taxonID	type	description	language	source
166387BCFFF5FFDD9A04FE11FD646C46.taxon	description	Redescription. Body. Small to medium-sized (3.5 – 10.0 mm), oblong ovate, convex, glabrous or elytra pubescent. Complete body usually coloured in various shades of yellow, orange or brown, elytra often with paler apical half; antennae, tibiae, tarsi, head, pronotum or part of underside black in some species. Male. Head (Fig. 1). Labrum transverse, subrectangular, with rounded anterior angles, anterior margin usually shallowly concave, dorsally with several setiferous pores. Anterior part of head subtriangular, more or less elevated, posterior tip with distinctly raised nasal keel. Frontal tubercles subtriangular, with anterior angles produced to interantennal space, apically separated from each other by nasal keel, basally by thin furrow, from frons separated by impressed line. Interocular space wide, 1.25 – 1.60 times as wide as transverse diameter of eye. Interantennal space narrow, 0.60 – 1.00 times as wide as transverse diameter of antennal socket. Eyes small to medium sized, rounded or shortly elliptical, convex. Vertex wide, convex. Mandibles with four teeth. Apical maxillar palpomere conical. Antennae (Fig. 6) with 11 antennomeres, filiform, 0.75 – 1.25 times as long as body, antennomere III 1.5 – 4 times longer than anntenomere II, apical antennomeres 6 – 8 times longer than wide. Prothorax. Pronotum subquadrate to transverse, wider than long, ratio of length to width 1.30 to 1.80, anterior margin straight or almost straight, posterior margin rounded or straight medially, lateral margins more or less rounded, anterior margin unbordered, posterior margin thinly bordered, lateral margins with wider border, sometimes chanelled. Surface glabrous, almost impunctate or finely to distinctly punctate, moderately convex, sometimes laterally with slightly elevated callosity, always with short shallow transverse or oblique impression behind anterior angles (Fig. 2). Procoxal cavities opened behind (Fig. 3), intercoxal prosternal process thin, low anteriorly, gradually increasing posteriorly. Scutellum small, triangular, with apex sharp or more or less rounded. Mesothorax slender, posteriorly divergent, in middle of posterior margin with thin sharp procession. Metathorax wide, convex (Fig. 4). Elytra. Elongate, convex, widest in posterior third, usually glabrous but in several species almost whole elytra or their posterior half densely covered with short pale semierected setae (sometimes partly abraded), punctation dense, fine, confused, without postscutellar impression. Epipleura impunctate, wide basally, in basal quarter slightly extended, narrowed in middle part, slender posteriorly, gradually disappearing at apex (Fig. 5). Elytral base distinctly wider than pronotal base, humeral calli well developed. Lateral and basal borders well developed, basal border ending near scutellum. Macropterous. Legs moderately thin (Fig. 7), apices of middle and metatibiae with spine. Tarsi thin, first tarsomeres of all legs elongate triangular, base of metatarsomere I not darkened, about as long as two following metatarsomeres combined. Claws appendiculate (Fig. 8). Abdomen. Posterior margin of last ventrite concave, sometimes straight in middle of concavity, without two incisions or with only very short indicated incisions, extreme posterior margin bent downwards. Surface in front of apical concavity transversely rounded and impressed. Structure of aedeagus variable, from relatively simple to very complicated (see definitions of species groups), with large elongate basal orifice and internally with one or two large and long sclerites. Female. Abdomen usually more convex, last ventrite entire or with wide shallow subtriangular incision. Pygidium entire or with narrow wedge-shaped incision. Spermatheca: nodulus globular or subglobular, cornu relatively narrow, distinctly inserted into nodulus, apex with distinct small appendix, spermathecal duct basally wider. Vaginal palpi, tignum and sternite VIII as in Figs 9 – 10.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF5FFDD9A04FE11FD646C46.taxon	diagnosis	Diagnosis. The genus Hesperopenna belongs to the subtribe Luperina (Galerucinae: Luperini) and is characterised by combination of the following characters: filiform antennae, anterior margin of pronotum unbordered, procoxal cavities open, apices of meso- and metatibiae with spine, metatarsomere I about as long as two following metatarsomeres combined, claws appendiculate and usually complicated structure of aedeagus. In habitus, some oblong ovate species of Calomicrus (polyphyletic in current concept) and Erganoides Jacoby, 1903 are similar mainly to smaller species of Hesperopenna. In both Calomicrus and Erganoides pronotum is regularly convex, without any oblique impression behind anterior angles, which is typical for Hesperopenna. Last abdominal ventrite of male is trilobed with relatively short but always well developed incisions in both Calomicrus and Erganoides while the incisions are either missing or only indicated in Hesperopenna. In general appearance Hesperopenna species may resemble also some Oriental genera / species of the section Monoleptites (e. g. Monolepta, Ochralea Clark, 1865, Paleosepharia Laboissière, 1936, etc.) but can be easily distinguished by shorter metatarsomere I (typically elongated in Monoleptites, cf. WILCOX 1973, WAGNER 2004).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF5FFDD9A04FE11FD646C46.taxon	distribution	Distribution (based partly on specimens of formally undescribed species studied by the author). Vietnam, Thailand, Laos, Cambodia, South China, eastern provincies of India, Indonesia (Sumatra, Java, Bali), Malaysia (Peninsular Malaysia, Sarawak). Most known species are accumulated in continental SE Asia. Previous records from Malaysia (MOHAMEDSAID & KIMOTO 1993; MOHAMEDSAID 1995, 1997, 1998, 1999, 2000, 2001; MOHAMEDSAID & HOLLOWAY 1999) and the Philippines (MEDVEDEV 1995) deal with species excluded here from Hesperopenna.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF0FFDE9BC1FB5BFD7C6F9D.taxon	description	Body length 4.2 – 5.7 mm. Body completely orange or tibiae and metaventrite black. Antennae longer than body, antennomere III about 1.5 – 3.5 times longer than anntenomere II. Pronotum 1.3 – 1.4 times as wide as long, covered with fine dense punctures. Elytra glabrous. Aedeagus simple, long, thin, with long internal sclerite (Fig. 67).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF0FFDE9BC1FB5BFD7C6F9D.taxon	discussion	Species included. Hesperopenna annulicornis (Jacoby, 1896) (comb. nov.), H. antennalis (Kimoto, 1989) (comb. nov.), H. fulvicollis (Jacoby, 1896) (comb. nov.), H. pallida (Jacoby, 1894) (comb. nov.) and at least two undescribed species.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF0FFDE9BC1FE79FCC2699F.taxon	description	Larger species, body length 6.6 – 10.0 mm. Body completely yellow, orange or brown, legs completely pale or tibiae and tarsi black, antennae usually gradually darkened from antennomere III or IV, or black with first two antennomeres pale, or, rarely, antennae completely yellow. Antennomere III about 1.5 – 2.0 times longer than anntenomere II. Pronotum less transverse, 1.40 – 1.60 times as wide as long, widest in anterior third, lustrous, with very fine, almost indistinct punctures, lateral margins moderately rounded, channeled, posterior margin rounded. Scutellum triangular with apex shortly rounded. Elytra lustrous (in species with pubescent elytra the places with setae semiopaque), glabrous or partly covered with short dense setae. Pygidium of females often incised at apex, in some species apex pointed. Aedeagus long, robust, subtubular, with basal orifice large and long, with two endophallic sclerites: one very large, with spoon-like apex and distinct ridges ventrally, always protruding aedeagus, second thin, usually hidden inside the aedeagus (Figs 11 – 18).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF0FFDE9BC1FE79FCC2699F.taxon	discussion	Species included. Hesperopenna medvedevi nom. nov. (for flava Medvedev & Dang, 1981), H. tibialis (Kimoto, 1989) (comb. nov.) (= merkli Medvedev, 2000, syn. nov.), H. persimilis (Kimoto, 1989) (comb. nov.) and 5 new species described below.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF0FFDE9BC1FC4AFC1B6E8C.taxon	description	Body length 5.1 – 7.1 mm. Body completely yellow or with tibiae, tarsi and / or head black. Antennomere III about 1.5 – 3.0 times longer than anntenomere II. Pronotum transverse, 1.6 – 1.8 times as wide as long, finely punctate. Elytra glabrous. Aedeagus dorsally with two thin lateral processes, incision between them very deep, reaching basal third of aedeagus, ventral side apically with or without hook-like process (Fig. 66).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF0FFDE9BC1FC4AFC1B6E8C.taxon	discussion	Species included. Hesperopenna vietnamica (Medvedev, 2000) (comb. nov.), H. thailandica (Kimoto, 1989) (comb. nov.) and at least four undescribed species.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF0FFDE9BC1FA74FEFC6C42.taxon	description	Body length 3.2 – 5.0 mm. Body completely orange. Antennomere III about 1.3 – 1.5 times longer than anntenomere II. Pronotum transverse, 1.6 – 2.2 times as wide as long, covered with fine dense punctures. Elytra glabrous. Aedeagus relatively short, simple (Fig. 68).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF0FFDE9BC1FA74FEFC6C42.taxon	discussion	Species included. Hesperopenna minor (Kimoto, 1989) (comb. nov.) and at least two undescribed species.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF1FFDF9B27FF37FC586B86.taxon	description	Body length 3.8 – 6.5 mm. Body completely orange or tibiae and tarsi or head black. Antennomere III about 1.3 – 1.5 times longer than anntenomere II. Pronotum 1.6 – 2.0 times as wide as long, covered with almost indistinct to fine dense punctures Aedeagus ventrally with long projection from basal part directed anteriorly (Fig. 69).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF1FFDF9B27FF37FC586B86.taxon	discussion	Species included. Hesperopenna bicolor (Kimoto, 1989) (comb. nov.), H. shinsakui nom. nov. (for Calomicrus bicolor Kimoto, 1989) and at least two undescribed species.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF1FFDF9B27FE40FEFC68F6.taxon	description	Body length 3.7 – 7.0 mm. Antennomere III about two times longer than anntenomere II. Body completely orange, tibiae sometimes black. Pronotum transverse, 1.45 – 1.55 times as wide as long, distinctly covered with fine dense punctures, often with two or four small slightly elevated areas on the disc. Elytra glabrous. Aedeagus with relatively short proximal part, apex often bent down (Fig. 70).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF1FFDF9B27FE40FEFC68F6.taxon	discussion	Species included. Hesperopenna granulicollis (Kimoto, 1989) (comb. nov.), H. fulva (Kimoto, 1977) (comb. nov.), H. flava (Jacoby, 1892) (comb. nov.) (= kimotoi Warchałowski, 1991, syn. nov.), H. romantsovi nom. nov. (for Levnma thailandica Medvedev & Romantsov, 2013) (comb. nov.), H. malayana (Medvedev & Romantsov, 2013) (comb. nov.) and at least six undescribed species.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFFFFD29AC4FE17FE966BC6.taxon	description	(Figs 11, 19, 27, 34, 41, 48, 56 – 57)	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFFFFD29AC4FE17FE966BC6.taxon	materials_examined	Type locality. Vietnam, Baktkhay, 50 km N of Thai Nguyen. Type material. HOLOTYPE: ♂, ‘ Вьетнам горы / Тхaй- Нгуен 50 км [p] / 21. [h] y [p] I. [h] 196 [p] 3 [h] Кабаков [w, p] // Holotypus [p] / Hesperopenna flava / L. Medv. et D-Dap [red label, h] ’ (LMCM). Additional material examined (8 spec.). VIETNAM: NINH BINH PROV.: Cuc Phong N. P., 20 ° 21 ′ 10 ″ N 105 ° 35 ′ 00 ″ E, 440 m, 24. - 28. iv. 2012, 4 ♂♂ 3 ♀♀, E. Jendek leg. (JBCB); Cuc Phuong, 3. – 10. v. 1966, 1 ♀, G. Topál leg. (ZMHB).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFFFFD29AC4FE17FE966BC6.taxon	description	Description. Body length: ♂♂ 7.6 – 8.5 mm (holotype 7.6 mm), ♀♀ 8.0 – 9.0 mm. Male (Fig. 48, holotype Fig. 56). Body completely pale brown, antennae pale brown or darkened from antennomere IV, apical half of elytra slightly paler than rest of body. Interocular space 1.5 times as wide as transverse diameter of eye. Antennae 0.85 times as long as body, length ratio of antennomeres I to XI equals 15 - 4 - 7 - 13 - 14 - 14 - 16 - 15 - 15 - 14 - 15. Pronotum 1.50 – 1.55 times as broad as long. Elytra 1.7 times as long as wide (measured at humeral calli), 0.70 times as long as body. Apical half of elytra densely covered with short pale setae. Length ratio of metatarsomeres I – IV equals 12 - 6 - 6 - 9. Last ventrite posteriorly without two incisions (Fig. 19). Aedeagus: dorsal side with long thin incision separating slightly wider and longer left part from narrower right part; ventral side apically with wide subtriangular process, in middle with distinct sharp keel (Fig. 11). Female. Pronotum 1.48 – 1.58 times as broad as long. Apex of last ventrite with wide triangular incision (Fig. 27), pygidium with U-shaped incision and slightly concave posterior margins near incision (Fig. 34). Spermatheca: nodulus globular, cornu regularly rounded, apical appendage bent up (Fig. 41).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFFFFD29AC4FE17FE966BC6.taxon	diagnosis	Differential diagnosis. Due to the apical half of elytra covered with short pale setae and completely brown legs, H. medvedevi nom. nov. is similar to H. sipekorum sp. nov. and H. arnoldi sp. nov. In H. arnoldi sp. nov. at least the apical 2 / 3 of elytra are covered with short pale setae, while only the apical half is covered with setae in H. sipekorum sp. nov. and H. medvedevi. Aedeagi of all three species share a thin long incision in the middle of dorsal side. Ventral side of aedeagus in H. medvedevi is more or less symmetrical, apex formed by subtriangular process with a rounded tip, lateral incisions short and of the same depth (Fig. 11). On the other hand, ventral sides of aedeagus in H. sipekorum sp. nov. and H. arnoldi sp. nov. have assymetrical apex with a deep U-shaped (H. arnoldi sp. nov.) or V-shaped (H. sipekorum sp. nov.) incision on the right and very short incision on the left (Figs 14, 17). Females of these three species can be distinguished by apex of pygidium with an U-shaped incision which is large and deep in H. sipekorum sp. nov., very small in H. arnoldi sp. nov., and of medium size in H. medvedevi (Figs 34, 37, 39).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFFFFD29AC4FE17FE966BC6.taxon	distribution	Distribution. North Vietnam (MEDVEDEV & DANG 1981, present paper).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFFFFD29AC4FE17FE966BC6.taxon	discussion	Comments. Hesperopenna flava Medvedev & Dang, 1981 is a junior secondary homonym of H. flava (Jacoby, 1892) (comb. nov.), thus a replacement name H. medvedevi nom. nov. is proposed.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFCFFD39A0BFE17FDB76BA1.taxon	description	(Figs 12, 20, 28, 35, 42, 49, 58 – 59)	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFCFFD39A0BFE17FDB76BA1.taxon	materials_examined	Type locality. ‘ Laos: Sedone Prov., Pakse’. Type material examined. HOLOTYPE: ♂, ‘ LAOS: / Sedone Prov. / Pakse / 31. V. 196 [p] 5 [w, h] // Native Collector / BISHOP MUS. [w, p] // HOLOTYPE [red label, p] // Calomicrus / persimilis / n. sp. [w, h] // PHOTO [red label, p] // 14964 [pink label, h] ’ (BPBM). PARATYPES: 1 ♂ 1 ♀, ‘ LAOS: / Sedone Prov. / Pakse / 31. V. 196 [p] 5 [w, h] // Native Collector / BISHOP MUS. [w, p] // PARATOPOTYPE [blue label, p] // Calomicrus / persimilis / n. sp. [w, h] ’ (BPBM); 2 ♂♂, ‘ LAOS: / Ile de Khong / 17. IV. 1965 [w, p] // J. L. Gressitt / Collector / BISHOP MUSEUM [w, p] // PARATYPE [blue label, p] // Calomicrus / persimilis / n. sp. [w, h] ’ (BPBM). Additional material examined (11 spec.). THAILAND: LOEI PROV.: Phu Kradung N. P., 16 ° 52´N 101 ° 49´E, 1000 m, 16. - 17. v. 1999, 1 ♀, M. Říha leg. (JBCB). PHETCHABUN PROV.: Lom Sak – Dan Sai, 17. – 19. v. 1993, 6 ♂♂ 3 ♀♀, Pacholátko & Dembický leg. (JBCB, JVCJ). LAOS: VIENTIANE PROV.: Vientiane, 1 ♂, Vitalis leg. (NMPC).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFCFFD39A0BFE17FDB76BA1.taxon	description	Description. Body length: ♂♂ 6.6 – 7.0 mm, ♀♀ 7.2 – 7.5 mm. Male (Fig. 49, paratype Fig. 58). Body completely orange, only apices of mandibles black. Interocular space 1.5 times as wide as transverse diameter of eye. Antennae 0.80 times as long as body, length ratio of antennomeres I to XI equals 11 - 4 - 5 - 13 - 12 - 12 - 12 - 13 - 12 - 12 - 13. Pronotum 1.42 – 1.54 times as broad as long. Elytra glabrous, 1.75 times as long as wide (measured at humeral calli), 0.70 times as long as body. Length ratio of metatarsomeres I – IV equals 9 - 4 - 4 - 7. Last ventrite with two very small incisions posteriorly, depression on the disc anteriorly surrounded with U-shaped ridge (Fig. 20). Aedeagus: dorsal side with deep asymmetrical incision on left side, on right side with short rounded process and wide shallow subtriangular incision; ventral side gradually tapering to triangular apex, ventral surface flat (Fig. 12). Female. Apex of last ventrite widely rounded (Fig. 28), pygidium with obtusangulate apex (Fig. 35). Spermatheca: nodulus subglobular, cornu regularly rounded with very long apical appendix (Fig. 42).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFCFFD39A0BFE17FDB76BA1.taxon	diagnosis	Differential diagnosis. Due to glabrous elytra and uniformly brown legs Hesperopenna persimilis is similar to H. helferi sp. nov. and H. bonifaci sp. nov. but differs from both species in the subapical impression on the last ventrite in male, anteriorly surrounded with an U-shaped ridge (without ridge in H. helferi sp. nov. and H. bonifaci sp. nov.) (Figs 20, 23, 24) and simple triangular apex of the ventral side of aedeagus (ventral side of aedeagus asymmetrical with a deep U-shaped incision in H. helferi sp. nov. and H. bonifaci sp. nov. – incision on the right side in H. helferi sp. nov., on the left side in H. bonifaci sp. nov.) (Figs 12, 15, 16). Females of H. persimilis and H. helferi sp. nov. can be separated by the shape of pygidium: apex obtusangulate in H. persimilis while with a small U-shaped incision in H. helferi sp. nov. (Figs 35, 38).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFCFFD39A0BFE17FDB76BA1.taxon	distribution	Distribution. Laos (KIMOTO 1989, present paper), Vietnam (KIMOTO 1989), Thailand (MOHAMEDSAID & CONSTANT 2007, present paper).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFDFFD69A1EFE37FEE168E6.taxon	description	(Figs 13, 21, 29, 36, 43, 50, 60 – 61)	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFDFFD69A1EFE37FEE168E6.taxon	materials_examined	Type localities. Liroetiella tibialis: ‘ Laos: Khammouane Prov., Phon Tiou’. Martinella merkli: ‘ Laos, Bolikhamsay Prov., Phou Khao Khouay NBCA, Tad Leuk Waterfall’. Type material examined. Liroetiella tibialis: PARATYPES: 1 ♂, ‘ LAOS: / Khammouane Prov. / Phon Tiou / 17. V. 1965 [w, p] // Native Collector / Collector [w, p] // PARATOPOTYPE [blue label, p] // Liroetiella / tibialis / n. sp. [w, h] // PHOTO [red label, p] ’ (KMNH); 1 ♀, ‘ Laos / Umg. Vientiane / III. - VI. 1963 [w, p] // PARATYPE [blue label, p] // Liroetiella / tibialis / n. sp. [w, p] // Zool. Staatsslg. / München [pale blue label, p] ’ (ZSM). Martinella merkli: PARATYPES: 1 ♂, ‘ LAOS, Bolikhamsay Prov., / Phou Khao Kouay NBCA, / Tad Leuk Waterfall, 280 m, [w, p] // at light, No. 46, / 11 - 12. IV. 1998, / leg. O. Merkl & G. Csorba [w, p] // PARATYPUS [p] / Martinella [h] / merkli m. [h] / L. Medvedev det. 19 [p] 99 [red label, h] ’ (LMCM); 1 ♀, ‘ LAOS, Bolikhamsay Prov., / Phou Khao Kouay NBCA, / Tad Leuk Waterfall, 280 m, [w, p] // at light, No. 46, / 11 - 12. IV. 1998, / leg. O. Merkl & G. Csorba [w, p] // PARATYPUS [p] / Martinella [h] / merkli m. [h] / L. Medvedev det. 19 [p] 99 [red label, h] // MUSEO GENOVA / Ex. coll. Medvedev / Acquisto XI. 2003 [w, p] ’ (MCSN). Additional material examined (16 spec.). LAOS: VIENTIANE PROV.: Lao Pako env., 55 km NE of Vientiane, 200 m, 1. - 4. v. 2004, 6 ♂♂ 2 ♀♀, J. Bezděk leg. (JBCB); Vientiane env., iii. – vi. 1963, 1 ♀, without additional data (ZSM); KHAMMOUAN PROV.: 17 ° 43 ′ N 105 ° 09 ′ E, 500 – 600 m, 22. v. – 8. vi. 2001, 1 ♀, E. Jendek & O. Šauša leg. (JBCB). VIETNAM: DONG NAI PROV.: Nam Cat Tien N. P., 1. – 15. v. 1994, 2 ♀♀, P. Pacholátko & L. Dembický leg. (NMW). MALAYSIA: Federal Malay States, 1909, 1 ♂, C. J. Brooks leg. (BMNH). PERAK: 40 km SE of Ipoh, Banjaran Titi Wangsa, Ringlet, 900 m, 25. iii – 3. iv. 2002, 1 ♂ 2 unsexed specimens, P. Čechovský leg. (RBCN, UACG).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFDFFD69A1EFE37FEE168E6.taxon	description	Description. Body length: ♂♂ 7.9 – 9.8 mm, ♀♀ 7.5 – 9.3 mm. Male (Fig. 50, paratype of Martinella merkli Fig. 60). Body yellowish orange, apices of mandibles black, antennae completely yellow, epipleura either completely yellowish orange or extreme lateral margin of epipleura in basal third brownish. Femora yellowish, tibiae and tarsi black, last tarsomere and claws dark brown. Interocular space 1.5 times as wide as transverse diameter of eye. Antennae 0.8 times as long as body, length ratio of antennomeres I to XI equals 17 - 5 - 7 - 16 - 18 - 16 - 17 - 16 - 15 - 14 - 15. Pronotum 1.52 – 1.65 times as broad as long. Elytra lustrous, glabrous, 1.75 times as long as wide (measured at humeral calli), 0.7 times as long as body. Length ratio of metatarsomeres I – IV equals 12 - 6 - 5 - 10. Last ventrite posteriorly without two incisions (Fig. 21). Aedeagus: dorsal side apically with short triangular asymmetrical incision; ventral side with very long thin incision placed right along longitudinal axis (Fig. 13). Female. Last ventrite with wide concavity at apex, lateral sides regularly rounded (Fig. 29), pygidium with sharp pointed tip (Fig. 36). Spermatheca: nodulus subglobular, cornu relatively thin, regularly rounded, slightly wider than nodulus (Fig. 43).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFDFFD69A1EFE37FEE168E6.taxon	diagnosis	Differential diagnosis. Having bicolorous legs and glabrous elytra, Hesperopenna tibialis is similar to H. zofka sp. nov. from Java. Both species also share similar structure of aedeagus with a very long basal orifice longer than the proximal part of aedeagus, which might indicate a close relationship. Hesperopenna tibialis can be distinguished by completely yellow antennae (black with first two antennomeres yellow in H. zofka sp. nov.) and epipleura completely yellowish orange or at most with brownish extreme lateral margin (always with black extreme lateral margin in basal half in H. zofka sp. nov.). Males of both species can be distinguished by long thin incision on the ventral side of aedeagus. While it is located on the right side in H. tibialis, it is placed medially in H. zofka sp. nov. (Figs 13, 18). Females of both species can be separated by the shape of pygidium: apex is distinctly pointed in H. tibialis while simply triangular in H. zofka sp. nov. (Figs 36, 40).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFDFFD69A1EFE37FEE168E6.taxon	distribution	Distribution. Laos (KIMOTO 1989, MEDVEDEV 2000, present paper), Thailand (KIMOTO 1989). Newly recorded from Vietnam and Peninsular Malaysia.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFFDFFD69A1EFE37FEE168E6.taxon	discussion	Comments. One specimen of Calomicrus flavus from Laos published by KIMOTO (1989) proved to be a female of H. tibialis. Record of Liroetiella tibialis from Thailand published by MOHAMEDSAID & CONSTANT (2007) refers to Hesperopenna vietnamica (relevant specimens examined in IRSN).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF8FFC89A80FCF0FD406881.taxon	description	(Figs 14, 22, 30, 37, 44, 51)	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF8FFC89A80FCF0FD406881.taxon	materials_examined	Type locality. China, Yunnan, Jinghong env., 21 ° 57.617 ′ N 100 ° 35.985 ′ E. Type material examined. HOLOTYPE: ♂, ‘ China, Yunnan, env. / Jinghong 12 / 19. vi. 2006 / N 21 ° 57,617´/ E 100 ° 35,985´1350 m / R. Novak leg. [w, p] ’ (NMPC). PARATYPES: 3 ♀♀, ‘ 12 - 21.6.2006 Čína / Jinghong 1550 mnm / N 21 ° 57, E 100 ° 35 / lgt. Kremitovský [w, p] ’ (JBCB); 1 ♂, ‘ NW Thailand, 1. - 7. V. / 1992, MAE HONG SON / Ban Si Lang / 1000 m, S. Bily leg. [w, p] ’ (NHMB); 3 ♂♂ 1 ♀, ‘ LAOS c. Bolikhamsai prov. / BAN NAPE-KAEW NUA-pass / 18 iv – 1 v 1998, 600 m. / N 18 ° 22,3´E 105 ° 09,1´/ lgt Sausa [w, p] ’ (RBCN, UACG); 1 ♀, ‘ LAOS north, 13 - 24. V. 1997 / 15 km NW Louang Namtha / N 21 ° 07.5, E 101 ° 21.0 / alt. 750 ± 100 m / E. Jendek & O. Šauša leg. [w, p] ’ (JBCB); 1 ♀, ‘ LAOS, / Luang Namtha / 1997 [w, p] ’ (FKCC). Material not included in the type series. 2 ♂♂, ‘ LAOS c. Bolikhamsai prov. / BAN NAPE-KAEW NUA-pass / 18 iv – 1 v 1998, 600 m. / N 18 ° 22,3´E 105 ° 09,1´/ lgt Sausa [w, p] ’ (RBCN, UACG). See Comments below.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF8FFC89A80FCF0FD406881.taxon	description	Description. Body length: ♂♂ 7.7 – 9.2 mm (holotype 8.4 mm), ♀♀ 7.7 – 10.0 mm. Male (holotype, Fig. 51). Body completely brown, apices of mandibles black, antennae gradually darkened from antennomere III, apical half of elytra slightly paler than rest of elytra. Interocular space 1.87 times as wide as transverse diameter of eye. Antennae 0.80 times as long as body, length ratio of antennomeres I to XI equals 14 - 5 - 7 - 15 - 16 - 16 - 16 - 16 - 16 - 14 - 15. Pronotum 1.60 times as broad as long. Elytra 1.77 times as long as wide (measured at humeral calli), 0.80 times as long as body. At least apical two thirds of elytra densely covered with short pale setae. Length ratio of metatarsomeres I – IV equals 12 - 5 - 5 - 9. Last ventrite posteriorly with two small incisions (Fig. 22). Aedeagus: dorsal side with long thin incision separating slightly wider and longer left part with wider apex from narrower and shorter right part with subtriangular apex; ventral side apically with tape-like slightly right-turn process in middle, right side with deep U-shaped incision, left side with short triangular incision (Fig. 14). Female. Apex of last ventrite with wide shallow concavity (Fig. 30), pygidium with small U-shaped incision (Fig. 37). Spermatheca: nodulus subglobular, cornu regularly rounded, wider than nodulus, apical appendage distinct, triangular (Fig. 44). Variability. Width / length ratio of pronotum in males varies between 1.51 – 1.60, in females 1.46 – 1.57.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF8FFC89A80FCF0FD406881.taxon	diagnosis	Differential diagnosis. Having most of elytral surface densely covered with short pale setae and completely brown legs, Hesperopenna arnoldi sp. nov. is similar to H. medvedevi nom. nov. and H. sipekorum sp. nov. Elytral setation covers at least the apical 2 / 3 of elytra while only the apical half in H. medvedevi and H. sipekorum sp. nov. All three species can be separated by the structure of the ventral side of aedeagus. Hesperopenna medvedevi nom. nov. has aedeagus more or less symmetrical ventrally, apex formed by a subtriangular process with a rounded tip, lateral incisions short and of the same depth, while H. arnoldi sp. nov. and H. sipekorum sp. nov. have aedeagus asymmetrical ventrally with a deep incision on the right and a shallow incision on the left. Proximal part of aedeagus of H. sipekorum sp. nov. is comparatively shorter and more robust than in H. arnoldi sp. nov. Moreover, ventral side of aedeagus of H. sipekorum sp. nov. forms a thin keel bent downwards (Figs 11, 14, 17). Females of these three species can be distinguished by apex of pygidium with an U-shaped incision which is large and deep in H. sipekorum sp. nov., very small in H. arnoldi sp. nov. and of medium size in H. medvedevi (Figs 34, 37, 39).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF8FFC89A80FCF0FD406881.taxon	etymology	Etymology. Dedicated to Ulf Arnold (Berlin, Germany), specialist in Chrysomelidae, who kindly allowed the study of specimens from his collection.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF8FFC89A80FCF0FD406881.taxon	distribution	Distribution. China (Yunnan), Laos, Thailand.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFF8FFC89A80FCF0FD406881.taxon	discussion	Comments. Two males from Laos are left out of the type series because the deep U-shaped incision in the apical part of the ventral side of aedeagus is placed oppositely (on the left side, while in all other males it is placed on the right side).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE6FFC89A86FD57FDCF6C39.taxon	description	(Figs 15, 23, 52)	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE6FFC89A86FD57FDCF6C39.taxon	materials_examined	Type locality. Myanmar, Kachin State, 25 km E Putao, Nansabon vill. Type material examined. HOLOTYPE: ♂, ‘ N Myanmar, 800 m, / 25 km E Putao, / Nansabon vill., 6 - 9. v. / 1998, S. Murzin leg. [w, p] ’ (NMPC). PARATYPE: 1 ♂, same data as in holotype (JBCB).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE6FFC89A86FD57FDCF6C39.taxon	description	Description. Body length: ♂♂ 8.7 – 9.5 mm (holotype 9.5 mm). Male (holotype, Fig. 52). Body completely orange brown, apical half of elytra slightly paler, apices of mandibles black. Interocular space 1.6 times as wide as transverse diameter of eye. Antennae 0.85 times as long as body, length ratio of antennomeres I to XI equals 19 - 5 - 8 - 18 - 18 - 18 - 18 - 17 - 17 - 15 - 17. Pronotum 1.55 times as broad as long. Elytra glabrous (only several pale setae on apical slopes), 1.85 times as long as wide (measured at humeral calli), 0.75 times as long as body. Length ratio of metatarsomeres I – IV equals 14 - 7 - 5 - 12. Last ventrite posteriorly with two small incisions (Fig. 23). Aedeagus: dorsal side medially with very long thin incision, apically pointed to two touching processes; ventral side apically asymmetrical, on left with U-shaped incision, on right with short process with rounded apex (Fig. 15). Female unknown. Variability. In paratype, pronotum 1.45 times as broad as long.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE6FFC89A86FD57FDCF6C39.taxon	diagnosis	Differential diagnosis. Having glabrous elytra and brown legs, Hesperopenna bonifaci sp. nov. is similar to H. helferi sp. nov. Both species can be distinguished by the structure of aedeagus which in H. bonifaci sp. nov. forms two thin touching processes dorsally and is deeply incised on the left side ventrally, while in H. helferi sp. nov. forms two wider processes dorsally and is deeply incised on the right side ventrally, moreover with a high median keel (Figs 15, 16).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE6FFC89A86FD57FDCF6C39.taxon	etymology	Etymology. Dedicated to Bonifác the black tom cat of Jiří Hájek and Květa Kalíková.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE6FFC89A86FD57FDCF6C39.taxon	distribution	Distribution. Myanmar (Kachin State).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE5FFCF9AE5FF30FDEF6846.taxon	description	(Figs 16, 24, 31, 38, 45, 53)	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE5FFCF9AE5FF30FDEF6846.taxon	materials_examined	Type locality. Myanmar, Tenasserim [= Tanintharyi Region]. Type material examined. HOLOTYPE: ♂, ‘ MUS. PRAGENSE / TENASSERIM / COLL. HELFER [w, p] ’ (NMPC). PARATYPES: 2 ♂♂ 1 ♀ and 1 spec. unsexed (abdomen missing), same data as in holotype (NMPC, 1 ♂ in JBCB).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE5FFCF9AE5FF30FDEF6846.taxon	description	Description. Body length: ♂♂ 7.9 – 8.5 mm (holotype 8.5 mm), ♀ 8.6 mm. Male (paratype, Fig. 53). Body completely brown, apical half of elytra somewhat paler, apices of mandibles black, antennae gradually darkened from antennomere III. Interocular space twice as wide as transverse diameter of eye. Antennae 0.75 times as long as body, length ratio of antennomeres I to XI equals 16 - 5 - 7 - 15 - 15 - 15 - 16 - 15 - 16 - 14 - 17. Pronotum 1.55 times as broad as long. Elytra glabrous, 1.65 times as long as wide (measured at humeral calli), 0.72 times as long as body. Length ratio of metatarsomeres 1 – 4 equals 12 - 7 - 5 - 11. Last ventrite posteriorly without two small incisions (Fig. 24). Aedeagus: dorsal side with long thin incision; apex ventrally with high thin keel in the middle and deep U-shaped incision on right side (Fig. 16). Female. Apex of last ventrite with three very small protrusions (median and two lateral) (Fig. 31), pygidium with small U-shaped incision at apex (Fig. 38). Spermatheca: nodulus relatively narrow, subtubular, slightly extended apically, cornu regularly rounded, slightly wider than nodulus, apical appendage quadrangular (Fig. 45).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE5FFCF9AE5FF30FDEF6846.taxon	diagnosis	Differential diagnosis. Hesperopenna helferi sp. nov. is similar to H. bonifaci sp. nov. Both species share glabrous elytra and uniformly brown legs and can be distinguished by the structure of aedeagus which in H. bonifaci sp. nov. forms two thin touching processes dorsally and is deeply incised on the left side ventrally, while in H. helferi sp. nov. forms two wider processes dorsally and is deeply incised on the right side ventrally, moreover with high median keel (Figs 15, 16).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE5FFCF9AE5FF30FDEF6846.taxon	etymology	Etymology. Dedicated to a Czech physician and explorer Jan Vilém Helfer (1810 – 1840) who collected the type series and whose collection is now deposited in NMPC.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE5FFCF9AE5FF30FDEF6846.taxon	distribution	Distribution. Myanmar (Tanintharyi Region).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE1FFC09AC8FD97FE7268A1.taxon	description	(Figs 1 – 10, 17, 25, 32, 39, 46, 54)	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE1FFC09AC8FD97FE7268A1.taxon	materials_examined	Type locality. India, Meghalaya, SW of Sohra, 29 ° 14´N 91 ° 40´E. Type material examined. HOLOTYPE: ♂, ‘ NE INDIA, Meghalaya, / SW of Sohra, 800 - 1000 m, / 29 ° 14´N 91 ° 40´E, / v. 2005, C. L. Peša leg. [w, p] ’ (NMPC). PARATYPES: 4 ♂♂ 8 ♀♀, same data as holotype (2 ♂♂ 2 ♀♀ in NMPC, 2 ♂♂ 6 ♀♀ in JBCB); 1 ♂ 1 ♀, ‘ NE INDIA, MEGHALAYA; / SW of CHERRAPUNJEE; / 25 ° 13´- 15´N; 91 ° 40´E; / 500 - 900 m; 11. - 12. v. 2004; / R. Businský leg. [w, p] ’ (JBCB); 2 ♂♂, ‘ INDIA, Meghalaya State, / E Khasi Hills, 11 km SW / Cherrapunjee, Laitkynsew, [w, p] // 810 m, 21 - 24. iv. 2008 / 25 ° 13´N, 91 ° 39´E, Fikáček / Podskalská & Šípek leg. [w, p] ’ (JBCB); 7 ♂♂ 10 ♀♀, ‘ NE INDIA, MEGHALAYA / SW of CHERRAPUNJEE / 25 ° 13´- 14´N 91 ° 40´E, 900 m / L. Dembický leg., 1. - 24. v. 2005 [w, p] // BMNH {E} / 2006 - 48 / L. Dembicky [w, p] ’ (BMNH); 2 ♂♂ 3 ♀♀, ‘ NE INDIA, MEGHALAYA, / SW of CHERRAPUNJEE / 25 ° 13´- 14´N 91 ° 40´E, / 5. - 24. v. 2005, 900 m / P. Pacholátko leg. [w, p] // BMNH {E} / 2006 - 48 / L. Dembicky [w, p] ’ (BMNH); 1 ♂ 1 ♀, ‘ NE INDIA, MEGHALAYA, / 8 km N of SHILLONG; / 25 ° 38´N 91 ° 54; ~ 1200 m / L. Dembický leg.; 7. - 9. v. 2004 [w, p] ’ (NHMB).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE1FFC09AC8FD97FE7268A1.taxon	description	Description. Body length: ♂♂ 7.1 – 8.9 mm (holotype 8.3 mm), ♀♀ 6.6 – 9.4 mm. Male (holotype, Fig. 54). Body orange brown, apices of mandibles black, antennae gradually darkened from antennomere III, apical half of elytra slightly paler than rest of elytra. Interocular space 1.75 times as wide as transverse diameter of eye. Antennae 0.85 times as long as body, length ratio of antennomeres I to XI equals 14 - 5 - 7 - 15 - 15 - 14 - 16 - 15 - 15 - 14 - 17. Pronotum 1.55 times as broad as long. Elytra 1.78 times as long as wide (measured at humeral calli), 0.70 times as long as body. Apical third of elytra densely covered with short pale setae. Length ratio of metatarsomeres I – IV equals 13 - 6 - 7 - 10. Last ventrite posteriorly without two small incisions (Fig. 25). Aedeagus: dorsal side with long thin incision; apex ventrally in the middle with high thin keel bent down and wide subtriangular incision on right side (Fig. 17). Female. Apex of last ventrite almost straight (Fig. 32), apex of pygidium with large Ushaped incision (Fig. 39). Spermatheca: nodulus subtubular, slightly wider apically, cornu regularly rounded, as wide as nodulus, apical appendage large, bent up (Fig. 46). Variability. Ratio width / length of pronotum in males varies between 1.44 – 1.56, in females 1.50 – 1.54.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE1FFC09AC8FD97FE7268A1.taxon	diagnosis	Differential diagnosis. Having dense elytral setation and completely brown legs H. sipekorum sp. nov. is similar to H. arnoldi sp. nov. and H. medvedevi nom. nov. While the setation covers at least the apical 2 / 3 of elytra in H. arnoldi sp. nov. it covers about the apical third to half in H. sipekorum sp. nov. and H. medvedevi. All these species can be also separated by the structure of the ventral side of aedeagus. Hesperopenna medvedevi has aedeagus more or less symmetrical ventrally, apex formed by a subtriangular process with a rounded tip, lateral incisions short and of the same depth, while H. arnoldi sp. nov. and H. sipekorum sp. nov. have aedeagus asymmetrical ventrally with a deep incision on the right and a shallow incision on the left. Proximal part of aedeagus of H. sipekorum sp. nov. is comparatively shorter and more robust than in H. arnoldi sp. nov. Moreover, ventral side of aedeagus of H. sipekorum sp. nov. forms a thin keel bent downwards (Figs 11, 14, 17). Females of these three species can be distinguished by apex of pygidium with an U-shaped incision which is large and deep in H. sipekorum sp. nov., very small in H. arnoldi sp. nov. and of medium size in H. medvedevi (Figs 34, 37, 39).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE1FFC09AC8FD97FE7268A1.taxon	etymology	Etymology. Dedicated to Hanka Podskalská and Petr Šípek (now married couple the Šípeks) who collected two paratypes.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFE1FFC09AC8FD97FE7268A1.taxon	distribution	Distribution. India (Meghalaya).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEEFFC19AB6FD00FE756886.taxon	description	(Figs 18, 26, 33, 40, 47, 55)	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEEFFC19AB6FD00FE756886.taxon	materials_examined	Type locality. Indonesia, Java, Bogor env. Type material examined. HOLOTYPE: ♂, ‘ West Java II. 1991 / BOGOR env. / R. Čermák lgt. [w, p] ’ (NMPC). PARATYPES: 2 ♂♂ 7 ♀♀, same data as in holotype (2 ♀♀ NMPC, 1 ♂ 3 ♀♀ in JBCP, 1 ♂ 2 ♀♀ in JSCP); 1 ♂ 1 ♀, ‘ ID-JAVA cent. Kudus env. / Colo- 1 km N of / Gunung Muria 1050 - 1200 m / 21. - 23.1.1998 / R. Červenka lgt. [w, p] ’ (JBCB); 1 ♀, ‘ Indonesia, Bali occ. / 6 km E of Gilimanuk / 31.1.1998, R. Červenka lgt. / pitfall baited traps [w, p] ’ (JBCB); 1 ♀, ‘ L. G. E. Kalshoven / Java [p] 700 [h] M. [p] / Djember [h] / 18. II [h] 19 [p] 19 [h] No [p] 842 [w, h] // 842 [w, h] // Le Moult vend. / via Reinbek / Eing. 1 – 1957 [w, p] ’ (ZMUH); 1 ♂, ‘ L. G. E. Kalshoven / Java …… M. [p] / Djember [h] / XI [h] 19 [p] 19 [h] No [p] 842 [w, h] // Le Moult vend. / via Reinbek / Eing. 1 – 1957 [w, p] ’ (ZMUH); 1 spec. unsexed, ‘ Java orient. / Montes Tengger / 4000´1890 / H. Fruhstorfer. [blue-gray label, p] ’ (RMNH).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEEFFC19AB6FD00FE756886.taxon	description	Description. Body length: ♂♂ 6.8 – 8.9 mm (holotype 6.9 mm), ♀♀ 7.6 – 9.4 mm. Male (holotype, Fig. 55). Body yellowish orange, apices of mandibles black, antennomeres I and II yellowish, rest of antennomeres black, extreme lateral margin of epipleura in basal half black. Femora yellowish, tibiae and tarsi black, last two tarsomeres and claws dark brown. Last ventrite posteriorly with two small incisions (Fig. 26). Interocular space 1.5 times as wide as transverse diameter of eye. Antennae 0.9 times as long as body, length ratio of antennomeres I to XI equals 15 - 5 - 7 - 14 - 14 - 15 - 15 - 15 - 15 - 13 - 15. Pronotum 1.40 times as broad as long. Elytra glabrous, 1.75 times as long as wide (measured at humeral calli), 0.75 times as long as body. Length ratio of metatarsomeres I – IV equals 12 - 6 - 4 - 9. Aedeagus: dorsal side apically with symmetrical triangular incision; ventral side with very long thin incision medially (Fig. 18). Female. Last ventrite with wide concavity at apex, lateral sides slightly concave (Fig. 33), pygidium with simple sharp triangular tip (Fig. 40). Spermatheca: nodulus large, globular, cornu regularly rounded, slightly narrower than nodulus (Fig. 47). Variability. Ratio width / length of pronotum varies between 1.40 – 1.55.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEEFFC19AB6FD00FE756886.taxon	diagnosis	Differential diagnosis. Hesperopenna zofka sp. nov. is very similar to H. tibialis. Both species share a similar structure of aedeagus with a very long basal orifice longer than the proximal part of aedeagus, bicolorous legs and glabrous elytra. Hesperopenna zofka sp. nov. differs from H. tibialis in black antennae with first two antennomeres yellow (completely yellow antennae in H. tibialis) and in epipleura always with black extreme lateral margin in the basal half (epipleura completely yellow or at most with brownish extreme lateral margin in H. tibialis). Males can be distinguished by the long thin incision on the ventral side of aedeagus which is placed medially in H. zofka sp. nov. while placed on the right side in H. tibialis (Figs 13, 18). Females of both species can be separated by the shape of pygidium: apex simply triangular in H. zofka sp. nov. while distinctly pointed in H. tibialis (Figs 36, 40).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEEFFC19AB6FD00FE756886.taxon	etymology	Etymology. Dedicated to Žofka the small black cat of Jiří Hájek and Květa Kalíková. Noun in apposition.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEEFFC19AB6FD00FE756886.taxon	distribution	Distribution. Indonesia (Java, Bali).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEAFFC49A5BFF31FDC3685C.taxon	materials_examined	Type locality. ‘ Malaysia, Sabah, Lembah Danum’. Type material. HOLOTYPE: ♂, ‘ SABAH: Lembah Danum / 27 - 31 Ogm 91 / Salleh, Zaidi, Mail, Lan [w, p] // HOLOTYPE [p] / Liroetiella / antennata n. sp. [h] / des. Mohamedsaid 199 [p] 2 [h] / & Kimoto [h] [white label with red margins] ’ (UKM). Additional material examined (1 spec.). MALAYSIA: Sabah, Lembah Danum, 5. - 8. xii. 1992, 1 ♂, Ismail, Yusof & Razali leg. (MSNV).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEAFFC49A5BFF31FDC3685C.taxon	distribution	Distribution. Malaysia: Sabah (MOHAMEDSAID & KIMOTO 1993; MOHAMEDSAID 1995, 1997, 1999; MOHAMEDSAID & HOLLOWAY 1999).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEAFFC49A01FD9BFDA2696A.taxon	materials_examined	Type locality. ‘ Malaysia, Sabah, Lembah Danum’. Type material. Not examined.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEAFFC49A01FD9BFDA2696A.taxon	distribution	Distribution. Malaysia: Sabah (MOHAMEDSAID 2001).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEAFFC49A01FD9BFDA2696A.taxon	discussion	Comments. After the tentative transfer of this species to Monolepta it is now a homonym of Monolepta apicalis (Sahlberg, 1823). However, I avoid to suggest a new name till its definitive placement in Monolepta is confirmed.	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEAFFC49A1CFC4DFD426EAA.taxon	materials_examined	Type locality. ‘ Malaysia, Perak, Temengor’. Type material. HOLOTYPE: ♂, ‘ PERAK: Temengor / Ekspedisi MNS-Belum / 29 - 30 Jan. 1994 / Salleh & Ismail [w, p] // HOLOTYPE [p] / Liroetiella / sallehnori n. sp. [h] / des. Mohamedsaid 199 [p] 8 [h] [white label with red margins] ’ (UKM). PARATYPE: 1 spec. unsexed, ‘ PERAK: Temengor / Ekspedisi MNS-Belum / 15 - 20 Nov. 1993 / Salleh, Ismail & Sham [w, p] // PARATYPE [p] / Liroetiella / sallehnori n. sp. [h] / des. Mohamedsaid 199 [p] 8 [h] [white label with red margins] ’ (UKM).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEAFFC49A1CFC4DFD426EAA.taxon	distribution	Distribution. Malaysia: Perak (MOHAMEDSAID 1998).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEAFFC59A00FB0DFE6C6B6F.taxon	description	(Figs 64 – 65)	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEAFFC59A00FB0DFE6C6B6F.taxon	materials_examined	Type locality. ‘ Malaysia, Perak, Temengor’. Type material. HOLOTYPE: ♂, ‘ PERAK: Temengor / Ekspedisi MNS-Belum / 29 Nov. - 5 Dis. 1993 / Ismail, Yusof, Bidi, Sayful [w, p] // HOLOTYPE [p] / Liroetiella / warisan n. sp. [h] / des. Mohamedsaid 199 [p] 8 [h] [white label with red margins] ’ (UKM). PARATYPES: 1 spec. unsexed, ‘ PERAK: Temengor / Ekspedisi MNS-Belum / 29 Nov. - 5 Dis. 1993 / Ismail, Yusof, Bidi, Sayful [w, p] // PARATYPE [p] / Liroetiella / warisan n. sp. [h] / des. Mohamedsaid 199 [p] 8 [h] [white label with red margins] ’ (UKM); 1 ♀, ‘ PERAK: Temengor / Ekspedisi MNS-Belum / 29 Nov. - 5 Dis. 1993 / Ismail, Yusof, Bidi, Sayful [w, p] // PARATYPE [p] / Liroetiella / warisan n. sp. [h] / des. Mohamedsaid 199 [p] 8 [h] [white label with red margins] // COLLEZIONE / DACCORDI M. [pale blue label, p] ’ (MSNV).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEAFFC59A00FB0DFE6C6B6F.taxon	distribution	Distribution. Malaysia: Perak (MOHAMEDSAID 1998), Sabah (MOHAMEDSAID 1999, 2000, MOHAMEDSAID & HOLLOWAY 1999).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEBFFC59AD0FE4AFD1768B5.taxon	description	(Figs 62 – 63)	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEBFFC59AD0FE4AFD1768B5.taxon	materials_examined	Type locality. ‘ Leyte, VISCA N Baybay’. Type material. HOLOTYPE: ♀, ‘ PHILIPPINEN: LEYTE / VISCA N Baybay 1991 / Untersu- IV. - IX. / chungsbelege A. ENGLER [yellow label, p] // Chrys. / Ast 10 [w, h] // Holotypus [red label, p] // HOLOTYPUS [p] / Liroetiella / englerae / m. [h] / L. Medvedev det. 199 [p] 3 [w, h] ’ (SMNS).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
166387BCFFEBFFC59AD0FE4AFD1768B5.taxon	distribution	Distribution. Philippines: Leyte (MEDVEDEV 1995).	en	Bezděk, Jan (2013): Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae 53 (2): 715-746, DOI: 10.5281/zenodo.5740035
