taxonID	type	description	language	source
34466794E3B35AA996FACD46E195FF5F.taxon	description	Fig. 18	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
34466794E3B35AA996FACD46E195FF5F.taxon	materials_examined	Material examined. Laos: Bolikhamsay Province. 22 exs., Nam Khading National Bio-Diversity Conservation Area, Tad Paloy campsite, 18 ° 23.17 ' N, 104 ° 09.65 ' E, alt. 300 m, forest stream, 8. - 11. VII. 2010 & 24. - 28. V. 2011, NHMB Basel Laos 2010 & 2011 Expeditions, M. Brancucci, M. Geiser, D. Hauk, A. Phantala & E. Vongphachan leg. (CGW, CRO, NMB, NMP) (Fig. 18).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
34466794E3B35AA996FACD46E195FF5F.taxon	distribution	Distribution. Laos: Vientiane and Bolikhamsay (first record) provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
1ADE86AE29DD50F28720770561AC9E3B.taxon	description	Figs 17, 19, 39 D	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
1ADE86AE29DD50F28720770561AC9E3B.taxon	materials_examined	Material examined. Thailand: Mukdahan Province. 1 ♂, Phu Pha Thoep N. P., small pools (23), 1. I. 2000, Mazzoldi leg. (CGW); Ubon Ratchathani Province. 1 ♂, 1 ♀, Nam Yuen District, Dom Pradit St. 240 (alt. 260 m), 22. V. 2021, R. Okada leg. (CRO, THNHM); Saraburi Province. 1 ♀, Kaeng Khoi District, Ched Khot St. 122 (alt. 140 m), 30. V. 2020, R. Okada leg. (CRO); Chonburi Province. 2 ♂♂, 1 ♀, Ban Bueng District, Khlong Kiu St. 261 (alt. 200 m), 6. XI. 2021, R. Okada leg. (CGW, CRO) (Fig. 17); 2 ♂♂, 1 ♀, Si Racha District, Bang Phra St. 269 (alt. 90 m), 26. XII. 2021, R. Okada leg. (CRO); Rayong Province. 4 ♂♂, Wang Chan District, Pa Yup Nai St 299 (alt. 200 m), 17. VII. 2022, R. Okada leg. (CRO, THNHM). Laos: Savannakhet Province. 203 exs., Phou Xang He NBCA, ca. 5 km SW Ban Pa Phaknau, 17 ° 00 ' N, 105 ° 38 ' E, alt. 250 - 400 m, 31. V. - 6. VI. 2011, NHMB Basel, Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala & E. Vongphachan leg. (CGW, CRO, NMB, NMP); Sekong Province. 3 ♀♀, ca. 51 km N Sekong (river) Ho Chi Minh trail, 15 ° 49.6 ' N, 106 ° 39.8 ' E, alt. 410 m, 14. - 15. V. 2010, J. Hajek leg. (NMP). Cambodia: Koh Hong Province. 12 exs., 20 km SW Koh Hong, Tatai River, 11 ° 34 ' N, 103 ° 07 ' E, alt. 50 - 300 m, 3. - 19. V. 2005, E. Jendek & O. Sausa leg. (CGW, NMB).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
1ADE86AE29DD50F28720770561AC9E3B.taxon	distribution	Distribution. Thailand: Mukdahan, Rayong, Trat, Ubon Ratchathani (first record), Saraburi (first record), and Chonburi (first record) provinces, Khao Yai NP [Nakhon Ratchatsima or Nakhon Nayok Province]; Laos (first record): Savannakhet and Sekong provinces; Cambodia (first record): Koh Hong Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0E028AD633F156FC96DC5E7C696DE758.taxon	description	Fig. 20	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0E028AD633F156FC96DC5E7C696DE758.taxon	materials_examined	Material examined. Thailand: Chonburi Province. 1 ♂, Si Racha District, Bang Phra St. 269 (alt. 90 m), 26. XII. 2020, R. Okada leg. (CRO) (Fig. 20).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0E028AD633F156FC96DC5E7C696DE758.taxon	distribution	Distribution. Thailand: Khao Yai NP [Nakhon Ratchasima or Nakhon Nayok Province] and Chonburi (first record) Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E3744614B5C25BF8985606CE116951EF.taxon	description	Figs 2, 7, 12, 42 C, 43 C	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E3744614B5C25BF8985606CE116951EF.taxon	diagnosis	Diagnosis. Microdytes eliasi sp. nov. very closely resembles M. maculatus (Motschulsky, 1860) (Figs 25, 41) in size and coloration but differs from this species by the more regularly oval habitus, the laterally expanded median lobe at apex and constricted tips of paramere (Figs 12, 42 C, 43 C) (see also comments in M. maculatus). It is also similar to M. feryi Wewalka, 2011 (Fig. 21) in habitus and coloration, but it is smaller and can be distinguished by male genitalia [compared with a male paratype from Myanmar, " Tenasserim, Birma Coll. V. Helfer National Museum Prague " (CGW)].	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E3744614B5C25BF8985606CE116951EF.taxon	description	Description. Measurements. TL = 1.64 - 1.85 mm (1.76 mm), TL-H = 1.49 - 1.65 mm (1.55 mm), MW = 1.12 - 1.25 mm (1.24 mm), TL / MW = 1.33 - 1.44 (1.42). Body regularly oval, moderately convex (Fig. 2). Coloration. Head reddish brown. Pronotum reddish brown, narrowly dark brown along anterior and posterior margins. Elytron dark brown with yellowish brown markings forming a distinct transverse band near base not reaching suture connected along lateral margin with a post-median transverse lateral band, with a small post-median spot near suture, and a triangular spot near apex (Figs 2, 7). Ventral surface of head, prothorax and elytral epipleuron yellowish brown; thorax and abdominal ventrites reddish brown to dark brown. Legs, antennae and palpi yellowish brown. Sculpture and structure. Head finely, sparsely, and relatively regularly punctured; anterior half to two-thirds finely microreticulate; clypeus not bordered. Pronotum quite regularly, sparsely, and fairly strongly punctured, with coarser punctures along posterior margin; without microreticulation; lateral margins finely bordered, regularly rounded. Elytron quite regularly, moderately densely and fairly strongly punctured, progressively finer and sparser towards lateral margin; without longitudinal rows of stronger punctures; highly polished and shining; without microreticulation. Ventral surface: metacoxae and metasternum strongly but sparsely punctured, abdomen finely and sparsely punctured; without microreticulation. Male. The two parts of the median lobe expanded laterally at apex in ventral aspect (Figs 12 A, 42 A); slightly curved in lateral aspect (Fig. 12 B). The tips of parameres twisted in lateral aspect (Fig. 12 C, D); constricted in ventral aspect (Fig. 43 C). Female. Without secondary sexual characters. Sclerotized spermatheca not found.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E3744614B5C25BF8985606CE116951EF.taxon	etymology	Etymology. This species is dedicated to Elias Bramboeck, Vienna, Austria. The species epithet is a name in the genitive singular.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E3744614B5C25BF8985606CE116951EF.taxon	distribution	Distribution. Thailand: Saraburi, Sa Kaeo, Chonburi, and Chanthaburi provinces; Cambodia: Koh Hong Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
31E418F324E952669D9F2B64F15A6BC2.taxon	description	Fig. 22	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
31E418F324E952669D9F2B64F15A6BC2.taxon	materials_examined	Material examined. Laos: Bolikhamsay Province. 20 exs., Nam Khading National Bio-Diversity Conservation Area, Tad Paloy campsite, 18 ° 23.17 ' N, 104 ° 09.65 ' E, alt. 300 m, forest stream, 8. - 11. VII. 2010 & 24. - 28. V. 2011, NHMB Basel Laos 2010 & 2011 Expeditions, M. Brancucci, M. Geiser, D. Hauk, A. Phantala & E. Vongphachan leg. (CGW, CRO, NMB, NMP) (Fig. 22).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
31E418F324E952669D9F2B64F15A6BC2.taxon	distribution	Distribution. Laos: Vientiane and Bolikhamsay (first record) provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
7F29A223BC0C56BEB0FC38B35495A93B.taxon	description	Fig. 23	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
7F29A223BC0C56BEB0FC38B35495A93B.taxon	materials_examined	Material examined. Thailand: Phetchabun Province. 1 ♂, Phu Hin Rongkla NP, small stream (7), 25. XII. 1999, Mazzoldi leg. (CGW) (Fig. 23); 4 ♂♂, 3 ♀♀, Lom Kao District, Ban Noen St. 294 (alt. 1620 m), 18. VI. 2022, R. Okada leg. (CGW, CRO, THNHM); Phitsanulok Province. 1 ♂, Nakhon Thai Distr., Phu Hin Rong Kla NP, in waterfall, 16 ° 59 ' 49.1 " N, 101 ° 00 ' 34.8 " E, 7. III. 2016, A. Damaska leg. (NMP).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
7F29A223BC0C56BEB0FC38B35495A93B.taxon	distribution	Distribution. Thailand: Phetchabun and Phitsanulok provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
5BA38C25DE725916ACB8998D206BB050.taxon	description	Figs 24, 39 A	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
5BA38C25DE725916ACB8998D206BB050.taxon	materials_examined	Material examined. Thailand: Chiang Mai Province. 8 exs., Mae Taeng District, Pa Pae St. 49 (alt. 1000 m), 15. VI. 2019, R. Okada leg. (CGW, CRO, THNHM); 1 ♂, same district, Kuet Chang St. 170 (alt. 520 m), 16. VIII. 2020, R. Okada leg. (CRO) (Fig. 24).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
5BA38C25DE725916ACB8998D206BB050.taxon	distribution	Distribution. Thailand: Nan and Chiang Mai (first record) provinces; Laos: Luang Namtha and Luang Prabang provinces; China: Yunnan Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0A7211761CF1522295AA7A35CFED94ED.taxon	description	Figs 3, 8, 13, 39 E	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0A7211761CF1522295AA7A35CFED94ED.taxon	diagnosis	Diagnosis. Microdytes jeenthongi sp. nov. resembles M. shunichii Sato, 1995 (Fig. 36) and M. zetteli Wewalka, 1997 (Fig. 37) in habitus, coloration and very finely punctured metasternum and metacoxae. From M. shunichii it differs by the reddish brown pronotum and very finely and sparsely punctured elytra. From M. zetteli it can be distinguished by larger size, less finely and sparsely punctured head.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0A7211761CF1522295AA7A35CFED94ED.taxon	description	Description. Measurements. TL = 1.77 - 1.82 mm (1.79 mm), TL-H = 1.23 - 1.26 mm (1.26 mm), MW = 1.18 - 1.23 mm (1.23 mm), TL / MW = 1.46 - 1.50 (1.49). Body regularly oval, moderately convex (Fig. 3). Coloration. Head reddish brown. Pronotum reddish brown, along the anterior margin narrowly and along the posterior margin more widely dark brown, especially medially. Elytron dark brown with a distinct transverse band at the base not reaching suture, continuing along the margin to a post-median transverse band and a triangular spot near the apex, and a small round post-median spot near suture (Figs 3, 8). Ventral surface of head and prothorax yellowish brown; thorax, abdominal ventrites and elytral epipleuron reddish brown to dark brown. Legs, antennae and palpi yellowish brown. Sculpture and structure. Head finely and sparsely, slightly irregularly punctured; anterior forth finely microreticulate; clypeus not bordered. Pronotum punctured moderately irregularly, finely, and sparsely, very coarsely along posterior margin; without microreticulation; lateral margins finely bordered, regularly rounded. Elytron very sparsely and finely punctured, progressively finer and sparser towards lateral margin; one longitudinal row of punctures distinct; highly polished and shining; without microreticulation. Ventral surface: metasternum and metacoxae very finely, sparsely, and irregularly punctured; abdomen almost without punctures; without microreticulation. Male. The two parts of the median lobe expanded and pointed at apex forming the unique arrowhead shape in ventral aspect (Fig. 13 A); almost straight and slightly expanded downwards at apex in lateral aspect (Fig. 13 B). Parameres moderate triangular in lateral aspect (Figs 13 C, D). Female. Unknown.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0A7211761CF1522295AA7A35CFED94ED.taxon	etymology	Etymology. This species is dedicated to Tadsanai Jeenthong, collection manager of National Science Museum, Thailand. The species epithet is a name in the genitive singular.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0A7211761CF1522295AA7A35CFED94ED.taxon	distribution	Distribution. Thailand: Chiang Mai Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E88287CB88D056EA9854CBECE0AAD8BB.taxon	description	Figs 25, 39 C, 41	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E88287CB88D056EA9854CBECE0AAD8BB.taxon	materials_examined	Material examined. Thailand: Chiang Mai Province. 6 ♂♂, 1 ♀, Chom Thong District, Ban Luang St. 165 (alt. 360 m), 15. VIII. 2020, R. Okada leg. (CGW, CRO) (Fig. 25); 9 exs., Mae Chaem District, Tha Pha St. 166 (alt. 720 m), 15. VIII. 2020, R. Okada leg. (CGW, CRO); 1 ♂, 1 ♀, Chiang Dao District, Ping Khong St. 226 (alt. 430 m), 20. III. 2021, R. Okada leg. (CRO); Chiang Rai Province. 3 ♀♀, Wiang Kaen District, Muang Yai St. 70 (alt. 470 m), 2. VIII. 2019, R. Okada leg. (CRO); Lampang Province. 5 ♂♂, 9 ♀♀, Thoen District, Mae Pa St. 197 (alt. 200 m), 15. XI. 2020, R. Okada leg. (CGW, CRO); Mae Hong Son Province. 3 ♂♂, Pai District, Mueang Paeng St. 93 (alt. 760 m), 1. XII. 2019, R. Okada leg. (CRO); 3 ♂♂, 2 ♀♀, Pha Bong, ca. 12 km S Mae Hong Son, 12. XI. 1995, H. Zettel leg (CGW, NMW); Phrae Province. 1 ♂, 15 km E Phrae, Mae Khaem, 16. XI. 1995, H. Zettel leg. (NMW); Uttaradit Province. 2 ♂♂, 1 ♀, Nam Pat District, Saen To St. 184 (alt. 210 m), 13. IX. 2020, R. Okada leg. (CRO); Phetchabun Province. 1 ♀, Huai Nam Phang, S Ban Nam Nao, 25. XI. 1995, H. Zettel leg. (NMW); Bueng Kan Province. 2 ♂♂, Seka District, Ban Tong St. 206 (alt. 190 m), 27. XII. 2020, R. Okada leg. (CGW, CRO); Nakhon Phanom Province. 12 exs, Bang Phaeng District, Phai Lom St. 203 (alt. 200 m), 27. XII. 2020, R. Okada leg. (CGW, CRO); Sakon Nakhon Province. 1 ♀, Charoen Sin District, Huai Yang St. 208 (alt. 320 m), 29. XII. 2020, R. Okada leg. (CRO); 2 ♂♂, 2 ♀♀, Tao Ngoi District, Nan Tan St. 210 (alt. 310 m), 29. XII. 2020, R. Okada leg. (CGW, CRO); Ubon Ratchathani Province. 1 ♂, 1 ♀, Si Mueang Mai District, Nam Thaeng St. 221 (alt. 190 m), 26. II. 2021, R. Okada leg. (CRO); 1 ♂, 1 ♀, Nam Yuen District, Dom Pradit St. 224 (alt. 190 m), 27. II. 2021, R. Okada leg. (CRO); Kanchanaburi Province. 1 ♂, Si Sawat District, Na Suan St. 28 (alt. 310 m), 12. I. 2019, R. Okada leg. (CRO). Laos: Luang Prabang Province. 4 exs., Thong Khan, 19 ° 35 ' N, 101 ° 58 ' E, alt. ca. 750 m, 11 - 12. V. 2002, V. Kuban leg. (CGW, NMB).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E88287CB88D056EA9854CBECE0AAD8BB.taxon	distribution	Distribution. India: southern Andaman Islands; Myanmar: Chin and Shan states, Saganing Division; Thailand: Chiang Mai, Chiang Rai (first record), Lampang (first record), Lamphun, Mae Hong Son, Uttaradit (first record), Phetchabun (first record), Bueng Kan (first record), Mukdahan, Nakhon Phanom (first record), Sakon Nakhon (first record), Ubon Ratchathani (first record), and Kantchanaburi provinces, Khao Yai NP [Nakhon Ratchasima or Nakhon Nayok Province]; Laos: Vientiane, Luang Prabang (first record) and Khammouan provinces; China: Yunnan Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
F6E632D1B8EB551B9F043CBB656211DC.taxon	description	Fig. 26	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
F6E632D1B8EB551B9F043CBB656211DC.taxon	materials_examined	Material examined. Thailand: Phetchabun Province. 8 ♂♂, 10 ♀♀, Lom Kao District, Ban Noen St. 187 (alt. 1620 m), 20. IX. 2020, R. Okada leg. (CGW, CRO, THNHM) (Fig. 26); 2 ♂♂, 2 ♀♀, same locality, St. 294, 18. VI. 2022, R. Okada Leg. (CRO, THNHM).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
F6E632D1B8EB551B9F043CBB656211DC.taxon	distribution	Distribution. Thailand: Phetchabun and Loei provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
B2456B3F39C75B3982D2EEF6964E097D.taxon	description	Figs 4, 9, 14, 38	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
B2456B3F39C75B3982D2EEF6964E097D.taxon	diagnosis	Diagnosis. Microdytes maximiliani sp. nov. resembles M. satoi Wewalka, 1997 in coloration of pronotum and elytra but differs from this species by darker head, larger size, a distinct impression on lateral side in anterior third of elytron (Fig. 38) and male genitalia. It is similar to M. paoloi Wewalka, 2011 in size and coloration of pronotum but can be distinguished by elytral markings, the impression on lateral side of elytron and male genitalia. The shape of median lobe of aedeagus of M. maximiliani is very similar to that of M. schoenmanni Wewalka, 1997, but this species is much smaller and differs in coloration, punctation and a missing impression on lateral side of elytron.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
B2456B3F39C75B3982D2EEF6964E097D.taxon	description	Description. Measurements. TL = 1.60 - 2.00 mm (1.86 mm), TL-h = 1.40 - 1.80 mm (1.61 mm), MW = 1.30 - 1.35 mm (1.30 mm), TL / MW = 1.23 - 1.43 (1.43). Body broad and regularly oval, quite convex (Figs 4, 38). Coloration. Head dark reddish brown, darker on vertex and along eyes. Pronotum dark brown. Elytron dark brown with yellowish brown markings forming a distinct transverse band near base not reaching suture and lateral margin, a post-median lateral spot often connected with a triangular spot near apex (Figs 4, 9). Elytral epipleuron and ventral surface dark brown to black. Legs, antennae and palpi reddish brown. Sculpture and structure. Head finely and sparsely punctured, stronger punctures along eyes and on vertex; with fine microreticulation missing on vertex; clypeus not bordered. Pronotum irregularly punctured with sparse fine punctures and strong punctures concentrated along posterior margin and sparsely on disc; without microreticulation; lateral margins finely bordered, scarcely rounded. Elytron quite regularly, moderately densely and fairly strongly punctured, with traces of two longitudinal rows of punctures; highly polished and shining; without microreticulation; with a distinct impression on lateral side in anterior third (Fig. 38). Ventral surface: metacoxae, metasternum and abdomen moderately strongly but very sparsely punctured; without microreticulation. Male. Median lobe of aedeagus in ventral aspect slightly tapered from base to apex, expanded before apex, apex truncate (Fig. 14 A); in lateral aspect straight, tapered in apical ninth, apex bent downwards (Fig. 14 B). Parameres narrow triangular in lateral aspect (Fig. 14 C, D). Female. Without secondary sexual characters. Sclerotized spermatheca not found.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
B2456B3F39C75B3982D2EEF6964E097D.taxon	etymology	Etymology. This species is dedicated to Maximilian Bramboeck, Vienna, Austria. The species epithet is a name in the genitive singular.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
B2456B3F39C75B3982D2EEF6964E097D.taxon	distribution	Distribution. Laos: Louang Namtha Province; China: Yunnan Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
ABD36E1BAF395CFE9DCC61C1A970DD2C.taxon	description	Fig. 27	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
ABD36E1BAF395CFE9DCC61C1A970DD2C.taxon	materials_examined	Material examined. Thailand: Nakhon Phanom Province. 1 ♂, 2 ♀♀, Bang Phaeng District, Phai Lom St. 203 (alt. 200 m), 27. XII. 2020, R. Okada leg. (CGW, CRO); Sakon Nakhon Province. 1 ♀, Tao Ngoi District, Nan Tan St. 210 (alt. 310 m), 29. XII. 2020, R. Okada leg. (CRO) (Fig. 27); Ubon Ratchathani Province. 1 ♀, Nam Yuen District, Dom Pradit St. 224 (alt. 190 m), 27. II. 2021, R. Okada leg. (CGW); 1 ♂, Si Mueang Mai District, Nam Thaeng St. 241 (alt. 190 m), 23. V. 2021, R. Okada leg. (THNHM). Laos: Khammouan Province. 1 ♂, Nakai-Nam Theun NPA, Ban Navang env., 17 ° 57 - 59 ' N, 105 ° 13 - 16 ' E, alt. 600 - 750 m, 18. - 21. V. 2012, NHMB Basel, Laos 2012 Expedition, M. Brancucci, M. Geiser, K. Phanthavong & S. Xayalath leg. (NMB); Savannakhet Province. 3 exs., Phou Xang He NBCA, ca. 5 km SW Ban Pa Phaknau, 17 ° 00 ' N, 105 ° 38 ' E, alt. 250 - 400 m, 31. V. - 6. VI. 2011, NHMB Basel, Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala & E. Vongphachan leg. (CGW, NMB).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
ABD36E1BAF395CFE9DCC61C1A970DD2C.taxon	distribution	Distribution. Thailand: Trat, Nakhon Phanom (first record), Sakon Nakhon (first record), and Ubon Ratchathani (first record) provinces, Khao Yai NP [Nakhon Ratchasima or Nakhon Nayok Province]; Laos: Khammouan, Savannakhet (first record) and Sekong provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
A442A1623EAA5C269C5E43F6FE525BFC.taxon	description	Fig. 28	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
A442A1623EAA5C269C5E43F6FE525BFC.taxon	materials_examined	Material examined. Thailand: Phetchabun Province. 1 ♂, paratypus, Phu Hin Rongkla NP, small stream, alt. 1250 m (4), 25. XII. 1999, Mazzoldi leg. (CMW) (Fig. 28).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
A442A1623EAA5C269C5E43F6FE525BFC.taxon	distribution	Distribution. Thailand: Phetchabun and Loei provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
556A83E1850053D881226F8B74C93A32.taxon	description	Fig. 29	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
556A83E1850053D881226F8B74C93A32.taxon	materials_examined	Material examined. Thailand: Chiang Rai Province. 3 ♂♂, Wiang Kaen District, Muang Yai St. 71 (alt. 440 m), 11. VIII. 2019, R. Okada leg. (CGW, CRO) (Fig. 29); Chiang Mai Province. 2 ♀♀, Mae Chaem District, Tha Pha St. 166 (alt. 720 m), 15. VIII. 2020, R. Okada leg. (CRO, THNHM).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
556A83E1850053D881226F8B74C93A32.taxon	distribution	Distribution. Thailand: Chiang Rai (first record), Chiang Mai (first record) and Phetchabun provinces; Philippines: Busuanga, Luzon, Mindoro, Palawan; Singapore; Indonesia: Java.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
197593CBCB385EBCBE29E151501763E4.taxon	description	Fig. 30	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
197593CBCB385EBCBE29E151501763E4.taxon	materials_examined	Material examined. Thailand: Phetchabun Province. 2 ♀♀, Lom Kao District, Ban Noen St. 187 (alt. 1620 m), 20. IX. 2020, R. Okada leg. (CRO); Uttaradit Province. 1 ♂, 1 ♀, Nam Pad District, Phu Soi Dao NP, pine forest, 12. VII. 2020, T. Jeenthong leg. (THNHM) (Fig. 30).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
197593CBCB385EBCBE29E151501763E4.taxon	distribution	Distribution. Thailand: Phetchabun and Uttaradit (first record) provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0A4BB3A407CE548D9DAED53A6D2C6516.taxon	description	Fig. 31	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0A4BB3A407CE548D9DAED53A6D2C6516.taxon	materials_examined	Material examined. Laos: Khammouan Province. 1 ♀, Ban Khon Ngeun, 18 ° 07 ' N, 104 ° 29 ' E, alt. ca 200 m, 19. - 31. V. 2001, Pacholatko leg. (CGW) (Fig. 31); Savannakhet Province. 1 ♂, Phou Xang He NBCA, ca. 5 km SW Ban Pa Phaknau, 17 ° 00 ' N, 105 ° 38 ' E, alt. 250 - 400 m, 31. V. - 6. VI. 2011, NHMB Basel, Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala & E. Vongphachan leg. (NMB).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
0A4BB3A407CE548D9DAED53A6D2C6516.taxon	distribution	Distribution. Laos: Khammuan and Savannakhet (first record) provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
C58B04105D2657D09BCA3D874C15E5E9.taxon	description	Figs 32, 39 B	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
C58B04105D2657D09BCA3D874C15E5E9.taxon	materials_examined	Material examined. Thailand: Chiang Mai Province. 2 ♀♀, Chiang Dao District, Ping Khong St. 226 (alt. 430 m), 20. III. 2021, R. Okada leg. (CRO); Lampang Province. 1 ♂, 1 ♀, Thoen District, Mae Pa St. 197 (alt. 200 m), 15. XI. 2020, R. Okada leg. (CRO); Mae Hong Son Province. 1 ♂, Pai District, Thung Yao St. 50 (alt. 610 m), 15. VI. 2019, R. Okada leg. (CRO); 2 ♂♂, 2 ♀♀, Muang Mae Hong Son District, Pha Bong St. 250 (alt. 480 m), 13. VI. 2021, R. Okada leg. (CGW, CRO) (Fig. 32); Phetchabun Province. 1 ♂, 2 ♀♀, Lom Sak District, Nam Chun St. 296 (alt. 200 m), 19. VI. 2022, R. Okada leg. (CRO); Kanchanaburi Province. 2 ♂♂, 1 ♀, Si Sawat District, Khao Chot St. 27 (alt. 420 m), 12. I. 2019, R. Okada leg. (CRO); 1 ♀, same district, Na Suan St. 28 (alt. 310 m), 12. I. 2019, R. Okada leg. (CRO). Laos: Khammouan Province. 1 ♂, 2 ♀♀, Nakai-NamTheun NPA, Ban Navang env., 17 ° 57 - 59 ' N, 105 ° 13 - 16 ' E, alt. 600 - 750 m, 18. - 21. V. 2012, NHMB Basel, Laos 2012 Expedition, M. Brancucci, M. Geiser, K. Phanthavong & S. Xayalath leg. (NMB); Savannakhet Province. 1 ♂, Phou Xang He NBCA, ca. 5 km SW Ban Pa Phaknau, 17 ° 00 ' N, 105 ° 38 ' E, alt. 250 - 400 m, 31. V. - 6. VI. 2011, NHMB Basel, Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala & E. Vongphachan leg. (NMB).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
C58B04105D2657D09BCA3D874C15E5E9.taxon	distribution	Distribution. Thailand: Chiang Mai, Lampang (first record), Mae Hong Son (first record), Phetchabun, Tak, Mukdahan, and Kanchanaburi (first record) provinces; Laos: Khammouan (first record), Savannakhet (first record) and Sekong provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E191489899C25EDA81A2D993849DADE4.taxon	description	Fig. 33	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E191489899C25EDA81A2D993849DADE4.taxon	materials_examined	Material examined. Myanmar: 1 ♂, Shan State, N Aungban, halfway between Pindaya and Ye'ngan, 20 ° 58.271 ' N, 96 ° 32.488 ' E, ca. alt. 1241 m, stream, 10. VI. 2004, Shaverdo leg. (CGW) (Fig. 33).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E191489899C25EDA81A2D993849DADE4.taxon	distribution	Distribution. India: Darjeeling; Nepal; Myanmar: Chin and Shan states, Saganing and Tanintharyi divisions; Thailand: Trat Province, Khao Yai NP [Nakhon Ratchasima or Nakhon Nayok Province]; Laos: Attapeu Province; China: Yunnan Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
B70D2C12B4DC50A0B553DCC9BD80B24A.taxon	description	Figs 34, 40 B	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
B70D2C12B4DC50A0B553DCC9BD80B24A.taxon	materials_examined	Material examined. Thailand: Nakhon Phanom Province. 1 ♂, 1 ♀, Bang Phaeng District, Phai Lom St. 203 (alt. 200 m), 27. XII. 2020, R. Okada leg. (CRO, THNHM) (Fig. 34). Laos: Savannakhet Province, 1 ♀, Phou Xang He NBCA, ca. 5 km SW Ban Pa Phaknau, 17 ° 00 ' N, 105 ° 38 ' E, alt. 250 - 400 m, 31. V. - 6. VI. 2011, NHMB Basel, Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala & E. Vongphachan leg. (NMB); Champasak Province. 2 ♂♂, 1 ♀, Bolavens Plateau, waterfall ca. 2 km E Tad Katamtok, 15 ° 08.1 ' N, 106 ° 38.8 ' E, alt. 415 m, 10. - 12. V. 2010, J. Hajek leg. (CGW, NMP).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
B70D2C12B4DC50A0B553DCC9BD80B24A.taxon	distribution	Distribution. Thailand: Nakhon Phanom (first record) and Sakon Nakhon provinces; Laos (first record): Savannakhet and Champasak provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
1D47CF0C5B8854CDAE37D1797CB021E7.taxon	description	Figs 5, 10, 15	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
1D47CF0C5B8854CDAE37D1797CB021E7.taxon	diagnosis	Diagnosis. Microdytes sekaensis sp. nov. resembles M. hainanensis Wewalka, 1997 and M. schwendingeri Wewalka, 1997 (Figs 34, 40 B) in habitus and size but differs by elytral markings and clypeus not bordered. Size and coloration of M. sekaensis sp. nov. is also similar to those of M. gabrielae Wewalka, 1997 (Fig. 23), but it is distinguishable from the latter by the more rounded-oval body and finely and sparsely punctured ventral surface.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
1D47CF0C5B8854CDAE37D1797CB021E7.taxon	description	Description. Measurements. TL = 1.34 - 1.36 mm (1.36 mm), TL-h = 0.92 - 0.93 mm (0.93 mm), MW = 0.99 - 1.01 mm (1.01 mm), TL / MW = 1.35 - 1.36 (1.35). Body broadly oval, strongly convex (Fig. 5). Coloration. Head yellowish brown. Pronotum yellowish brown, along posterior margin narrowly dark brown. Elytron reddish brown to dark brown with two yellowish brown spots: one basal spot not reaching base, one anterior lateral spot continuing along the lateral margin to a post-median lateral spot and a transverse spot near apex, also with a round median spot near suture (Figs 5, 10). Ventral surface predominantly yellowish brown. Legs, antennae and palpi yellowish brown. Sculpture and structure. Head very finely and sparsely, slightly irregularly punctured; entirely and distinctly microreticulate; clypeus not bordered. Pronotum quite irregularly, finely, and sparsely punctured on the disc, additionally with coarser punctures at medial part of posterior margin; without microreticulation; lateral margins finely bordered, regularly rounded. Elytron fairly finely, regularly, and moderately densely punctured, progressively finer and sparser towards lateral margin; two longitudinal rows of punctures moderately distinct; highly polished and shining; without microreticulation. Ventral surface: metacoxae and metasternum finely and sparsely punctured, epipleura and abdomen almost without punctures; without microreticulation. Male. Median lobe of aedeagus in ventral aspect slightly tapered from base to apex and narrowly constricted before apex, apex spoon shaped (Fig. 15 A); in lateral aspect slightly curved with an arrowhead shaped apical expansion (Fig. 15 B). Paramere narrowly trapezoid in lateral aspect (Fig. 15 C, D). Female. Without secondary sexual characters. Sclerotized spermatheca not found.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
1D47CF0C5B8854CDAE37D1797CB021E7.taxon	etymology	Etymology. This species is named after the district name of the type locality.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
1D47CF0C5B8854CDAE37D1797CB021E7.taxon	distribution	Distribution. Thailand: Bueng Kan Province; Laos: Bolikhamsay Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
FAAD1C24D633582CAA7DE3F55B9F8B09.taxon	description	Fig. 35	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
FAAD1C24D633582CAA7DE3F55B9F8B09.taxon	materials_examined	Material examined. Thailand: Chiang Mai Province. 1 ♂, 1 ♀, Chiang Dao District, Ping Khong St. 226 (alt. 430 m), 20. III. 2021, R. Okada leg. (CRO); Chiang Rai Province. 1 ♂, Wiang Kaen District, Muang Yai St. 71 (alt. 440 m), 2. VIII. 2019, R. Okada leg. (CRO); Uthai Thani Province. 5 ♂♂, 2 ♀♀, Ban Rai District, Kaen Makrut St. 128 (alt. 440 m), 20. VI. 2020, R. Okada leg. (CGW, CRO, THNHM) (Fig. 35). Laos: 1 ♂, Luang Prabang Province, Thong Khan, 19 ° 35 ' N, 101 ° 58 ' E, alt. ca. 750 m, 11. - 12. V. 2002, V. Kuban leg. (NMB).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
FAAD1C24D633582CAA7DE3F55B9F8B09.taxon	distribution	Distribution. Thailand: Chiang Mai, Chiang Rai (first record), Mae Hong Son, Phetchabun, Uthai Thani (first record), Chaiyaphum and Sakon Nakhon provinces; Laos: Luang Prabang Province; China: Yunnan Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
44C6F2709CAB5C19B4AC75C7B962567D.taxon	description	Fig. 36	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
44C6F2709CAB5C19B4AC75C7B962567D.taxon	materials_examined	Material examined. Thailand: Phetchabun Province. 1 ♀, Lom Kao District, Ban Noen St. 187 (alt. 1620 m), 20. IX. 2020, R. Okada leg. (CRO); 3 ♂♂, 2 ♀♀, same locality, St. 294, 18. VI. 2022, R. Okada leg. (CGW, CRO, THNHM). Laos: Luang Namtha Province. 62 exs., 10 km E Muang Sing, Ban Oudomsinh / B. Nam Det / B. Nam Mai, 21 ° 09 - 10 ' N, 101 ° 13 - 15 ' E, 750 - 1400 m, 14. - 20. V. 2011, NHMB Basel, Laos 2011 Expedition, D. Hauk & M. Geiser (CGW, CRO, NMB, NMP) (Fig. 36); Luang Prabang Province. 1 ♂, 5 km W Ban Song Cha, 20 ° 33 - 34 ' N, 102 ° 14 ' E, 1200 m, 24. - 30. IV. 1999, V. Kuban leg. (NMB); Khammouan Province. 2 ♂♂, Nakai-Nam Theun NPA, Ban Navang env., 17 ° 57 - 59 ' N, 105 ° 13 - 16 ' E, alt. 600 - 750 m, 18. - 21. V. 2012, NHMB Basel, Laos 2012 Expedition, M. Brancucci, M. Geiser, K. Phanthavong & S. Xayalath leg. (NMB); Savannakhet Province. 1 ♀, Phou Xang He NBCA, ca. 5 km SW Ban Pa Phaknau, 17 ° 00 ' N, 105 ° 38 ' E, alt. 250 - 400 m, 31. V. - 6. VI. 2011, NHMB Basel, Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala & E. Vongphachan leg. (CGW, NMB, NMP); Attapeu Province. 1 ♂, 1 ♀, Dong Amphan National Bio-Diversity Conservation Area, Nong Fa (crater lake) env., 15 ° 05.9 ' N, 107 ° 25.6 ' E, alt. ca. 1160 m, 30. IV. - 6. V. 2010, J. Hajek leg. (NMP); 1 ♀, Thong Kai Ohk, Ban Kachung (Mai) env., 15 ° 01 - 02 ' N, 107 ° 26 - 27 ' E, 1200 - 1450 m, 10. - 24. VI. 2011, NHMB Basel Laos 2011 Expedition, M. Geiser, D. Hauk, A. Phantala & E. Vongphachan leg. (NMB).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
44C6F2709CAB5C19B4AC75C7B962567D.taxon	distribution	Distribution. Thailand: Nan, Chiang Mai, and Phetchabun (first record) provinces; Laos: Phongsali (first record), Luang Namtha (first record), Oudomxai, Luang Prabang (first record), Vientiane, Khammouan, Savannakhet (first record), and Attapeu (first record) provinces; China: Yunnan Province, Hong Kong; Vietnam: Vinh Phuc Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E631B55B5C055D5EBBEF1A3FD015833B.taxon	description	Figs 6, 11, 16, 40 A	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E631B55B5C055D5EBBEF1A3FD015833B.taxon	diagnosis	Diagnosis. Microdytes ubonensis sp. nov. is similar to M. boukali Wewalka, 1997 and M. lotteae Wewalka, 1998 in habitus, size, and pronotal coloration but differs from these species by having yellowish head and the elytral markings with distinct post-median spot near suture and finely and sparsely punctured ventral surface. M. ubonensis sp. nov. also resembles M. huangyongensis Bian et al., 2015 in habitus and coloration but it differs in size and male genitalia.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E631B55B5C055D5EBBEF1A3FD015833B.taxon	description	Description. Measurements. TL = 1.25 - 1.40 mm (1.37 mm), TL-h = 1.08 - 1.19 mm (1.18 mm), MW = 0.87 - 0.95 mm (0.95 mm), TL / MW = 1.42 - 1.47 (1.44). Body regularly oval, moderately convex (Fig. 6). Coloration. Head yellowish brown. Pronotum dark brown, indistinctly yellowish brown at lateral sides. Elytron dark brown with yellowish brown markings forming a distinct transverse band at base not reaching suture, with a post-median lateral spot and a triangular spot near apex and often with a distinct longitudinal post-median spot near suture (Figs 6, 11). Ventral surface of head and prothorax yellowish brown; thorax, abdominal ventrites and elytral epipleuron reddish brown to dark brown. Legs, antennae and palpi yellowish brown. Sculpture and structure. Head finely, sparsely, and relatively regularly punctured; almost entirely microreticulate; clypeus not bordered (Fig. 40 A). Pronotum quite regularly, sparsely, and fairly strongly punctured, coarser punctures along the posterior margin; without microreticulation; lateral margins very finely bordered, regularly rounded. Elytron moderately regularly and densely punctured, progressively finer and sparser towards lateral margin; longitudinal rows of stronger punctures not distinct; highly polished and shining; without microreticulation. Ventral surface: metacoxae and metasternum finely and sparsely punctured, abdomen without punctures; without microreticulation. Male. Median lobe of aedeagus in ventral aspect equally tapered from base to apex, with a small swelling at apex (Fig. 16 A); in lateral aspect tapered from base to apex, with a small ventral hook at apex (Fig. 16 B). Paramere narrow triangular in lateral aspect (Fig. 16 C, D). Female. Without secondary sexual characters. Sclerotized spermatheca not found.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E631B55B5C055D5EBBEF1A3FD015833B.taxon	etymology	Etymology. This species is named after the popular provincial name of the type locality.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
E631B55B5C055D5EBBEF1A3FD015833B.taxon	distribution	Distribution. Thailand: Ubon Ratchathani Province; Laos: Savannakhet, Champasak, and Vientiane provinces.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
4A95238CB1495BED9839DA08716D810B.taxon	materials_examined	Material examined. Laos: Attapeu Province. 3 ♂♂, 2 ♀♀, Dong Amphan National Bio-Diversity Conservation Area, Nong Fa (crater lake) env., 15 ° 05.9 ' N, 107 ° 25.6 ' E, alt. ca. 1160 m, 30. IV. - 6. V. 2010, J. Hajek leg. (CGW, NMP).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
4A95238CB1495BED9839DA08716D810B.taxon	distribution	Distribution. Laos (first record): Attapeu Province; China: Hainan Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
67258B2EE5335F90B5BAB80FA058D96B.taxon	description	Fig. 37	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
67258B2EE5335F90B5BAB80FA058D96B.taxon	materials_examined	Material examined. Thailand: Chiang Mai Province. 12 exs., Mae Taeng District, Pa Pae St. 49 (alt. 1000 m), 15. VI. 2019, R. Okada leg. (CGW, CRO, THNHM); 2 ♂♂, Mae Chaem District, Tha Pha St. 136 (alt. 720 m), 4. VII. 2020, R. Okada leg. (CGW, CRO); 1 ♂, same locality St. 166 (alt. 720 m), 15. VIII. 2020, R. Okada leg. (THNHM) (Fig. 37); Chiang Rai Province. 1 ♀, Wiang Kaen District, Muang Yai St. 71 (alt. 440 m), 2. VIII. 2019, R. Okada leg. (CRO). Laos: 4 exs., Luang Prabang Province, 5 km W Ban Song Cha, 20 ° 33 - 4 ' N, 102 ° 14 ' E, alt. ca. 1200 m, 1. - 16. V. 1999, V. Kuban leg. (CGW, NMB).	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
67258B2EE5335F90B5BAB80FA058D96B.taxon	distribution	Distribution. Myanmar: Shan State; Thailand: Chiang Mai, Chiang Rai (first record) and Mae Hong Son provinces; Laos: Luang Prabang Province.	en	Okada, Ryohei, Jaitrong, Weeyawat, Wewalka, Guenther (2023): A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae). ZooKeys 1159: 87-119, DOI: http://dx.doi.org/10.3897/zookeys.1159.99218, URL: http://dx.doi.org/10.3897/zookeys.1159.99218
