taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
C8A5014707175FF7A796491D5BD7CAC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473088	https://doi.org/10.3897/vz.75.e167366.figure4	Figure 4. Photographs in life of Ichnotropis bivittata depicting the closely spaced yellow spots just posterior to the forelimbs. Specimens photographed at A Cuito town (PEM R 23530), Angola; B Cambau (FKH - 0833), Angola; C Cangandala National Park, Angola; D Luando Reserve (CHL 0675), Angola; and E Humpata (PEM R 17934 – I. b. pallida), Angola. Photographs: A – Werner Conradie; B, C, D – Pedro Vaz Pinto; E – William R. Branch.	Figure 4. Photographs in life of Ichnotropis bivittata depicting the closely spaced yellow spots just posterior to the forelimbs. Specimens photographed at A Cuito town (PEM R 23530), Angola; B Cambau (FKH - 0833), Angola; C Cangandala National Park, Angola; D Luando Reserve (CHL 0675), Angola; and E Humpata (PEM R 17934 – I. b. pallida), Angola. Photographs: A – Werner Conradie; B, C, D – Pedro Vaz Pinto; E – William R. Branch.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
C8A5014707175FF7A796491D5BD7CAC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473089	https://doi.org/10.3897/vz.75.e167366.figure5	Figure 5. Syntype (ZMB 5827) of Ichnotropis bivittata from Duque de Bragança [= Calandula], Malanje Province, Angola. Photographs of body in A dorsal and B ventral views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Frank Tillack.	Figure 5. Syntype (ZMB 5827) of Ichnotropis bivittata from Duque de Bragança [= Calandula], Malanje Province, Angola. Photographs of body in A dorsal and B ventral views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Frank Tillack.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
C8A5014707175FF7A796491D5BD7CAC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473090	https://doi.org/10.3897/vz.75.e167366.figure6	Figure 6. Holotype (BE _ RMCA _ Vert. R. 14641) of Ichnotropis capensis nigrescens from Bolobo, Democratic Republic of the Congo. Photographs of body in A dorsal and B ventral views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Max Benito.	Figure 6. Holotype (BE _ RMCA _ Vert. R. 14641) of Ichnotropis capensis nigrescens from Bolobo, Democratic Republic of the Congo. Photographs of body in A dorsal and B ventral views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Max Benito.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
C8A5014707175FF7A796491D5BD7CAC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473091	https://doi.org/10.3897/vz.75.e167366.figure7	Figure 7. Topotypic (PEM R 17934) Ichnotropis bivittata pallida from Humpata, Huila Province, Angola. Photographs of body in A dorsal and B ventral views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Werner Conradie.	Figure 7. Topotypic (PEM R 17934) Ichnotropis bivittata pallida from Humpata, Huila Province, Angola. Photographs of body in A dorsal and B ventral views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Werner Conradie.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
C8A5014707175FF7A796491D5BD7CAC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473086	https://doi.org/10.3897/vz.75.e167366.figure2	Figure 2. Above: Scatterplots of the first two principal components for Ichnotropis, based on the full Dataset 1 (A) and reduced Dataset 2 (B). Arrows indicate the loadings of morphological variables (see Materials and Methods for abbreviations). The green star marks the holotype of Ichnotropis capensis nigrescens (BE _ RMCA _ Vert. R. 14641), and the brown star marks the holotype of Ichnotropis overlaeti (BE _ RMCA _ Vert. R. 9691). Below: Box plots showing C snout – vent length (SVL), D head width (HW), and E head height (HH). Species are represented by different colours, and species groups are enclosed by polygons, as indicated in the legend.	Figure 2. Above: Scatterplots of the first two principal components for Ichnotropis, based on the full Dataset 1 (A) and reduced Dataset 2 (B). Arrows indicate the loadings of morphological variables (see Materials and Methods for abbreviations). The green star marks the holotype of Ichnotropis capensis nigrescens (BE _ RMCA _ Vert. R. 14641), and the brown star marks the holotype of Ichnotropis overlaeti (BE _ RMCA _ Vert. R. 9691). Below: Box plots showing C snout – vent length (SVL), D head width (HW), and E head height (HH). Species are represented by different colours, and species groups are enclosed by polygons, as indicated in the legend.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
C8A5014707175FF7A796491D5BD7CAC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473087	https://doi.org/10.3897/vz.75.e167366.figure3	Figure 3. Records of specimens from the Ichnotropis bivittata group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ib – I. bivittata, Ibp – I. bivittata pallida, Ic – I. chapini, Im – I. microlepidota, Icn – I. capensis nigrescens, Io – I. overlaeti and It – I. tanganicana.	Figure 3. Records of specimens from the Ichnotropis bivittata group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ib – I. bivittata, Ibp – I. bivittata pallida, Ic – I. chapini, Im – I. microlepidota, Icn – I. capensis nigrescens, Io – I. overlaeti and It – I. tanganicana.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
B08CEE791EB95AD8B7EB0ABAE6E2C34D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473104	https://doi.org/10.3897/vz.75.e167366.figure20	Figure 20. Photographs in life of Ichnotropis capensis sensu lato from A Cuando River source, Angola (adult male), B Cuanavale River source, Angola (adult female), C Xai-Xai, Mozambique (adult male), D Ngonye Falls, Zambia (adult female), E Zambezi Region, Namibia (adult male), F Kosi Bay, South Africa (adult male). Photographs: A – D – Werner Conradie; E, F – William R. Branch.	Figure 20. Photographs in life of Ichnotropis capensis sensu lato from A Cuando River source, Angola (adult male), B Cuanavale River source, Angola (adult female), C Xai-Xai, Mozambique (adult male), D Ngonye Falls, Zambia (adult female), E Zambezi Region, Namibia (adult male), F Kosi Bay, South Africa (adult male). Photographs: A – D – Werner Conradie; E, F – William R. Branch.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
B08CEE791EB95AD8B7EB0ABAE6E2C34D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473105	https://doi.org/10.3897/vz.75.e167366.figure21	Figure 21. Ichnotropis capensis sensu lato (PEM R 27396) from middle Quembo River bridge camp, Moxico Province, Angola. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Werner Conradie.	Figure 21. Ichnotropis capensis sensu lato (PEM R 27396) from middle Quembo River bridge camp, Moxico Province, Angola. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Werner Conradie.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
B08CEE791EB95AD8B7EB0ABAE6E2C34D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473106	https://doi.org/10.3897/vz.75.e167366.figure22	Figure 22. Syntype (ZMB 6123) of Ichnotropis macrolepidota from Lourenço Marques [= Maputo], Mozambique. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Frank Tillack.	Figure 22. Syntype (ZMB 6123) of Ichnotropis macrolepidota from Lourenço Marques [= Maputo], Mozambique. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Frank Tillack.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
B08CEE791EB95AD8B7EB0ABAE6E2C34D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473107	https://doi.org/10.3897/vz.75.e167366.figure23	Figure 23. Syntype (BMNH 1946.8. 4.23) of Ichnotropis longipes from Mazoë, Zimbabwe. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Patrick Campbell.	Figure 23. Syntype (BMNH 1946.8. 4.23) of Ichnotropis longipes from Mazoë, Zimbabwe. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Patrick Campbell.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
B08CEE791EB95AD8B7EB0ABAE6E2C34D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473103	https://doi.org/10.3897/vz.75.e167366.figure19	Figure 19. Geographic records of the Ichnotropis capensis group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ic – I. capensis, Ilc – I. longicorpa sp. nov., Ilp – I. longipes, and Im – I. macrolepidota.	Figure 19. Geographic records of the Ichnotropis capensis group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ic – I. capensis, Ilc – I. longicorpa sp. nov., Ilp – I. longipes, and Im – I. macrolepidota.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
284FB215FB1A56028F821F613482C363.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473092	https://doi.org/10.3897/vz.75.e167366.figure8	Figure 8. Holotype (AMNH 10674) of Ichnotropis chapini from Aba, Haut-Uele Province, Democratic Republic of the Congo. Photographs of body in A dorsal and B ventral views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Lauren Vonnahme.	Figure 8. Holotype (AMNH 10674) of Ichnotropis chapini from Aba, Haut-Uele Province, Democratic Republic of the Congo. Photographs of body in A dorsal and B ventral views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Lauren Vonnahme.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
284FB215FB1A56028F821F613482C363.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473086	https://doi.org/10.3897/vz.75.e167366.figure2	Figure 2. Above: Scatterplots of the first two principal components for Ichnotropis, based on the full Dataset 1 (A) and reduced Dataset 2 (B). Arrows indicate the loadings of morphological variables (see Materials and Methods for abbreviations). The green star marks the holotype of Ichnotropis capensis nigrescens (BE _ RMCA _ Vert. R. 14641), and the brown star marks the holotype of Ichnotropis overlaeti (BE _ RMCA _ Vert. R. 9691). Below: Box plots showing C snout – vent length (SVL), D head width (HW), and E head height (HH). Species are represented by different colours, and species groups are enclosed by polygons, as indicated in the legend.	Figure 2. Above: Scatterplots of the first two principal components for Ichnotropis, based on the full Dataset 1 (A) and reduced Dataset 2 (B). Arrows indicate the loadings of morphological variables (see Materials and Methods for abbreviations). The green star marks the holotype of Ichnotropis capensis nigrescens (BE _ RMCA _ Vert. R. 14641), and the brown star marks the holotype of Ichnotropis overlaeti (BE _ RMCA _ Vert. R. 9691). Below: Box plots showing C snout – vent length (SVL), D head width (HW), and E head height (HH). Species are represented by different colours, and species groups are enclosed by polygons, as indicated in the legend.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
284FB215FB1A56028F821F613482C363.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473087	https://doi.org/10.3897/vz.75.e167366.figure3	Figure 3. Records of specimens from the Ichnotropis bivittata group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ib – I. bivittata, Ibp – I. bivittata pallida, Ic – I. chapini, Im – I. microlepidota, Icn – I. capensis nigrescens, Io – I. overlaeti and It – I. tanganicana.	Figure 3. Records of specimens from the Ichnotropis bivittata group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ib – I. bivittata, Ibp – I. bivittata pallida, Ic – I. chapini, Im – I. microlepidota, Icn – I. capensis nigrescens, Io – I. overlaeti and It – I. tanganicana.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
F151507D3A5851ADB7FAAC88776348DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473099	https://doi.org/10.3897/vz.75.e167366.figure15	Figure 15. Photographs in life of juvenile Ichnotropis grandiceps. Specimens photographed from A Ngonye Falls (TM 86237), Zambia and B Chitokoloki, Zambia. Photographs: A – Darren Pietersen; B – Frank Willems.	Figure 15. Photographs in life of juvenile Ichnotropis grandiceps. Specimens photographed from A Ngonye Falls (TM 86237), Zambia and B Chitokoloki, Zambia. Photographs: A – Darren Pietersen; B – Frank Willems.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
F151507D3A5851ADB7FAAC88776348DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473100	https://doi.org/10.3897/vz.75.e167366.figure16	Figure 16. Ichnotropis grandiceps (RE 211206 D 1 / NMNW R 12212) specimen from Khaudum, Namibia. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Photographs: Werner Conradie.	Figure 16. Ichnotropis grandiceps (RE 211206 D 1 / NMNW R 12212) specimen from Khaudum, Namibia. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Photographs: Werner Conradie.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
F151507D3A5851ADB7FAAC88776348DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473097	https://doi.org/10.3897/vz.75.e167366.figure13	Figure 13. Holotype (BE _ RMCA _ Vert. R. 9691) of Ichnotropis overlaeti from Kapanga, Haut-Katanga Province, Democratic Republic of the Congo. Photographs of body in A ventral and B dorsal (note the evenly spaced white dorsolateral spots indicated by the arrows) views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Max Benito.	Figure 13. Holotype (BE _ RMCA _ Vert. R. 9691) of Ichnotropis overlaeti from Kapanga, Haut-Katanga Province, Democratic Republic of the Congo. Photographs of body in A ventral and B dorsal (note the evenly spaced white dorsolateral spots indicated by the arrows) views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Max Benito.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
08667256A08C5902B4CF3B9BB2A2F97A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473108	https://doi.org/10.3897/vz.75.e167366.figure24	Figure 24. Photographs in life of Ichnotropis longicorpa sp. nov. Photographs of A adult male (PEM R 23410) from Lungwebungu River camp bridge crossing, Angola, and B adult female (PEM R 23531) from Sombanana village, Angola. Photographs: Werner Conradie.	Figure 24. Photographs in life of Ichnotropis longicorpa sp. nov. Photographs of A adult male (PEM R 23410) from Lungwebungu River camp bridge crossing, Angola, and B adult female (PEM R 23531) from Sombanana village, Angola. Photographs: Werner Conradie.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
08667256A08C5902B4CF3B9BB2A2F97A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473109	https://doi.org/10.3897/vz.75.e167366.figure25	Figure 25. Holotype (PEM R 23410) of Ichnotropis longicorpa sp. nov. from Lungwebungu River camp bridge crossing, Moxico Province, Angola. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Werner Conradie.	Figure 25. Holotype (PEM R 23410) of Ichnotropis longicorpa sp. nov. from Lungwebungu River camp bridge crossing, Moxico Province, Angola. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Werner Conradie.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
08667256A08C5902B4CF3B9BB2A2F97A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473103	https://doi.org/10.3897/vz.75.e167366.figure19	Figure 19. Geographic records of the Ichnotropis capensis group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ic – I. capensis, Ilc – I. longicorpa sp. nov., Ilp – I. longipes, and Im – I. macrolepidota.	Figure 19. Geographic records of the Ichnotropis capensis group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ic – I. capensis, Ilc – I. longicorpa sp. nov., Ilp – I. longipes, and Im – I. macrolepidota.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
22C20657A1BE521C8EACCEAEF5DF86DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473093	https://doi.org/10.3897/vz.75.e167366.figure9	Figure 9. Photographs in life of Ichnotropis microlepidota (MHNCUP-REP 0983) from Serra do Moco, Huambo Province, Angola (adapted from Benito et al. 2025). Photographs in A dorsolateral view of the full body and B lateral view of the head. Photographs: Pedro Vaz Pinto.	Figure 9. Photographs in life of Ichnotropis microlepidota (MHNCUP-REP 0983) from Serra do Moco, Huambo Province, Angola (adapted from Benito et al. 2025). Photographs in A dorsolateral view of the full body and B lateral view of the head. Photographs: Pedro Vaz Pinto.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
22C20657A1BE521C8EACCEAEF5DF86DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473094	https://doi.org/10.3897/vz.75.e167366.figure10	Figure 10. Ichnotropis microlepidota (MHNCUP-REP 0983) specimen from Serra do Moco, Huambo Province, Angola (adapted from Benito et al. 2025). Photographs of body in A dorsal and B ventral views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Max Benito.	Figure 10. Ichnotropis microlepidota (MHNCUP-REP 0983) specimen from Serra do Moco, Huambo Province, Angola (adapted from Benito et al. 2025). Photographs of body in A dorsal and B ventral views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Max Benito.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
22C20657A1BE521C8EACCEAEF5DF86DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473087	https://doi.org/10.3897/vz.75.e167366.figure3	Figure 3. Records of specimens from the Ichnotropis bivittata group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ib – I. bivittata, Ibp – I. bivittata pallida, Ic – I. chapini, Im – I. microlepidota, Icn – I. capensis nigrescens, Io – I. overlaeti and It – I. tanganicana.	Figure 3. Records of specimens from the Ichnotropis bivittata group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ib – I. bivittata, Ibp – I. bivittata pallida, Ic – I. chapini, Im – I. microlepidota, Icn – I. capensis nigrescens, Io – I. overlaeti and It – I. tanganicana.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
59681C2BC5B251A2A570E125D6B18DE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473101	https://doi.org/10.3897/vz.75.e167366.figure17	Figure 17. Photographs in life of Ichnotropis robusta sp. nov. Photographs of A adult male (PEM R 23482) from Cuando River source, Angola; B adult female (PEM R 23362) from the road between Cuanavale River source camp and Samanunga village, Angola; C juvenile (PEM R 23304) from 4 km upstream of Cuanavale River source lake, Angola; and D juvenile (PEM R 23300) from the road between Cuanavale River source camp and Samanunga village, Angola. Photographs: A, B, C – Werner Conradie; D – Luke Verburgt.	Figure 17. Photographs in life of Ichnotropis robusta sp. nov. Photographs of A adult male (PEM R 23482) from Cuando River source, Angola; B adult female (PEM R 23362) from the road between Cuanavale River source camp and Samanunga village, Angola; C juvenile (PEM R 23304) from 4 km upstream of Cuanavale River source lake, Angola; and D juvenile (PEM R 23300) from the road between Cuanavale River source camp and Samanunga village, Angola. Photographs: A, B, C – Werner Conradie; D – Luke Verburgt.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
59681C2BC5B251A2A570E125D6B18DE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473102	https://doi.org/10.3897/vz.75.e167366.figure18	Figure 18. Holotype (PEM R 23420) of Ichnotropis robusta sp. nov. from Cuando River source, Moxico Province, Angola. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Werner Conradie.	Figure 18. Holotype (PEM R 23420) of Ichnotropis robusta sp. nov. from Cuando River source, Moxico Province, Angola. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Werner Conradie.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
59681C2BC5B251A2A570E125D6B18DE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473098	https://doi.org/10.3897/vz.75.e167366.figure14	Figure 14. Records of the Ichnotropis grandiceps group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ig – I. grandiceps, and Ir – I. robusta sp. nov.	Figure 14. Records of the Ichnotropis grandiceps group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ig – I. grandiceps, and Ir – I. robusta sp. nov.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
CBA5F90B078E5964B4D1B7B663DFDD50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473095	https://doi.org/10.3897/vz.75.e167366.figure11	Figure 11. Photographs in life of Ichnotropis tanganicana from across its range, depicting the evenly spaced dorsolateral blue spots. Photographs from A Rukwa, Tanzania; B Cambua, Democratic Republic of the Congo; C Rumphi, Malawi (https://www.inaturalist.org/observations/146895735); D Nyika National Park, Malawi (https://www.inaturalist.org/observations/146684850); E Kindingi, Lake Tanganyika (MTSN 9947), Democratic Republic of the Congo; and F Upemba National Park, Democratic Republic of the Congo (https://www.inaturalist.org/observations/249778421). Photographs: David Lloyd-Jones; B – Colin Tilbury, C – Marc Henrion, D – Tim Brammer, E – Wandege Muninga, D – Naftali Honig.	Figure 11. Photographs in life of Ichnotropis tanganicana from across its range, depicting the evenly spaced dorsolateral blue spots. Photographs from A Rukwa, Tanzania; B Cambua, Democratic Republic of the Congo; C Rumphi, Malawi (https://www.inaturalist.org/observations/146895735); D Nyika National Park, Malawi (https://www.inaturalist.org/observations/146684850); E Kindingi, Lake Tanganyika (MTSN 9947), Democratic Republic of the Congo; and F Upemba National Park, Democratic Republic of the Congo (https://www.inaturalist.org/observations/249778421). Photographs: David Lloyd-Jones; B – Colin Tilbury, C – Marc Henrion, D – Tim Brammer, E – Wandege Muninga, D – Naftali Honig.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
CBA5F90B078E5964B4D1B7B663DFDD50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473096	https://doi.org/10.3897/vz.75.e167366.figure12	Figure 12. Holotype (BMNH 1946.9. 3.49) of Ichnotropis tanganicana from ‘ East coast of Lake Tanganyika’, Tanzania. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Photographs: Patrick Campbell.	Figure 12. Holotype (BMNH 1946.9. 3.49) of Ichnotropis tanganicana from ‘ East coast of Lake Tanganyika’, Tanzania. Photographs of body in A ventral and B dorsal views, and head in C dorsal, D lateral and E ventral views. Photographs: Patrick Campbell.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
CBA5F90B078E5964B4D1B7B663DFDD50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473097	https://doi.org/10.3897/vz.75.e167366.figure13	Figure 13. Holotype (BE _ RMCA _ Vert. R. 9691) of Ichnotropis overlaeti from Kapanga, Haut-Katanga Province, Democratic Republic of the Congo. Photographs of body in A ventral and B dorsal (note the evenly spaced white dorsolateral spots indicated by the arrows) views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Max Benito.	Figure 13. Holotype (BE _ RMCA _ Vert. R. 9691) of Ichnotropis overlaeti from Kapanga, Haut-Katanga Province, Democratic Republic of the Congo. Photographs of body in A ventral and B dorsal (note the evenly spaced white dorsolateral spots indicated by the arrows) views, and head in C dorsal, D lateral and E ventral views. Scale bars represent 10 mm. Photographs: Max Benito.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
CBA5F90B078E5964B4D1B7B663DFDD50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1473087	https://doi.org/10.3897/vz.75.e167366.figure3	Figure 3. Records of specimens from the Ichnotropis bivittata group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ib – I. bivittata, Ibp – I. bivittata pallida, Ic – I. chapini, Im – I. microlepidota, Icn – I. capensis nigrescens, Io – I. overlaeti and It – I. tanganicana.	Figure 3. Records of specimens from the Ichnotropis bivittata group, based on all literature records (open circles), examined material (closed circles) and genetically analysed material (white centres). Respective type localities are indicated by arrows: Ib – I. bivittata, Ibp – I. bivittata pallida, Ic – I. chapini, Im – I. microlepidota, Icn – I. capensis nigrescens, Io – I. overlaeti and It – I. tanganicana.	2025-11-21	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke		Zenodo	biologists	Conradie, Werner;Keates, Chad;Greenbaum, Eli;Lobón-Rovira, Javier;Tolley, Krystal A.;Benito, Max;Vaz Pinto, Pedro;van Breda, Reuben V.;Verburgt, Luke			
