taxonID	type	description	language	source
0987463F2E2D509F88594C782A30D269.taxon	description	Figure 5; Tables 2, S 2	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
0987463F2E2D509F88594C782A30D269.taxon	diagnosis	Diagnosis. Medium-sized gecko, SVL at least 68.8 mm in adult female; supralabials 9 – 12 and infralabials nine; 12 or 13 lamellae beneath the digit IV of manus; 17 lamellae beneath digit IV of pes; 17 feebly keeled and bluntly conical tubercles across mid dorsum and 32 paravertebral tubercles; 36 smooth mid-ventral scales between ventrolateral folds; at least 10 small precloacal pores in female; head on top pale-brown with purplish tinge; dark-brown postorbital stripe continuing to above the ear opening; a dark-brown cross bar with irregular edges on nape; indistinct pale-yellow patch on occipital region; dorsally and laterally neck and back pale yellowish-brown with dark-brown irregular reticulation.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
0987463F2E2D509F88594C782A30D269.taxon	description	Description of holotype. Holotype well preserved except an incision below left axilla ventrolaterally. Snout-vent length 68.8 mm. Head moderately large (HL / SVL = 0.26), dorsoventrally depressed, longer than width (HW / HL = 0.65), oval, distinct from neck, broader at occipital region; snout tip rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat; a longitudinal furrow on dorsal surface of the snout; snout short (SO / HL = 0.39), longer than orbit (OD / SO = 0.59); nostril rounded, opening directed posterolaterally; ear opening oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous, supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at the anterodorsal region; rostral wide, a short groove at the middle on top; rostral connected with nasals, supranasals and first supralabials; a single scale between the supranasals, larger than the rest of the granular snout scales; granular scales at parietal, occipital and temporal region intermixed with slightly larger rounded and bluntly conical granular tubercles starting from the level of posterior margin of the upper eyelids, smaller in parietal region, size increases towards nape; supralabial 12 on right side and nine on left side, supralabials count up to midorbit is eight on right side and six on left side, size decreases towards angle of jaw; a series of narrow, enlarged scales above the supralabials between nostril and anterior orbital border; mental triangular, connected with first infralabials and inner postmentals; nine infralabials on both side, size decreases towards angle of jaw; inner pair of postmentals are larger than the outer postmentals; posterior margin of the inner postmentals bordered by eight granular scales of different size; two rows of slightly enlarged, narrow scales, larger than gular scales present along the infralabials starting below the outer postmentals, posteriorly size decreases; gular scales granular, juxtaposed, homogeneous, size increases towards the throat where they become imbricate. Habitus slender (BW / SVL = 0.16, TRL / SVL = 0.47), dorsoventrally depressed; dorsal scales granular, rounded, heterogeneous, intermixed with densely placed rounded, weakly keeled and bluntly conical tubercles, irregularly arranged, continues to seventh segment of the tail, size increases towards posterior body and pronounced at the base of tail; 17 dorsal tubercles across mid dorsum; 32 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid, subimbricate to imbricate, largest on posterior part of the belly, 36 mid-ventral scales between ventrolateral folds; 10 small precloacal pores arranged in an inverted “ V ” shaped continuous series, followed by a series eight unpored, enlarged scales below it. Forelimbs and hindlimbs slender (FL / SVL = 0.14, CL / SVL = 0.18); digits strongly inflected at the joints, all bearing large recurved claw and enlarged subdigital lamellae; lamellae count beneath digit IV of right and left manus (given as basal + distal) is 5 + 7 and 5 + 8 respectively; lamellae count beneath digit IV of both right and left pes (given as basal + distal) is 8 + 9; dorsal scales on forelimbs heterogeneous in size; hind arm scales smooth and subimbricate; forearm scales small and granular towards proximal and towards distal end it is smooth, cycloid and imbricate; forearm scales intermixed with enlarged feebly keeled tubercles; dorsal scales of hindlimbs heterogeneous, intermixed with densely placed large, rounded and bluntly conical tubercles; scales on inner lateral and dorsolateral side of the thighs smooth, large and imbricate, rest of the scales are granular; scales on tibia are small, granular juxtaposed; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous in shape and size, granular juxtaposed; ventral scales of hind limbs smaller than those of belly, smooth, cycloid and subimbricate; scales on the knee, above cloaca and on thigh below the level of precloacal pores smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate. Tail complete, posterior 6 mm regenerated (TL = 64 mm), slender, gradually tapering towards tip; tail segments indistinct; dorsal scales small, granular, juxtaposed at the base, posteriorly size of the scales increases, flat, smooth, subimbricate, heterogeneous in shape and size; enlarged feebly keeled scales up to seventh segment of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; four bluntly conical spurs on both sides of the tail base.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
0987463F2E2D509F88594C782A30D269.taxon	etymology	Etymology. The specific epithet is a toponym derived from the name of the hill range “ Barail ” where the type locality of the species lies.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
0987463F2E2D509F88594C782A30D269.taxon	distribution	Distribution and natural history. During our two-day survey we only located a single individual of this species. Thus, the new species is currently known only from the type locality, in Peren District, Nagaland, India. We recorded this species on the trunk of a small tree at a height of approximately 2 m from the ground in the Athibung Reserve Forest at approximately 20: 00 hrs on 14 August 2021. The forest type is semi-evergreen with relatively little anthropogenic pressure.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
507D9749BFBD54018DADAD5538DE287C.taxon	description	Figures 10, S 3; Tables 2, S 4	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
507D9749BFBD54018DADAD5538DE287C.taxon	distribution	Distribution and natural history. The type locality, Xizang (Tibet), China by C. cayuensis also appears to be widely distributed in Arunachal Pradesh, India (Fig. 3 B), within an elevational range of 410 – 1200 m a. s. l. Cyrtodactylus cayuensis is widespread in the foothills of Arunachal Pradesh from the western part of the state, at Seijusa (Pakke Tiger Reserve) to the eastern part (Kamlang Tiger Reserve) and it has successfully spread across major tributaries of the Brahmaputra River, i. e., the Subansiri, Siang, Dibang, Lohit up to at least north of Noa-Dihing. Populations of C. cayuensis have been recorded in degraded evergreen forest as well as in undisturbed semi-evergreen and evergreen forests. Cyrtodactylus cayuensis is particularly abundant in the vegetation close to the picturesque Glaw Lake in the Kamlang Tiger Reserve. We observed individuals at night on shrubs and bedrocks (sedimentary rock) along small forest streams. We also encountered them on small trees up to a 2.0 m inside the forest at Jengging, Kamlang Tiger Reserve and Mehao Wildlife Sanctuary.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
CAD2A307ECEF5751BF30BB86E58CF3CE.taxon	description	Figure 4; Tables 2, S 2	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
CAD2A307ECEF5751BF30BB86E58CF3CE.taxon	diagnosis	Diagnosis. A medium-sized gecko (SVL up to at least 64.7 mm); 10 supralabials; 9 – 12 infralabials; 16 bluntly conical and feebly keeled dorsal tubercles; 26 – 29 paravertebral tubercles; 35 or 36 midventral scales rows between the weak ventrolateral folds; no precloacal groove; six or seven precloacal pores in continuous series; 12 – 15 total subdigital lamellae beneath digit IV of pes; dark brown irregular cross bars or reticulation on dorsum.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
CAD2A307ECEF5751BF30BB86E58CF3CE.taxon	description	Description of the holotype. Holotype well preserved except single incision below left axilla ventrolaterally. Snout-vent length 63.9 mm. Head moderately large (HL / SVL = 0.26), dorsoventrally depressed, longer than width (HW / HL = 0.68), distinct from neck, broader at occipital region; snout tip rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat; a longitudinal furrow on dorsal surface of the snout; snout short (SO / HL = 0.4), longer than orbit (OD / SO = 0.7); nostril nearly rounded, opening directed posterolaterally; ear opening oval and oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous, supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at approximately anterior one third of it; rostral wide, a short groove at the middle on top; rostral connected with nasals, supranasals, internasal and first supralabials; two scale between the supranasals, larger than the rest of the granular snout scales; granular scales at parietal region and occipital region intermixed with slightly large rounded granular tubercles starting from the level of posterior margin of the upper eyelids, size increases towards nape; supralabial 10 on both side, seven supralabials upto midorbit, size decreases towards angle of jaw; a series of narrow, enlarged scales above the supralabials between nostril and anterior orbital border; mental slightly wider than rostral (RW / MW = 0.8), triangular, connected with first infralabials, inner postmentals; nine infralabials on both sides, size decreases towards angle of jaw; first infralabials connected with mental, second infralabial, inner and outer postmentals; inner pair of postmentals are larger than the outer postmentals; posterior margin of the inner postmentals bordered by eight granular scales of different size; two rows of enlarged scales along the infralabials starting below the outer postmentals, posteriorly size decreases, elongated and narrow; rest of the gular scales are small, granular juxtaposed, homogeneous, size increases towards the throat where they become imbricate. Habitus slender (BW / SVL = 0.2, TRL / SVL = 0.35), dorsoventrally depressed; dorsal scales granular, rounded, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, up to fourth segment of the tail, size increases towards posterior body and pronounced; 16 dorsal tubercles across mid dorsum; 26 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate, largest on belly; 36 mid-ventral scales between ventrolateral fold; seven precloacal pores arranged in an inverted “ V ” shaped continuous series, followed by five unpored, large scales below it, largest at the apex. Forelimbs and hindlimbs slender (FL / SVL = 0.13, CL / SVL = 0.16); digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae; lamellae beneath digit IV of right and left manus (given as basal + distal) is 5 + 8 and 5 + 9 respectively; lamellae beneath digit IV of right and left pes (given as basal + distal) is 5 + 10 and 5 + 9 respectively; dorsal scales on forelimbs smooth and subimbricate, small and granular at elbow and proximal end of forearm; forearm scales intermixed with rounded large tubercles; dorsal scales of hindlimbs heterogeneous, intermixed with large, rounded and bluntly conical tubercles; horizontally upper half of the thigh scales are smooth, large and subimbricate, those on lower half small granular; scales on tibia are small, granular juxtaposed; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous in shape and size, granular juxtaposed; scales on ventral side of hindlimbs smaller than those of belly, smooth, cycloid and subimbricate, but on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate. Tail regenerated (TL = 68 mm), slender, gradually tapering towards tip, segments indistinct, dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size; large feebly keeled scales upto fourth segment of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal and widest at the middle, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; two and three bluntly conical spurs on right and left side of the tail base respectively.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
CAD2A307ECEF5751BF30BB86E58CF3CE.taxon	etymology	Etymology. The specific epithet is derived from Kiphire, a District of Nagaland from where the type series of this species were collected.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
CAD2A307ECEF5751BF30BB86E58CF3CE.taxon	distribution	Distribution and natural history. We recorded Cyrtodactylus kiphire sp. nov. from Kiphire forest colony, Kiphire District, Nagaland, India. The area is characterised as subtropical forest with regenerating jhum forest dominated by Castanopsis indica (Roxb. ex Lindl.) A. DC., Quercus sp., Itea macrophylla Wall. ex Roxb., Albizia chinensis (Osbeck) Merr .. The holotype was collected from on a forest trail shrub (approximately 2.0 m from the ground) at approximately 23: 00 hrs. The paratype was collected from a rocky slope along the edge of a stream.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
739EA53FD4D75FBFB2DB62ECAC41A8D3.taxon	description	Figure 6; Tables 2, S 2	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
739EA53FD4D75FBFB2DB62ECAC41A8D3.taxon	diagnosis	Diagnosis. Medium-sized gecko, SVL at least 59.5 mm in adult male; 10 supralabials; eight or nine infralabials; 21 bluntly conical and feebly keeled tubercles across midbody; 37 paravertebral tubercles; 36 mid-ventral scales between ventrolateral folds; 11 or 12 subdigital lamellae beneath digit IV of manus; 13 – 16 subdigital lamellae beneath digit IV of pes; seven precloacal pores arranged in a continuous series; six irregular shaped dark-brown cross bands on back between axilla and groin; tail dorsally with eight dark-brown and seven pale-brown bands arranged alternatively.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
739EA53FD4D75FBFB2DB62ECAC41A8D3.taxon	description	Description of the holotype. Specimen well preserved except an incision below left axilla ventrolaterally. Snout-vent length 59.5 mm. Head moderately large (HL / SVL = 0.26), oval, dorsoventrally depressed, longer than width (HW / HL = 0.71), distinct from neck, broader at occipital region; snout tip rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat; a longitudinal furrow on dorsal surface of the snout; snout short (SO / HL = 0.39), longer than orbit (OD / SO = 0.68); nostril nearly rounded, opening directed posterolaterally; ear opening rounded; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous; supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at the anterodorsal region; rostral wide, a short groove at the middle on top, rostral connected with nasals, supranasals, an internasal and first supralabials, two scales between the supranasals paced longitudinally, granular scales at parietal, occipital and temporal region intermixed with slightly large rounded granular tubercles starting from the level of posterior margin of the upper eyelids, those on temporal region are slightly larger than that of occipital region, size of the tubercles increases towards nape; 10 supralabial scales on both sides, size decreases towards angle of jaw; 8 supralabials up to midorbit on both sides; mental nearly triangular, connected with first infralabials, inner postmentals; nine infralabials on right side and eight on left side, size decreases towards angle of jaw; first infralabials connected with mental, second infralabial, inner and outer postmentals; inner pair of postmentals are larger than the outer postmentals; two rows of slightly enlarged scales along the infralabials starting below the outer postmentals, posteriorly size of those decreases; rest of the gular scales are small, granular juxtaposed, nearly homogeneous, size increases towards the throat where they become imbricate. Habitus slender (BW / SVL = 0.16, TRL / SVL = 0.49), dorsoventrally depressed, dorsal scales granular, rounded, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, up to third segment of the tail, size increases towards posterior body and pronounced at the base of tail; 21 dorsal tubercles across mid dorsum; 37 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate; 36 mid-ventral scales between ventrolateral fold; seven precloacal pores arranged in an inverted “ V ” shaped continuous series, followed by a series of five unpored enlarged scales below it, largest at the apex, an unpored scale equal size to PcP present in continuous with the PcP in both ends; scales above the PcP larger than those of belly scales. Forelimbs and hindlimbs slender (FL / SVL = 0.14, CL / SVL = 0.17), digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae; lamellae beneath digit IV of right and left manus (given as basal + distal) is 6 + 5 and 6 + 6 respectively; lamellae beneath digit IV of both right and left pes (given as basal + distal) is 6 + 7 and 7 + 9 respectively; dorsal scales on forelimbs heterogeneous is size, mostly granular; proximal scales on upper arm are smaller than that of lower arm; scales near elbow on lower arm smooth and subimbricate; upper arm scales granular at proximal part and at the distal end it is smooth, cycloid and imbricate; dorsal scales of hindlimbs heterogeneous, intermixed with slightly enlarged, feebly keeled and bluntly conical tubercles; scales on inner lateral side of the thighs are smooth, large and subimbricate, those on dorsal side are small granular; scales on tibia are small, granular juxtaposed; ventral scales of forelimbs granular, juxtaposed, heterogeneous; scales on palm heterogeneous in shape and size, granular juxtaposed; ventral scales on hindlimbs heterogeneous; most of the thigh scales are smooth, cycloid and subimbricate, but on the knee and below the level of precloacal pores, scales are smaller and granular; scales above the vent granular; tibia scales smooth, nearly homogeneous, cycloid and subimbricate; scales on soles heterogeneous, granular, juxtaposed to subimbricate. Tail complete (TL = 67 mm), slender, gradually tapering towards tip, segments indistinct, dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size, large feebly keeled scales up to third segment of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; three bluntly conical spurs on both sides of the tail base.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
739EA53FD4D75FBFB2DB62ECAC41A8D3.taxon	etymology	Etymology. This species is named after Manipur state in India.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
739EA53FD4D75FBFB2DB62ECAC41A8D3.taxon	distribution	Distribution and natural history. Cyrtodactylus manipurensis sp. nov. is currently only recorded from the type locality. We collected a single individual at 18: 00 hrs on 25 July 2022 near Lamdan Kabui village. It was perched on a shrub at a height of approximately 1.5 m, on the road connecting Leimatak and Charoikhullen. The habitat is secondary forest, with Zingiber sp. cultivation and settlements.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
6B05318011F35D7C95E621A314732FC0.taxon	description	Figure 8; Tables 2, S 2	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
6B05318011F35D7C95E621A314732FC0.taxon	diagnosis	Diagnosis. Medium-sized gecko (SVL 57.5 – 70.7 mm in males and SVL 54.8 – 69.3 mm in females); supralabials and infralabials 8 – 11; dorsum with weakly keeled and bluntly conical tubercles, 29 – 36 paravertebral tubercles between the level of axilla and level of groin; 17 – 19 dorsal tubercle rows at mid body; 33 – 40 mid ventral scale rows; seven to nine precloacal pores in males and 8 – 10 small precloacal pores in females in a continuous series, PcP much smaller in females than that of males; 12 – 14 subdigital lamellae on finger IV and 11 – 17 subdigital lamellae on toe IV; irregular dark-brown spots or stripes on dorsum.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
6B05318011F35D7C95E621A314732FC0.taxon	description	Description of holotype. Holotype well preserved except a ventrolateral incision below left axilla. Snout-vent length 65.3 mm. Head moderately large (HL / SVL = 0.27), dorsoventrally depressed, ovoid in shape, longer than width (HW / HL = 0.66), distinct from neck, broader at occipital region; snout rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat, a longitudinal furrow on dorsal surface of the snout, snout short (SO / HL = 0.4), longer than orbit (OD / SO = 0.71); nostril semicircular, opening directed posterolaterally; ear opening oval and oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids nearly homogeneous, granular juxtaposed; supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at the anterodorsal region; rostral wide, a short groove at the middle on top, rostral connected with nasals, supranasals, internasals and first supralabials; granular scales at parietal, occipital and temporal region intermixed with slightly large, rounded and bluntly conical tubercles, dense in occipital and temporal region and size increases towards nape; supralabial 11 on right and 10 on left side, supralabials up to midorbit eight on right and seven on left side, size decreases towards angle of jaw; a series of scales slightly larger than the loreal scales present above the supralabials, posteriorly size decreased; mental as wide as rostral, nearly triangular, connected with first infralabials, inner postmentals; nine infralabials on right and eight on left side, size decreases towards angle of jaw; inner pair of postmentals are larger than the outer postmentals; two rows of enlarged scales along the infralabials starting below the outer postmentals, posteriorly size of those decreases; rest of the gular scales are small, granular juxtaposed, homogeneous, size increases towards throat where they becomes imbricate. Habitus slender (BW / SVL = 0.17, TRL / SVL = 0.47), dorsoventrally depressed; dorsal scales granular, rounded, heterogeneous, intermixed with rounded, irregularly arranged weakly keeled and bluntly conical tubercles, these tubercles continues to fourth segment of the tail, size increases towards posterior body and pronounced; 18 dorsal tubercles across mid dorsum; 36 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate, largest towards belly; 40 mid-ventral scales between ventrolateral fold; seven precloacal pores arranged in an inverted “ V ” shaped continuous series, followed by five unpored, large scales below it, one enlarged unpored scale present on right end of the PcP series. Forelimbs and hindlimbs slender (FL / SVL = 0.15, CL / SVL = 0.18); digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae; lamellae beneath digit IV of right and left manus (given as basal + distal) is 5 + 8; lamellae beneath digit IV of right and left pes (given as basal + distal) is 6 + 8 and 5 + 8 respectively; dorsal scales on forelimbs smooth, subimbricate and heterogeneous, small and granular at elbow, scales on forearm nearly rounded while those on hind arm are posteriorly tapering; forearm scales intermixed with enlarged rounded and bluntly conical tubercles; dorsal scales of hindlimbs heterogeneous, intermixed with large, rounded and bluntly conical tubercles, dense than those on forelimbs; horizontally upper half of the thigh scales are smooth, large and subimbricate, those on lower half small granular; scales on tibia are small, granular juxtaposed; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous in shape and size, granular juxtaposed; scales on ventral side of hindlimbs smaller than those of belly, smooth, cycloid and subimbricate, but on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate. Tail regenerated (TL = 79 mm), slender, gradually tapering towards tip, segments indistinct, in the original part of the tail dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size; in the regenerated part of the tail, scales are irregular in shape and size; large feebly keeled scales upto fourth segment of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; two and four bluntly conical spurs on right and left side of the tail base respectively.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
6B05318011F35D7C95E621A314732FC0.taxon	etymology	Etymology. The specific epithet is a toponym named after its type locality Namdapha Tiger Reserve in Arunachal Pradesh, India.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
6B05318011F35D7C95E621A314732FC0.taxon	distribution	Distribution and natural history. So far, Cyrtodactylus namdaphaensis sp. nov. has been recorded within an elevational range of 400 – 650 m a. s. l. inside Namdapha Tiger Reserve. All the localities are south of Noa-Dihing River in Changlang District, Arunachal Pradesh, India. We recorded this species between May and September 2022. Individuals were recorded on tree bark, ferns, and riparian vegetation and along forest trails between 18: 00 – 23: 00 hrs. The forest type can be classified as Assam Valley Tropical Evergreen Forest. The area had a distinct understory with a thick covering of leaf-litter. Currently the species has been recorded from 25 Mile, Burma Nullah (40 Mile), Gibbons Land, Motijheel trail and at Hornbill camp located at the north bank of Noa-Dihing River within the Namdapha Tiger Reserve. We also recorded this species on rocks and vegetation near Kamlang River at Sinabrai, near the Kamlang Tiger Reserve, at an elevation of 470 m a. s. l. We observed juveniles of this species on the forest floor in the leaf litter during May, 2022. During 2023, adult individuals were seen on the ferns overhanging first order streams. At the slightest disturbance, the lizards would drop into the thick vegetation below. Other arboreal reptile taxa from the area included Ptyctolaemus sp., Japalura sp. and Pareas sp.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
0F6F61B9C7765FC09DEBF74295D25A34.taxon	description	Figure 7; Tables 2, S 2	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
0F6F61B9C7765FC09DEBF74295D25A34.taxon	diagnosis	Diagnosis. Medium-sized gecko (SVL at least 61.2 mm in adult male and 72 – 74.1 mm in adult females); 9 – 12 supralabials; 8 – 11 infralabials; 18 – 20 bluntly conical and feebly keeled dorsal tubercles; 29 – 34 paravertebral tubercles; 38 or 39 midventral scale rows between the weak ventrolateral folds; no precloacal groove; at least 27 precloacofemoral pores in continuous series in male and 10 – 16 small precloacal pores in females; short dark-brown bars between a pair of dorsolateral stripes or enlarged irregular dark-brown spots present on dorsum.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
0F6F61B9C7765FC09DEBF74295D25A34.taxon	description	Description of holotype. Holotype well preserved except an incision below left axilla ventrolaterally. Snout-vent length 61.2 mm. Head moderately large (HL / SVL = 0.27), dorsoventrally depressed, longer than width (HW / HL = 0.68), distinct from neck, broader at occipital region, snout tip rounded in both dorsal and lateral view, loreal region convex, canthus rostralis rounded, indistinct, interorbital space flat, a longitudinal furrow on dorsal surface of the snout, snout short (SO / HL = 0.4), longer than orbit (OD / SO = 0.66), nostril nearly rounded, opening directed posterolaterally, ear opening oval and oblique, scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed, scales on upper eyelids heterogeneous, supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anterior and posterior end decreases, rostral wide, a short groove at the middle on top, rostral connected with nasals, supranasals, an internasal and first supralabials, two scales between the supranasals, larger than the rest of the granular snout scales, granular scales at parietal region and occipital region intermixed with slightly large rounded granular tubercles, dense in occipital and temporal region and size increases towards nape, nine supralabials on right and 10 on left side, size decreases towards angle of jaw, supralabials up to midorbit seven on right and eight on left side, a series of narrow and slightly elongated scales above the supralabials between nostril and anterior border of the orbit, mental as wide as rostral, triangular, connected with first infralabials, inner postmentals, nine infralabials on both side, size decreases towards angle of jaw, first infralabials connected with mental, second infralabial, inner and outer postmentals, inner pair of postmentals are larger than the outer postmentals, posterior margin of the inner postmentals bordered by seven granular scales of different size, two or three rows of enlarged scales along the infralabials starting below the outer postmentals, posteriorly size of those decreases, elongated and narrow, rest of the gular scales are small, granular juxtaposed, homogeneous, size increases towards the throat where they become imbricate. Habitus slender (BW / SVL = 0.17, TRL / SVL = 0.41), dorsoventrally depressed, dorsal scales granular, rounded, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, starting from occipital region to seventh segment of the tail, size increases towards posterior body, pronounced at sacrum and base of the tail, 18 dorsal tubercles across mid dorsum, 29 paravertebral tubercles, ventrolateral fold weak, ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate, largest on belly, 39 mid-ventral scales between ventrolateral fold, 27 precloacal femoral (PcFP) pores in a continuous series (Fig. 7 I), followed by eight unpored, large scales below the PcFP at the middle. Forelimbs and hindlimbs slender (FL / SVL = 0.14, CL / SVL = 0.19); digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae; lamellae beneath digit IV of right and left manus (given as basal + distal) is 5 + 8 and 5 + 9 respectively; lamellae beneath digit IV of right and left pes (given as basal + distal) is 4 + 10 and 5 + 10 respectively; dorsal scales on forelimbs heterogeneous, granular juxtaposed; dorsal scales of hindlimbs heterogeneous, granular, intermixed with densely placed large, rounded and bluntly conical tubercles, scales on inner lateral side of the thighs near knee are subimbricate; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous in shape and size, granular; scales on ventral side of hindlimbs nearly equal to those of belly, smooth, cycloid and subimbricate, but on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate. Tail complete (TL = 70 mm), slender, gradually tapering towards tip, segments indistinct, dorsal scales granular, juxtaposed, flat, smooth, heterogeneous in shape and size; enlarged feebly keeled scales up to eight segments of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
0F6F61B9C7765FC09DEBF74295D25A34.taxon	etymology	Etymology. The specific epithet is a toponym derived from the name “ Ngengpui Wildlife Sanctuary ” of Mizoram state from where the type series of the species were collected.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
0F6F61B9C7765FC09DEBF74295D25A34.taxon	distribution	Distribution and natural history. Cyrtodactylus ngengpuiensis sp. nov. new species is currently only known from the type locality, the Ngengpui Wildlife Sanctuary, Lawngtlai District, Mizoram, India. The forest is characterised as tropical semi-evergreen to moist evergreen forest. The forest is dominated by Dipterocarpus spp., palms, canes and rattans. Individuals of C. ngengpuiensis sp. nov. were collected during 6 – 9 September 2021 at 21: 00 – 22: 00 hrs. Individuals were recorded in bamboo thickets, on tree buttresses and trunks, and amongst ferns and rocks along the banks of evergreen forest streams.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
CB20AD95DD3E53A58D7FC3B88F1BCC17.taxon	description	Figure 9; Tables 2, S 2	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
CB20AD95DD3E53A58D7FC3B88F1BCC17.taxon	diagnosis	Diagnosis. Medium-sized gecko (SVL 70.1 – 72.1 mm in females); males unknown; supralabials 8 – 12 and infralabials 9 – 12; tubercles on dorsum weakly keeled and bluntly conical, 26 – 32 paravertebral tubercles between the level of axilla and level of groin; 15 or 16 dorsal tubercle rows at mid body; 40 – 45 mid ventral scale rows; 8 – 10 small precloacal pores in a continuous series; 13 – 17 subdigital lamellae on finger IV and 14 – 19 subdigital lamellae on toe IV; six or seven irregular and broken dark-brown bands on dorsum between the level of axilla and level of groin, or irregular dark-brown reticulation on dorsum.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
CB20AD95DD3E53A58D7FC3B88F1BCC17.taxon	description	Description of the holotype. Holotype well preserved except an incision below left axilla ventrolaterally. Snout-vent length 72.1 mm. Head moderately large (HL / SVL = 0.26), dorsoventrally depressed, longer than width (HW / HL = 0.72), distinct from neck, broader at occipital region; snout rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat, a longitudinal furrow on dorsal surface of the snout, snout short (SO / HL = 0.39), longer than orbit (OD / SO = 0.73); nostril nearly rounded, opening directed posterolaterally; ear opening oval and oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous; supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at approximately anterior one third of it; rostral wide, a short groove at the middle on top, rostral connected with nasals, supranasals, an internasal and first supralabials; a single scale between the supranasals, larger than the rest of the granular snout scales; granular scales at parietal region and occipital region intermixed with slightly large rounded granular tubercles, dense in occipital and temporal region and size increases towards nape; supralabial eight on right and 11 on left side, supralabials upto midorbit seven on right and eight on left side, size decreases towards angle of jaw; a series of scales nearly rounded, granular and slightly larger than the loreal scales present above the supralabials; mental as wide as rostral, triangular, connected with first infralabials, inner postmentals; 10 infralabials on right and nine on left side, size decreases towards angle of jaw, first infralabials connected with mental, second infralabial, inner and outer postmentals; inner pair of postmentals are larger than the outer postmentals, posterior margin of the inner postmentals bordered by eight granular scales of different size; one or two rows of enlarged scales along the infralabials starting below the outer postmentals, posteriorly size of those decreases, elongated and narrow; rest of the gular scales are small, granular juxtaposed, homogeneous, size increases towards throat where they become imbricate. Habitus slender (BW / SVL = 0.18, TRL / SVL = 0.45), dorsoventrally depressed; dorsal scales granular, rounded, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, starting from occipital region to fourth segment of the tail, size increases towards posterior body, pronounced at sacrum and base of the tail; 15 dorsal tubercles across mid dorsum; 26 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate, largest on belly; 40 mid-ventral scales between ventrolateral fold; very small, 10 precloacal pores arranged in an inverted “ V ” shaped continuous series, followed by 7 unpored, large scales below it, largest at the apex. Forelimbs and hindlimbs slender (FL / SVL = 0.14, CL / SVL = 0.17); digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae, lamellae beneath digit IV of right and left manus (given as basal + distal) is 6 + 11 and 6 + 10 respectively, the second most lower lamellae of the basal series of digit IV of both sides are divided (not included in lamellar count); lamellae beneath digit IV of right and left pes (given as basal + distal) is 7 + 11 and 6 + 10 respectively; dorsal scales on forelimbs heterogeneous, granular juxtaposed, smooth and subimbricate at distal end of forearm; forearm intermixed with a few rounded, large tubercles; dorsal scales of hindlimbs granular intermixed with large, rounded, bluntly conical tubercles; scales on thigh towards knee smooth subimbricate; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous, granular juxtaposed; scales on hindlimbs smaller than those of belly, smooth, cycloid and subimbricate; scales on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate. Tail complete, regenerated (TL = 80 mm), slender, gradually tapering, segments indistinct; dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size; large feebly keeled scales up to fourth segment of the tail at the distal end of each segment, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; below cloaca scales are granular, smaller than the rest of the tail scales; no enlarged plate like series of subcaudal scales; three bluntly conical spurs on each side of the tail base present.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
CB20AD95DD3E53A58D7FC3B88F1BCC17.taxon	etymology	Etymology. The specific epithet is a toponym derived from the name of the river “ Siang ”. The type locality of this species lies in the Siang valley of Arunachal Pradesh.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
CB20AD95DD3E53A58D7FC3B88F1BCC17.taxon	distribution	Distribution and natural history. Cyrtodactylus siangensis sp. nov. is currently only known from the type locality, Bodak, East Siang District. It appears to be uncommon, and likely a habitat generalist. The holotype was encountered on a shrub at the edge of Kalek stream (a tributary of Siang River) at a height of approximately one meter above the ground, at 18: 15 hrs. The stream at the time of survey was flowing at a moderately speed, was approximately 10 m in width and contained boulders (sedimentary rock) and fallen logs. The two paratypes were recorded on a small tree at a height of 2.0 m on the slope of a hill close to the collection site for the holotype. The type locality is ca. 300 m from Siang River and approximately 150 m from the road connecting Pasighat and Yinkiong.	en	Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V., Das, Abhijit (2024): Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453-486, DOI: 10.3897/vz.74.e124752
