; Laos: Savannakhet, Champasak, and Vientiane provinces.</p></div>	https://treatment.plazi.org/id/E631B55B5C055D5EBBEF1A3FD015833B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
4A95238CB1495BED9839DA08716D810B.text	4A95238CB1495BED9839DA08716D810B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes wewalkai Bian & Ji 2009	<div><p>Microdytes wewalkai Bian &amp; Ji, 2009</p><p>Microdytes wewalkai Bian &amp; Ji, 2009: 37; Wewalka 2011: 36; Nilsson and Hájek 2023: 212.</p><p>Type locality.</p><p>China, Hainan, Lingtou (Ledong) County, Jianfengling NR.</p><p>Material examined.</p><p>Laos: Attapeu Province. 3♂♂, 2♀♀, Dong Amphan National Bio-Diversity Conservation Area, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.42667&amp;materialsCitation.latitude=15.098333" title="Search Plazi for locations around (long 107.42667/lat 15.098333)">Nong Fa</a> (crater lake) env., 15°05.9'N, 107°25.6'E, alt. ca. 1160 m, 30.IV.-6.V.2010, J. Hájek leg. (CGW, NMP) .</p><p>Comments.</p><p>Specimens from Laos have two transverse yellowish red markings on both sides of the middle of pronotum while in specimens from China (Hainan) it is completely black, but no other differences have been observed.</p><p>Distribution.</p><p>Laos (first record): Attapeu Province; China: Hainan Province.</p></div>	https://treatment.plazi.org/id/4A95238CB1495BED9839DA08716D810B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
67258B2EE5335F90B5BAB80FA058D96B.text	67258B2EE5335F90B5BAB80FA058D96B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdytes zetteli Wewalka 1997	<div><p>Microdytes zetteli Wewalka, 1997</p><p>Fig. 37</p><p>Microdytes zetteli Wewalka, 1997: 41; Wewalka 2011: 34; Nilsson and Hájek 2023: 212.</p><p>Type locality.</p><p>Thailand, Chiang Mai Province, Doi Suthep.</p><p>Material examined.</p><p>Thailand: Chiang Mai Province. 12 exs., Mae Taeng District, Pa Pae St. 49 (alt. 1000 m), 15.VI.2019, R. Okada leg. (CGW, CRO, THNHM) ; 2♂♂, Mae Chaem District, Tha Pha St. 136 (alt. 720 m), 4.VII.2020, R. Okada leg. (CGW, CRO) ; 1♂, same locality St. 166 (alt. 720 m), 15.VIII.2020, R. Okada leg. (THNHM) (Fig. 37) ; Chiang Rai Province. 1♀, Wiang Kaen District, Muang Yai St. 71 (alt. 440 m), 2.VIII.2019, R. Okada leg. (CRO) .</p><p>Laos: 4 exs., Luang Prabang Province, 5 km W Ban Song Cha, 20°33-4'N, 102°14'E, alt. ca. 1200 m, 1.-16.V.1999, V. Kubáň leg. (CGW, NMB) .</p><p>Distribution.</p><p>Myanmar: Shan State; Thailand: Chiang Mai, Chiang Rai (first record) and Mae Hong Son provinces; Laos: Luang Prabang Province.</p><p>Additional species known from Thailand, Laos, or Cambodia</p></div>	https://treatment.plazi.org/id/67258B2EE5335F90B5BAB80FA058D96B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Okada, Ryohei;Jaitrong, Weeyawat;Wewalka, Guenther	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
