identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
71BBD70E0A835650AC93222401D43D0A.text	71BBD70E0A835650AC93222401D43D0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalipaludina delavayana (Heude 1889) Zhang & Du & von Rintelen 2023	<div><p>Dalipaludina delavayana (Heude, 1889) comb. nov.</p><p>Paludina delavayana Heude, 1889: 47 ("lacu Ta-li fou", Lake Erhai).</p><p>Sinotaia delavayana - Qian et al. 2014.</p><p>Material examined.</p><p>2 syntypes, IZCAS-FG-492553, IZCAS-FG-492554; 1 paratype, MCZ-Mala-167342; 8 specimens in ZMB, collected by Le-Jia Zhang from around 0.5 to 1 metres depth in the north shore of Lake Cibi in Augest 2018 , 2 specimens in ZMB, collected by Jiao-Wei Ning from around 1 to 2 metres depth in Lake Haixihai; 17 specimens in NJW, collected by Jiao-Wei Ning from around 1 to 2 metres depth in Lake Cibi .</p><p>Description.</p><p>Shell (Fig. 3A-H) large, conical, thin, greenish yellow to dark olive in colour; up to six whorls at adulthood, including one relatively smooth protoconch whorl, apex acute, spire high; each teleoconch whorls with one strong keel at suture, above suture smooth or with many weak spiral threads, sometimes with additional two to three stronger cords, base of shell with many thin weak spiral threads, weak spiral cords and threads with short periostracal hairs; aperture ovate, less than half of shell in height, lip thin and simple, umbilicus narrow, sometimes bordered by a keel.</p><p>Operculum (Fig. 3I) corneous, ovate, rather thin, yellow, with reddish brown nuclear region, exterior surface of operculum rather smooth, inner opercular region relatively small, nuclear region smooth.</p><p>Radular (Fig. 7A, B) central tooth with one broad central denticle and with four small sharp cusps on either side; lateral tooth with one broad central denticle and four small sharp cusps on either side; inner marginal tooth with one broad central denticle and three small sharp cusps on either side; outer marginal tooth with 10 to 11 small sharp cusps.</p><p>Remarks.</p><p>This species differs from the other Dalipaludina species by having a larger shell with a higher spiral. It may be difficult to distinguish from the smooth form of Dalipaludina oxytropoides from Lake Lugu based on morphology. However, this species is not recorded from Lake Lugu, and can be distinguished from D. oxytropoides based on differences in COI gene sequences. The recommended Chinese name of this species is “德拉维理田螺” .</p><p>Habitat and distribution.</p><p>shallow water area with sandy and muddy bottoms in Erhai, Cibi and Haixihai, Yunnan, China.</p></div>	https://treatment.plazi.org/id/71BBD70E0A835650AC93222401D43D0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Le-Jia;Du, Li-Na;von Rintelen, Thomas	Zhang, Le-Jia, Du, Li-Na, von Rintelen, Thomas (2023): A new genus of river snails, Dalipaludina (Gastropoda, Viviparidae), endemic to the Yunnan Plateau of SW China. Zoosystematics and Evolution 99 (2): 285-297, DOI: http://dx.doi.org/10.3897/zse.99.102586, URL: http://dx.doi.org/10.3897/zse.99.102586
BA528F39739C55BBA13C8BA168E48C31.text	BA528F39739C55BBA13C8BA168E48C31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalipaludina occidentalis (Annandale 1924) Zhang & Du & von Rintelen 2023	<div><p>Dalipaludina occidentalis (Annandale, 1924) comb. nov.</p><p>Lecythoconcha malleata f. occidentalis Annandale, 1924: 415 ("Ho-ching and Shihku", Heqing County and Shigu town, Lijiang City, Yunnan, China).</p><p>Viviparus occidentalis - Yen 1942.</p><p>Material examined.</p><p>8 specimens in ZMB, collected by local fishermen from 3 to 5 metres depth in the south shore of Lake Jianhu. February 2020 ; 4 specimens in ZMB, collected by Le-Jia Zhang from 0.5 metres depth in an unnamed pond on the Sheshan hill in Lijiang City in April 2022 ; 19 specimens in NJW, collected by Jiao-Wei Ning from Lake Jianhu .</p><p>Description.</p><p>Shell (Fig. 5A-G) large, broadly conical or sub globose, thin, olive to dark brown in colour; up to six whorls at adulthood, including one relatively smooth protoconch whorl, apex acute; teleoconch whorls inflated, occasionally shouldered, with one strong keel at suture, above suture with many weak spiral threads, sometimes with additional two to three stronger cords, base of shell with many thin weak spiral threads, weak spiral threads and cords with short periostracum setae; aperture ovate, slightly less than half of shell in height, lip thin and simple, umbilicus narrow, sometimes bordered by akeel.</p><p>Operculum (Fig. 5H) corneous, ovate, rather thin, yellow, with red nuclear region, exterior surface of operculum rather smooth, inner opercular region relatively small, nuclear region smooth.</p><p>Radular (Fig. 7E, F) central tooth with one broad central denticle and four small sharp cusps on either side; lateral tooth with one broad central denticle and four small sharp cusps on either side; inner marginal tooth with one broad central denticle and four small sharp cusps on either side; outer marginal tooth with 9 to 10 small sharp cusps.</p><p>Remarks.</p><p>This species differs from the other species of Dalipaludina by having rather inflated, shouldered whorls. Annandale (1924) presented a photo of the type specimen. The shell morphology and COI phylogeny of the specimens collected from an unnamed pond in Lijiang City, which is very close to the type locality, confirm that this species should be assigned to Dalipaludina . The recommended Chinese name of this species is “滇西理田螺” .</p><p>Habitat and distribution.</p><p>Shallow ponds with muddy bottoms and abundant aquatic plants in Lijiang City, Yunnan, China; Shallow to medium-depth water area with muddy bottoms in Lake Jianhu, Jianchuan County, Yunnan, China.</p></div>	https://treatment.plazi.org/id/BA528F39739C55BBA13C8BA168E48C31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Le-Jia;Du, Li-Na;von Rintelen, Thomas	Zhang, Le-Jia, Du, Li-Na, von Rintelen, Thomas (2023): A new genus of river snails, Dalipaludina (Gastropoda, Viviparidae), endemic to the Yunnan Plateau of SW China. Zoosystematics and Evolution 99 (2): 285-297, DOI: http://dx.doi.org/10.3897/zse.99.102586, URL: http://dx.doi.org/10.3897/zse.99.102586
4F71F8C5709B5EE097E5B5C7E25A4C5B.text	4F71F8C5709B5EE097E5B5C7E25A4C5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalipaludina oxytropoides (Heude 1889) Zhang & Du & von Rintelen 2023	<div><p>Dalipaludina oxytropoides (Heude, 1889) comb. nov.</p><p>Paludina oxytropoides Heude, 1889: 176 ('lacu prope Tchao-tong’ = lake near Zhaotong City,Yunnan Province, China).</p><p>Vivipara oxytropoides - Kobelt 1909: 128.</p><p>Margarya sp. - Du et al. 2012: 45.</p><p>Margarya oxytropoides - Zhang et al. 2015: 777; Wiese et al. 2022.</p><p>Material examined.</p><p>2 syntypes, IZCAS-FG-492586, IZCAS-FG-492587; 1 paratype, MCZ-Mala-167343; 91 specimens in ZMB, collected by Frank Riedel from around 1 to 6 metres depth in all over the Lake Lugu in September and October 2014 ; 5 specimens in ZMB, collected by Le-Jia Zhang from 1 metre depth in the east shore of Lake Lugu in April 2022 .</p><p>Description.</p><p>Shell (Fig. 4A-H) large, conical, thin, greenish yellow to dark brown in colour; up to six whorls at adulthood, including one relatively smooth protoconch whorl, apex acute; each teleoconch whorls with one strong keel at suture, above suture with many rather weak spiral threads, sometimes with additional two to three strong cords, base of shell with many thin weak spiral threads, weak spiral cords and threads with short periostracum setae; aperture ovate, less than half of shell in height, lip thin and simple, umbilicus narrow, sometimes bordered by a keel.</p><p>Operculum (Fig. 4I) corneous, ovate, thin to relatively thick, yellow to reddish brown, with darker coloured nuclear region, exterior surface of operculum rather smooth, inner opercular region relatively small, nuclear region smooth or with small grains.</p><p>Radular (Fig. 7C, D) central tooth with one broad central denticle and three to four small sharp cusps on either side; lateral tooth with one broad central denticle and three to four small sharp cusps on either side; inner marginal tooth with one broad central denticle and four small sharp cusps on either side; each outer marginal tooth with 10 to 11 small sharp cusps.</p><p>Remarks.</p><p>This species has the most variable shell within Dalipaludina . The form with strong cords of this species can be easily distinguished from the other Dalipaludina species. It used to be considered as a species of Margarya with a distribution in Lake Dianchi as well (Zhang et al. 2015). However, the COI-based phylogeny showed that " Margarya oxytropides " from Dianchi is not a species of Dalipaludina, but indeed a member of the Cipangopaludina / Margarya complex. The syntypes of Paludina oxytropoides Heude, 1889 from Zhaotong in Northeast Yunnan show a strong keel at the suture, two strong keels above the suture, a relatively smooth shell base with many weak spiral keels, and a narrow umbilicus with a keel around it. It is distinct from the " Margarya oxytropides " from Lake Dianchi but quite similar to the strongly keeled form of D. oxytropoides from Lake Lugu. Meanwhile, the shell morphology of " Margarya oxytropides " from Lake Dianchi matches the description and figure of Cipangopaludina dianchiensis Zhang, 1990 endemic to Lake Dianchi (Zhang 1990). Therefore, instead of D. oxytropoides, " Margarya oxytropides " from Lake Dianchi is here recognised as Margarya dianchiensis (Zhang, 1990) comb. nov. Paludina oxytropoides Heude, 1889 is reclassified as a member of Dalipaludina . The recommended Chinese name of this species is “尖龙骨理田螺” .</p><p>Habitat and distribution.</p><p>shallow to medium-depth water area with sandy bottoms in Lake Lugu, Yunnan/Sichuan, China; lakes in Zhaotong, Yunnan, China.</p></div>	https://treatment.plazi.org/id/4F71F8C5709B5EE097E5B5C7E25A4C5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Le-Jia;Du, Li-Na;von Rintelen, Thomas	Zhang, Le-Jia, Du, Li-Na, von Rintelen, Thomas (2023): A new genus of river snails, Dalipaludina (Gastropoda, Viviparidae), endemic to the Yunnan Plateau of SW China. Zoosystematics and Evolution 99 (2): 285-297, DOI: http://dx.doi.org/10.3897/zse.99.102586, URL: http://dx.doi.org/10.3897/zse.99.102586
CE87BE4DBC805A9A8681908E7B0EA7C3.text	CE87BE4DBC805A9A8681908E7B0EA7C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalipaludina pyramidella (Du, Yang & Chen 2011) Zhang & Du & von Rintelen 2023	<div><p>Dalipaludina pyramidella (Du, Yang &amp; Chen, 2011) comb. nov.</p><p>Trochotaia pyramidella Du, Yang &amp; Chen, 2011: 85-89 (Yousuo village of Er-Yuan County, Yunnan, China).</p><p>Material examined.</p><p>Holotype, KIZ-DLN20100035; 19 paratypes, KIZ-DLN20100036 to KIZ-DLN20100054; 3 specimens in ZMB, 7 specimens in NJW, collected by Jiao-Wei Ning from 1 metre depth in the ponds of Yousuo Village, Eryuan County.</p><p>Description.</p><p>Shell (Fig. 6A-D) large, conical, thin, greenish brown to brown in colour; up to six whorls at adulthood, including one relatively smooth protoconch whorl, apex acute, spire low; each teleoconch whorls with one strong keel at suture, above suture smooth, with two rings of short periostracum setae, base of shell smooth, with many rather thin weak spiral threads; aperture ovate, almost half of shell in height, lip thin and simple, umbilicus narrow, sometimes bordered by a keel.</p><p>Operculum (Fig. 6E) corneous, ovate, rather thin, yellow or orange, with red nuclear region, exterior surface of operculum rather smooth, inner opercular region relatively small, nuclear region smooth.</p><p>Radular (according to Du et al. 2011) central tooth with one broad central denticle and three to four small sharp cusps on either side; lateral tooth with one broad central denticle and four small sharp cusps on either; inner marginal tooth with one broad central denticle and two small sharp cusps on either side; outer marginal tooth with 9 small sharp cusps.</p><p>Remarks.</p><p>This species differs from the other species of Dalipaludina by having a relatively smooth shell with lower spire. Du et al. (2011) assigned this species to Trochotaia mainly based on the strongly keeled shell with lower spire. However, characters of the operculum, embryonic shells, the reduced number of cusps on the outer marginal teeth (9 in D. pyramidella vs 14-16 in Trochotaia trochoides) and the distribution in Yunnan strongly support its assignment to Dalipaludina . Species of Trochotaia have a very different operculum according to Zhang and von Rintelen (2021). This species is also quite similar to Cipangopaludina miyagii Kuroda, 1941 endemic to Kaohsiung, S Taiwan, China. It can be distinguished from C. miyagii by having a thinner shell, a shell base with many rather thin weak spiral threads, and a narrowly open umbilicus with a keel around it. The recommended Chinese name of this species is “塔形理田螺” .</p><p>Habitat and distribution.</p><p>shallow ponds with muddy bottoms and aquatic plant Ottelia acuminata (Gagnep.) in Yousuo Village, Eryuan County, Yunnan, China.</p></div>	https://treatment.plazi.org/id/CE87BE4DBC805A9A8681908E7B0EA7C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Le-Jia;Du, Li-Na;von Rintelen, Thomas	Zhang, Le-Jia, Du, Li-Na, von Rintelen, Thomas (2023): A new genus of river snails, Dalipaludina (Gastropoda, Viviparidae), endemic to the Yunnan Plateau of SW China. Zoosystematics and Evolution 99 (2): 285-297, DOI: http://dx.doi.org/10.3897/zse.99.102586, URL: http://dx.doi.org/10.3897/zse.99.102586
BC487F66D49F50AFA5982C2056D81931.text	BC487F66D49F50AFA5982C2056D81931.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalipaludina Zhang 2023	<div><p>Dalipaludina Zhang gen. nov.</p><p>Type species.</p><p>Paludina delavayana Heude, 1889.</p><p>Etymology.</p><p>“Dali” refers to the ancient Dali Kingdom (大理国) mostly situated in modern Yunnan, China; “paludina” refers the assignment to Viviparidae . The recommended Chinese name is “理田螺” .</p><p>Diagnosis.</p><p>Shell large, thin but solid; apex acute; teleoconch whorls with strong keel at suture, above suture relatively smooth or with several weak spiral threads or strong spiral cords, many thin and weak spiral threads on base; umbilicus narrow, sometimes bordered by a keel; exterior surface of operculum rather smooth, inner opercular region relatively small, nuclear region of operculum smooth, sometimes with small grains; outer marginal tooth with 9 to 11 small sharp cusps.</p><p>Comparative remarks.</p><p>Dalipaludina gen. nov. resembles Cipangopaludina (widely distributed in East Asia), Ussuripaludina Zatravkin &amp; Bogatov, 1987 (endemic to Far East), Heterogen Annandale, 1921 (endemic to Japan and Korea), and Torotaia Haas, 1939 (endemic to Philippines, Sulawesi and New Guinea). It differs from Cipangopaludina, Ussuripaludina, Heterogen and Torotaia by having one strong keel at the suture, a shell base with many thin and weak spiral threads, a narrow umbilicus sometimes bordered by a keel, and operculum characters. The molecular phylogeny supports that Dalipaludina is distinct from Cipangopaludina, Ussuripaludina, Heterogen and Torotaia . The location of the testis in male Dalipaludina specimens in the mantle cavity supports the classification of this genus in the subfamily Bellamyinae . There are four known species of Dalipaludina .</p><p>Distribution.</p><p>This genus is endemic to the lakes or ponds in northwest to northeast Yunnan, China.</p></div>	https://treatment.plazi.org/id/BC487F66D49F50AFA5982C2056D81931	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Le-Jia;Du, Li-Na;von Rintelen, Thomas	Zhang, Le-Jia, Du, Li-Na, von Rintelen, Thomas (2023): A new genus of river snails, Dalipaludina (Gastropoda, Viviparidae), endemic to the Yunnan Plateau of SW China. Zoosystematics and Evolution 99 (2): 285-297, DOI: http://dx.doi.org/10.3897/zse.99.102586, URL: http://dx.doi.org/10.3897/zse.99.102586
