taxonID	type	description	language	source
1509C719C6434350FDF6FBA5FE67C48D.taxon	diagnosis	Diagnosis (updated from Townes 1969 and Gauld 1991) Small to large insects (fore wing length 6 – 25 mm), usually orange or yellow interspersed with black maculae; clypeus very small, not subdivided, often with a subapical ridge bearing median and / or lateral weak to strong tubercles; ventral margin of clypeus thin, slightly concave, often with lateral corners produced; mandible not twisted, stout, strongly constricted basally, then weakly tapered, apically bidentate with lower tooth pointed (may appear chisel-like when worn) and upper tooth blunt or chisellike; occipital carina complete to nearly absent, when ventrally present joining hypostomal carina above mandibular base; hypostomal carina strongly raised above mandible base; subocular sulcus absent, but malar area granulate; pronotum long, mid-dorsally with a deep invagination and with thin margin projecting beyond invagination; antero-lateral margin of pronotum broadly rounded in front, its lower corner acute and slightly in-turned; epomia weak to moderately strong, contiguous with and diverging from anterior pronotal margin; mesoscutum with strong transverse rugosities; notauli anteriorly strong and meeting in centre of mesoscutum, defining an anteriorly abruptly rounded median lobe; mesopleuron moderately long, epicnemial carina most often present laterally and reaching to or above lower corner of pronotum, its dorsal end evanescent; mesopleural suture weakly angled centrally; submetapleural carina weak to strong, more or less complete; propodeum of moderate length, without carina dorsally but pleural carina distinct; tarsal claws simple, large; fore wing with areolet open, 2 m – cu slightly basal to distinctly apical to rs – m; hind wing with distal abscissa of Cu present, joining Cu & cu – a near or at junction with M; mid trochantellus with a ventral longitudinal ridge; metasoma moderately slender, tergites smooth to finely sculptured; tergite 1 fused with its sternite and without glymma; apical margins of tergites 3 – 5 hardly to strongly concave; female with sternites 2 – 4 (but see E. brianfisheri sp. nov.) with an anterior pair of tubercles, and with an apical truncate horn-like process on last tergite, ovipositor stout, laterally compressed, 4 – 5 x longer than hind tibia; male with metasoma slenderer and subgenital plate elongate, posteriorly rather concave.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6424356FDC9FD81FB15C32A.taxon	description	urn: lsid: zoobank. org: act: 1 C 872313 - 52 C 3 - 4 D 81 - B 8 AD- 94 AA 2 C 724 EAC Fig. 1	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6424356FDC9FD81FB15C32A.taxon	diagnosis	Diagnosis Overall color mottled, pale yellow and dark brown to black; wings hyaline, apically infuscate; face rugulose-punctate, strongly protruding medially; clypeus sparsely punctate, with median subapical tubercle strong and lateral subapical tubercle weak, ventral margin strongly produced laterally; frons rugulose with a blunt Y-shaped mid-longitudinal carina; antenna with 33 flagellomeres; mesopleuron with shallow transverse striations; epicnemial carina hardly reaching level of mesopleural pit; subtegular ridge convex; fore wing with 2 m – cu opposite rs – m; tergite 1 slender; sternite 1 with an additional pair of basal tubercles; tergites 1 – 2 smooth, following tergites densely and finely punctate; apical margins of tergites 3 – 5 moderately concave. CT 3.3; ML 0.7; POL 2.0; OOL 1.5; Fl 1 4.2; Fl 15 1.8; Fl 32 1.2; T 1 1.7; OT 3.9. Differential diagnosis Species characterized by the exceptional mottled coloration and the presence of antero-basal tubercles on sternite 1 (cf. taxonomic remarks). All other species of Epirhyssa lack the antero-basal tubercles.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6424356FDC9FD81FB15C32A.taxon	etymology	Etymology Dedicated to Brian Fisher, friend and expert organizer of expeditions into central Africa, including the CAR expedition that discovered this species.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6424356FDC9FD81FB15C32A.taxon	materials_examined	Type material Holotype CENTRAL AFRICAN REPUBLIC: ♀, Préfecture Sangha-Mbaéré, Réserve Spéciale de Forêt Dense de Dzanga-Sangha, 12.7 km, 326 ° NW of Bayanga, 3 ° 00.27 ’ N 16 ° 11.55 ’ E, 420 m, 13 May 2001, S. van Noort, Sweep CAR 01 – S 158, Lowland Rainforest, SAM – HYM – P 048018 (SAMC).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6424356FDC9FD81FB15C32A.taxon	description	Description Female (holotype) B 18.6; A 11.0; F 12.8. COLOR. Head pale yellow with black to dark brown parts: mandible, malar space, median tubercle of clypeus, a thin mid-longitudinal line on upper face, frons, vertex, and most of occiput; mesosoma pale yellow with most of mesoscutum black, and testaceous to brown parts: anterior margin of pronotum, propleuron, most of mesopleuron but subtegular ridge and two lateral patches, mesosternum, ventral margin of metapleuron and anterior third of propodeum; metasoma dark brown with pale yellow markings: tergites 1 – 2 medially, tergite 3 – 4 posteriorly; sternites pale yellow, more or less extensively testaceous anteriorly; antenna dark brown with scape somewhat pale maculated; ovipositor sheath brown; fore and mid legs testaceous with coxae and tibiae mostly pale yellow, hind leg testaceous with coxa partly pale yellow and femur with an outer longitudinal pale yellow stripe; wings hyaline, fore wing anteriorly infuscate from pterostigma to apex. HEAD. Densely setose with long hairs, hairs shorter on upper back head; face rugulose punctate, strongly protruding medially; clypeus sparsely punctate, subapical ridge with median tubercle strong and lateral tubercle weak, ventral margin strongly produced laterally; mandible stout; frons transversely rugose with a blunt Y-shaped mid-longitudinal carina below median ocellus, inner margins of toruli not expanded backwards; vertex and temple sparsely punctate but inter-ocellar area rugose; ocellar triangle wide; occipital carina mid-dorsally interrupted; antenna with 33 flagellomeres. MESOSOMA. Entirely covered with dense and long hairs; mesosoma moderately densely punctate, but mesonotum deeply transversely striate, anterior half of pronotum smooth and mesopleuron somewhat transversely strigose medially; epomia moderate; epicnemial carina hardly reaching level of mesopleural pit; apex of subtegular ridge convex, not flanged laterally; submetapleural carina strong, slightly and evenly narrowed toward apex. WINGS. Fore wing with 2 m – cu opposite rs – m, cu – a almost opposite Rs & M, and Rs strongly bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a close to junction with M, with a basal group of two hamuli on distal abscissa of R 1. METASOMA. Tergite 1 moderately slender; sternite 1 with a basal pair of submedian tubercles, additional to usual ones on sternites 2 – 4; tergites 1 – 2 mostly smooth, following tergites densely and finely punctate with fine transverse striations medially; apical margins of tergites 3 – 5 moderately concave. Male Unknown.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6424356FDC9FD81FB15C32A.taxon	distribution	Distribution Central African Republic.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6424356FDC9FD81FB15C32A.taxon	discussion	Taxonomic remarks The presence of tubercles on sternite 1 is quite unexpected in Epirhyssa, while it is one of the features defining the Oriental genus Triancyra Baltazar, 1961. However, we refrain from placing this new species in Triancyra for the following reasons. Besides the geographical separation from the known distribution area of Triancyra, the majority of morphological characters exhibited by this new species do not fit the current generic definition for Triancyra. For example, E. brianfisheri sp. nov. exhibits subequal mandibular teeth (vs. upper tooth narrower in Triancyra), a pterostigma that is long and slender (vs. up to 5 x longer than high), hind wing with a basal group of two hamuli (vs. usually three), tergite 1 long and slender (vs. stout, 1.3 x longer than wide) (Townes, 1969). Hence, we are of the opinion that the species fits more comfortably within Epirhyssa than in Triancyra, but the possession of additional tubercles in Epirhyssa species underlines the need for a global phylogenetic revision of the whole subfamily. Epirhyssa is indeed poorly defined and probably not monophyletic (G. Broad, pers. comm.).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C644434BFDCBF9D8FEDDC426.taxon	description	urn: lsid: zoobank. org: act: 94 C 30 C 69 - E 07 C- 4 C 48 - BB 79 - 3 B 3 D 903665 EE Fig. 2	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C644434BFDCBF9D8FEDDC426.taxon	diagnosis	Diagnosis Bright yellow overall with isolated dorsal black markings; wings hyaline, slightly infuscate apically; face rugulose-punctate, moderately protruding dorsally; clypeus punctate, with median subapical tubercle moderate, ventral margin strongly produced laterally; frons smooth with a sharp mid-longitudinal carina reaching median ocellus and lateral curved carina toward lateral ocellus; antenna with more than 35 flagellomeres; mesopleuron with epicnemial carina reaching level of mesopleural pit; subtegular ridge concave, laterally flanged; fore wing with 2 m – cu slightly apical to rs – m; tergite 1 slender; tergites 1 – 5 smooth, following tergites densely and finely punctate; apical margins of tergites 3 – 7 strongly concave. CT 2.7; ML 0.5; POL 1.3; OOL 1.6; Fl 1 4.2; Fl 15 1.8; T 1 1.7; OT 5.2. Differential diagnosis A mostly bright yellow species with hyaline wings, otherwise characterized by the combination of the punctate clypeus, the concave and flanged subtegular ridge, and the sharp mid-longitudinal carina on the frons, which separates this species from other yellow species with hyaline wings (these species have a strigose or hardly sculptured clypeus, and all of them except E. uelensis have no flange on the subtegular ridge).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C644434BFDCBF9D8FEDDC426.taxon	etymology	Etymology Dedicated to Gavin Broad for his constant support and his most valuable advice.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C644434BFDCBF9D8FEDDC426.taxon	materials_examined	Type material Holotype CAMEROON: ♀, Nkoemvon, 02 ° 48 ’ N 11 ° 08 ’ E, 16 Mar. – 4 May 1980. Ms D. Jackson (BMNH).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C644434BFDCBF9D8FEDDC426.taxon	description	Description Female (holotype) B 15.7; A NA; F 12.1. COLOR. Bright yellow overall with black markings: mandible, mid-apical tubercle on clypeus, vertex, temple, occiput mid-dorsally, mesoscutum but postero-lateral margins and a central macula, and a small mid-posterior macula on tergites 3 – 6; flagellum and ovipositor sheath dark brown; wings hyaline, slightly infuscate apically, venation brown. HEAD. Face rugulose punctate, moderately protruding dorsally; clypeus punctate, with a moderate subapical median tubercle, ventral margin strongly produced laterally; mandible stout; frons smooth; inner margins of toruli expanded backwards into two converging lamellar carinae, carinae then fused into a sharp mid-longitudinal carina reaching median ocellus; frons also with an additional lateral curved carina from posterior margin of torulus to next to lateral ocellus; inter-ocellar area rugose, remainder of vertex and temple finely punctate; occipital carina mid-dorsally evanescent; antenna with more than 35 flagellomeres (apices broken). MESOSOMA. Mesosoma moderately densely punctate, but mesonotum deeply transversely striate and anterior half of pronotum smooth; epomia very weak; epicnemial carina reaching level of mesopleural pit; apex of subtegular ridge concave, strongly flanged laterally; submetapleural carina slightly and evenly narrowed toward apex. WINGS. Fore wing with 2 m – cu slightly apical to rs – m, cu – a slightly apical to Rs & M, and Rs strongly bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a close to junction with M. METASOMA. Tergite 1 moderately slender; tergites 1 – 5 mostly smooth, apex of tergite 5 and following tergites finely and densely punctate; apical margins of tergites 3 – 7 strongly concave. Male Unknown.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C644434BFDCBF9D8FEDDC426.taxon	distribution	Distribution Cameroon.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6594349FDEEFEDDFCB4C678.taxon	description	Fig. 3	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6594349FDEEFEDDFCB4C678.taxon	diagnosis	Diagnosis Bright yellow overall with isolated dorsal black markings; wings hyaline, slightly infuscate apically; face deeply rugulose-punctate, moderately protruding dorsally; clypeus nearly smooth, with median subapical tubercle strong, ventral margin produced laterally; frons laterally punctate and medially rugulose, with two weak submedian carinae diverging below median ocellus toward lateral ocelli; antenna with 34 flagellomeres; mesopleuron with epicnemial carina reaching above level of mesopleural pit; subtegular ridge convex; fore wing with 2 m – cu slightly apical to rs – m; tergite 1 slender; tergites 1 – 2 smooth, following tergites shallowly punctate with basal transverse striations; apical margins of tergites straight. CT 2.7; ML 0.7; POL 1.1; OOL 1.3; Fl 1 4.1; Fl 15 2.2; Fl 33 1.1; T 1 2.5; OT 4.2. Differential diagnosis A mostly bright yellow species with hyaline wings, otherwise characterized by the combination of the pair of submedian carinae on the frons diverging towards the lateral ocelli and the slender tergite 1. The other yellow species with hyaline wings have a stouter tergite 1 in the female, less than twice as long as apically wide.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6594349FDEEFEDDFCB4C678.taxon	materials_examined	Material examined Holotype DEMOCRATIC REPUBLIC OF CONGO: ♀, Eala [00 ° 40 ’ N 18 ° 17 ’ E], Nov. 1935, J. Ghesquière, R. Dét. G 3330 (MRAC). Paratype DEMOCRATIC REPUBLIC OF CONGO: 1 ♂, Bambesa, Dec. 1946, P. L. Benoit (MRAC). Other material CAMEROON: 1 ♂, Korup, Dec. 1980 – Jan. 1981, Mrs D. Jackson (BMNH); 1 ♂, Korup, 1981, Mrs D. Jackson (BMNH).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6594349FDEEFEDDFCB4C678.taxon	description	Description Female (holotype) B 14.6; A 12.2; F 12.1. COLOR. Bright yellow overall, with black markings on mandible, vertex, upper occiput, and mesoscutum margins; antenna and ovipositor sheath dark brown; wings hyaline, apex hardly infuscate, venation pale brown. HEAD. Face subquadrate, deeply rugulose punctate and moderately protruding dorsally; clypeus almost smooth, with a strong subapical median tubercle, ventral margin produced laterally; mandible stout; frons laterally moderately punctate, medially smoothly rugulose; inner margins of toruli expanded backwards into two weak sinuate submedian carinae, carinae then diverging and reaching lateral ocelli; vertex and temple slightly and finely punctate; occipital carina complete but mid-dorsally faint; antenna with 34 flagellomeres. MESOSOMA. Mesosoma shallowly and moderately densely punctate, but mesonotum deeply transversely striate and anterior half of pronotum smooth; epomia very weak; epicnemial carina long, its evanescent dorsal end reaching above level of mesopleural pit; apex of subtegular ridge convex, not flanged laterally; submetapleural carina thick, evenly narrowed toward apex. WINGS. Fore wing with 2 m – cu slightly apical to rs – m, cu – a slightly apical to Rs & M, and Rs weakly bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a close to junction with M. METASOMA. Tergite 1 slender and smooth, tergite 2 mostly smooth with some superficial punctures laterally, tergites 3 – 6 shallowly punctate with basal transverse striations; apical margins of all tergites straight. Male (3 specimens) B 11.8 – 13.6; A 15.1 – 16.2; F 14.9 – 16.0. Similar to female but tergite 1 slenderer (T 1 2.5); the two males from Cameroon are significantly lighter, without dorsal black markings.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6594349FDEEFEDDFCB4C678.taxon	distribution	Distribution Democratic Republic of Congo. New record: Cameroon.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65B434FFDC3FC96FDD6C1F1.taxon	description	Fig. 4	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65B434FFDC3FC96FDD6C1F1.taxon	diagnosis	Diagnosis Bright yellow overall with black markings including entire mesosternum, metasoma fading to brown apically; wings hyaline, slightly infuscate apically; face rugulose-punctate, strongly protruding medially; clypeus nearly smooth, with median and lateral subapical tubercles moderate, ventral margin produced laterally; frons laterally punctate and medially striate, with a blunt Y-shaped mid-longitudinal carina; mesopleuron with epicnemial carina reaching above level of mesopleural pit; subtegular ridge convex; fore wing with 2 m – cu slightly basal to rs – m; tergite 1 of female stout (male unknown); tergites 1 – 3 smooth, following tergites shallowly punctate with basal transverse striations; apical margins of tergites 3 – 4 slightly concave. CT 3.6; ML 0.6; POL 1.6; OOL 1.4; Fl 1 3.4; Fl 15 NA; T 1 1.2 – 1.4; OT 4.4. Differential diagnosis A mostly bright yellow species with hyaline wings, otherwise characterized by the combination of the black mesosternum, vein 2 m – cu slightly basal to rs – m and the stout tergite 1 (at least in the female). All other yellow Afrotropical Epirhyssa species have the mesosternum concolorous with the remainder of the mesosoma and the vein 2 m – cu slightly apical to rs – m.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65B434FFDC3FC96FDD6C1F1.taxon	materials_examined	Material examined Holotype DEMOCRATIC REPUBLIC OF CONGO: ♀, Bambesa [03 ° 28 ’ N 25 ° 43 ’ E], Nov. 1933, J. V. Leroy (MRAC). Paratype DEMOCRATIC REPUBLIC OF CONGO: ♀, Ubangui-Bumba, Dec. 1939, H. de Saeger (MRAC).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65B434FFDC3FC96FDD6C1F1.taxon	description	Description Female (holotype) B 14.9; A NA; F 11.1. COLOR. Bright to dull yellow overall, with metasoma fading to brown toward apex, and with black markings: mandible, ventral margin of clypeus, ocellar area (macula connected to eye margin), occiput mid-dorsally, mesosternum, and a median macula on each mesoscutal lobe; antenna and ovipositor sheath dark brown; wings hyaline, slightly infuscate apically, venation pale brown. HEAD. Face rugulose-punctate, strongly bulging medially; clypeus smoothly sculptured with distinct lateral and median subapical tubercles, ventral margin strongly produced laterally; mandible short and stout; frons laterally sparsely punctate, medially transversely striate with a blunt Y-shaped midlongitudinal carina below median ocellus, inner margins of toruli not expanded backwards; temple and vertex finely punctate, inter-ocellar area somewhat rugose; occipital carina mid-dorsally interrupted; antenna broken in both specimens. MESOSOMA. Mesosoma shallowly and moderately densely punctate, but mesonotum deeply transversely striate and anterior half of pronotum smooth; epomia moderate; epicnemial carina long, its evanescent dorsal end reaching above level of mesopleural pit; apex of subtegular ridge convex, not flanged laterally; submetapleural carina anteriorly expanded, then distinctly narrowed toward apex. WINGS. Fore wing with 2 m – cu slightly basal to rs – m, cu – a slightly apical to Rs & M, and Rs weakly bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a close to junction with M. METASOMA. Tergite 1 stout and smooth, tergites 2 – 3 nearly smooth, following tergites finely and distinctly punctate with weak basal striations; apical margins of tergites 3 – 4 slightly concave, others straight. Male Unknown.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65B434FFDC3FC96FDD6C1F1.taxon	distribution	Distribution Democratic Republic of Congo.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65D434DFDF5FB0DFBE1C3FD.taxon	description	Fig. 5	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65D434DFDF5FB0DFBE1C3FD.taxon	diagnosis	Diagnosis Bright yellowish-orange overall with dark dorsal markings including apical half of metasoma; wings yellowish with apices strongly infuscate and an additional infuscate transverse subapical stripe on fore wing; face sparsely punctate, barely protruding; clypeus nearly smooth, without subapical tubercle, ventral margin truncate; frons smooth with a blunt Y-shaped mid-longitudinal carina; antenna with 37 flagellomeres; epicnemial carina hardly extending onto mesopleuron; subtegular ridge convex; fore wing with 2 m – cu opposite rs – m; tergite 1 stout; all tergites smooth, with fine isolated punctures; apical margins of tergites 3 – 5 strongly concave. CT 2.9; ML 0.5; POL 1.3; OOL 1.3; Fl 1 3.7; Fl 15 2.5; Fl 36 1.2; T 1 1.2. Differential diagnosis Species characterized by its coloration, with apical half of metasoma black, wings yellowish with black patterns; otherwise characterized by the combination of the sparsely punctate face, the absence of subapical protuberances on the clypeus, the ventral margin of the clypeus not produced laterally, and the epicnemial carina nearly absent laterally. It is the only yellow Afrotropical species with banded wings.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65D434DFDF5FB0DFBE1C3FD.taxon	materials_examined	Material examined Holotype DEMOCRATIC REPUBLIC OF CONGO: ♂, Albertville [Kalemie, 05 ° 56 ’ N 29 ° 12 ’ E], Jul. 1918, R. Mayné (MRAC). Paratype DEMOCRATIC REPUBLIC OF CONGO: 1 ♂, Bambesa, Dec. 1946, P. L. Benoit (MRAC). Other material CENTRAL AFRICAN REPUBLIC: 1 ♂, Préfecture Sangha-Mbaéré, Parc National de Dzanga-Ndoki, 38.6 km, 173 ° S of Lidjombo, 2 ° 21.60 ’ N 16 ° 09.20 ’ E, 350 m, 27 May 2001, S. van Noort, handnet, Lowland Rainforest, CAR 01 - H 25, SAM – HYM – P 049437 (SAMC).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65D434DFDF5FB0DFBE1C3FD.taxon	description	Description Female Unknown. Male (3 specimens) B 14.2 – 16.7; A 13.6 – 14.9; F 11.5 – 13.3 (holotype: B 16.7; A 14.9; F 13.3). COLOR. Bright yellowish orange overall with dark brown to black markings: apical 2 / 3 of mandible, frons, vertex, temple, hind leg from femur and metasoma from tergite 4; flagellum dark brown; wings yellowish, apically strongly infuscate, fore wing with an additional transverse infuscate stripe at pterostigma level, venation yellow. HEAD. Face sub-quadrate, sparsely punctate, barely protruding dorsally but with a short mid-longitudinal ridge near toruli; clypeus mostly smooth with some punctures along dorsal margin, without distinct subapical tubercle, ventral margin truncate; mandible stout, both teeth truncate and chisel-like; upper head quite smooth; frons with a blunt Y-shaped mid-longitudinal carina below median ocellus, inner margins of toruli not expanded backwards; occipital carina mid-dorsally evanescent; antenna with 34 – 37 flagellomeres. MESOSOMA. Mesosoma shallowly and moderately densely punctate, but mesonotum deeply transversely striate and anterior half of pronotum smooth; epomia very weak; epicnemial carina very short, hardly extending laterally onto mesopleuron; apex of subtegular ridge convex, not flanged laterally; submetapleural carina anteriorly moderately expanded, then abruptly narrowed from anterior third. WINGS. Fore wing with 2 m – cu opposite rs – m, cu – a distinctly apical to Rs & M, Rs moderately bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a at its anterior third, remote from junction with M. METASOMA. Tergite 1 slender; all tergites smooth but with fine isolated punctures; apical margins of tergites 3 – 5 strongly concave.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65D434DFDF5FB0DFBE1C3FD.taxon	distribution	Distribution Democratic Republic of Congo. New record: Central African Republic.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65F4343FDE0F911FEB3C2CE.taxon	description	Fig. 6	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65F4343FDE0F911FEB3C2CE.taxon	diagnosis	Diagnosis Bright yellow overall with isolated dorsal black markings; wings hyaline, slightly infuscate apically; face sparsely punctate, almost flat; clypeus nearly smooth, with median and lateral subapical tubercles moderate, ventral margin produced laterally; frons smooth, with two submedian carinae diverging below median ocellus toward lateral ocelli; antenna with 36 flagellomeres; epicnemial carina laterally absent; subtegular ridge convex; fore wing with 2 m – cu slightly apical to rs – m; tergite 1 weakly slender in female, much more slender in male; tergites 1 – 3 smooth, following tergites sparsely punctate with basal transverse striations; apical margins of tergites 3 – 5 moderately concave. CT 2.5; ML 0.6; POL 2.6; OOL 1.3; Fl 1 4.4; Fl 15 2.3; Fl 35 1.1; T 1 1.4 – 1.5; OT 5.0. Differential diagnosis A mostly bright yellow species with hyaline wings, otherwise characterized by the combination of the flat and sparsely punctate face, the pair of submedian carinae on the frons and the absence of the epicnemial carina on the mesopleuron. The lateral part of the epicnemial carina is distinct in every other yellow species with hyaline wings.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65F4343FDE0F911FEB3C2CE.taxon	materials_examined	Material examined Lectotype MADAGASCAR: ♀, Rogez [48 ° 18 ’ S 48 ° 32 ’ E], Sep. 30, A. Seyrig, MNHN EY 8815. Paratypes 3 ♂♂, same label data.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65F4343FDE0F911FEB3C2CE.taxon	description	Description Female (holotype) B 18.2; A 15.1; L 14.7. COLOR. Bright yellow overall with infuscate to black markings: mandible, inter-ocellar area (macula connected to eye margin), dorsal margin of occiput, mesoscutal lobes mid-longitudinally, and scutoscutellar groove; antenna and ovipositor sheath mostly dark brown; wings nearly hyaline, apically slightly infuscate, venation testaceous. HEAD. Face sparsely punctate, almost flat; clypeus smoothly sculptured with moderate lateral and median subapical tubercles, ventral margin moderately produced laterally; mandible stout; upper head quite smooth to moderately and shallowly punctate, inter-ocellar area somewhat rugose; frons with a pair of submedian carinae, carinae abruptly diverging below median ocellus and following anterior margin of vertex, inner margins of toruli hardly produced backwards; occipital carina mid-dorsally interrupted; antenna with 36 flagellomeres. MESOSOMA. Mesosoma shallowly and moderately densely punctate, but mesonotum transversely striate and anterior half of pronotum smooth; epomia weak; epicnemial carina absent except for a short ventral remnant; apex of subtegular ridge convex, not flanged laterally; submetapleural carina strong and even all along. WINGS. Fore wing with 2 m – cu slightly apical to rs – m, cu – a slightly apical to Rs & M, and Rs moderately bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a close to junction with M. METASOMA. Tergite 1 moderately slender; tergites 1 – 3 smooth, following tergites finely and sparsely punctate with basal transverse striations; apical margins of tergites 3 – 5 moderately concave, others straight. Male (3 specimens) B 17.5 – 18.2; A 14.3 – 14.9; L 14.0 – 14.5. Similar to female but metasoma much more slender (T 1 2.0 – 2.2).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C65F4343FDE0F911FEB3C2CE.taxon	distribution	Distribution Madagascar.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6504340FDFCFEACFB5BC739.taxon	description	Fig. 7	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6504340FDFCFEACFB5BC739.taxon	diagnosis	Diagnosis Overall color mottled, black, yellow and testaceous; wings hyaline, slightly infuscate apically; face transversely striate, nearly flat; clypeus shallowly and sparsely punctate, without subapical tubercle, ventral margin strongly produced laterally; frons smooth, with a sharp mid-longitudinal carina reaching median ocellus and lateral curved carina toward lateral ocellus; antenna with 41 – 43 flagellomeres; epicnemial carina reaching lower level of speculum; fore wing with 2 m – cu opposite rs – m; tergite 1 slender; all tergites nearly to quite smooth, tergites 4 – 5 finely transversely striate basally; apical margins of tergites 3 – 6 strongly concave. CT 2.5; ML 0.5; POL 1.3; OOL 1.6; Fl 1 3.9; Fl 15 2.8; Fl 42 1.3; T 1 1.7; OT 7.2. Differential diagnosis Highly distinctive species of mottled color and large to very large size; otherwise characterized by the combination of the flat and sparsely punctate face, the sharp mid-longitudinal carina on the frons, the absence of a subapical ridge on the clypeus, and the very long ovipositor. No other known Epirhyssa species shares this color pattern.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6504340FDFCFEACFB5BC739.taxon	materials_examined	Material examined Holotype DEMOCRATIC REPUBLIC OF CONGO: ♀, Lulua [10 ° 37 ’ S 24 ° 54 ’ E], Kapanga, Apr. 1933, F. G. Overlaet, R. Dét. F 3330 (MRAC). Other material CAMEROON: 1 ♀, Nkoemvon, Jul. - Aug. 1980, Ms. D. Jackson (BMNH).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6504340FDFCFEACFB5BC739.taxon	description	Description Female (2 specimens) B 25.2 – 37.8; A 15.4 – 23.2; F 16.1 – 24.2. COLOR. Head yellow with mandible black and lower gena, vertex and upper occiput brownish testaceous; mesosoma black to dark brownish testaceous interspersed with yellow markings: pronotum dorsally and ventrally, mesopleuron anteriorly and posteriorly, and mesoscutum, scutellum, metanotum and propodeum centrally; legs testaceous to dark testaceous with fore and mid coxa yellow; metasoma light to dark testaceous, tergites 1 – 5 with apico-lateral large yellow spots; antenna and ovipositor sheath dark brown; wings nearly hyaline, apically infuscate, venation dark brown. HEAD. Face subquadrate, transversely striate and hardly bulging medially; clypeus shallowly and sparsely punctate, without subapical tubercle, ventral margin strongly produced laterally; mandible stout; upper head quite smooth; inner margins of toruli expanded backwards into two converging lamellar carinae, carinae then fused into a sharp mid-longitudinal carina reaching median ocellus; frons also with an additional lateral curved carina from posterior margin of torulus to next to lateral ocellus; occipital carina mid-dorsally interrupted; antenna with 41 – 43 flagellomeres. MESOSOMA. Mesosoma shallowly and moderately densely punctate, but mesonotum transversely striate and anterior half of pronotum smooth; epomia indistinct; epicnemial carina reaching lower level of speculum, mesopleural pit indistinct; apex of subtegular ridge convex, not flanged laterally; submetapleural carina moderate and even all along. WINGS. Fore wing with 2 m – cu opposite rs – m, cu – a slightly apical to Rs & M, and Rs short and moderately bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a at junction with M. METASOMA. Tergite 1 moderately slender; all tergites smooth to indistinctly sculptured but tergites 4 – 5 weakly striate basally; apical margins of tergites 3 – 6 strongly concave, others straight. Male Unknown.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6504340FDFCFEACFB5BC739.taxon	distribution	Distribution Democratic Republic of Congo. New record: Cameroon.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6504340FDFCFEACFB5BC739.taxon	discussion	Comments The species is only known from two female specimens. They exhibit a remarkable variation in size, the BMNH specimen being one third smaller than the impressively large MRAC holotype.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6524346FD2EFDD5FDCFC6C4.taxon	description	urn: lsid: zoobank. org: act: CF 997 FDF-A 6 A 7 - 49 A 1 - 9 DCA-C 5 F 781 CACEE 1 Fig. 8	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6524346FD2EFDD5FDCFC6C4.taxon	diagnosis	Diagnosis Pale yellow overall with isolated dorsal black markings; wings hyaline, slightly infuscate apically; face rugulose-punctate, weakly bulging; clypeus shallowly longitudinally strigose with a strong subapical median tubercle, ventral margin produced laterally; frons smooth, with a weak mid-longitudinal carina; antenna with 32 flagellomeres; mesopleuron with epicnemial carina reaching level of mesopleural pit; subtegular ridge convex; fore wing with 2 m – cu slightly apical to rs – m; tergite 1 slender; tergite 1 smooth, following tergites densely punctate; apical margins of all tergites straight. CT 3.8; ML 0.7; POL 1.3; OOL 1.2; Fl 1 4.5; Fl 15 1.8; Fl 31 1.0; T 1 1.7; OT 3.9. Differential diagnosis Pale yellow species of relatively small size, otherwise characterized by the combination of the weak mid-longitudinal carina on the frons, the entirely punctate tergite 2 and the slender tergite 1.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6524346FD2EFDD5FDCFC6C4.taxon	etymology	Etymology Refers to King Shaka Zulu, another kind of population regulator of the Zululand region. He would certainly have been proud of giving his name to an ichneumon.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6524346FD2EFDD5FDCFC6C4.taxon	materials_examined	Type material Holotype SOUTH AFRICA: ♀, Natal. 1.5 km E of Mtunzini, 28 ° 57 ’ S 31 ° 45 ’ E, 2831 Dd Umlalazi Nat. Res., Nov. 1978, R. M. Miller, indigenous forest, Malaise tr. (NMSA).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6524346FD2EFDD5FDCFC6C4.taxon	description	Description Female (holotype) B 11.9; A 9.6; F 8.7. COLOR. Pale dull yellow overall with black parts: mandible, ocellar area (macula isolated from eye margin), a median spot on each mesoscutal lobe, scuto-scutellar groove, a mid-longitudinal triangle at base of tergites 4 – 6; flagellum dark brown; hind tarsus and ovipositor sheath reddish-brown; wings hyaline, barely infuscate apically, venation yellowish. HEAD. Face subquadrate, weakly bulging mid-dorsally, rugulose punctate and laterally smoother; clypeus shallowly longitudinally strigose, with a strong and acute subapical median tubercle, ventral margin laterally produced; mandible stout; frons smooth; inner margins of toruli expanded backwards into two converging carinae, carinae then fused into a weak mid-longitudinal carina reaching median ocellus; vertex and temple moderately punctate; occipital carina complete; antenna with 32 flagellomeres. MESOSOMA. Mesosoma shallowly and moderately densely punctate, but mesonotum deeply transversely striate and anterior half of pronotum smooth; epomia moderate; dorsal faint apex of epicnemial carina reaching level of mesopleural pit; apex of subtegular ridge convex, not flanged laterally; submetapleural carina slightly and evenly narrowed toward apex. WINGS. Fore wing with 2 m – cu slightly apical to rs – m, cu – a slightly apical to Rs + M, Rs moderately bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a close to junction with M. METASOMA. Tergite 1 moderately slender, sparsely and shallowly punctate; tergite 2 and following densely punctate; apical margins of all tergites straight. Some other yellow species also sometimes have some punctures on tergite 2 (E. tombeaodiba), but the first tergite of the female is then distinctively stouter. Male Unknown.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6524346FD2EFDD5FDCFC6C4.taxon	distribution	Distribution South Africa (KwaZulu-Natal).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6544344FDF7FC7BFEDDC185.taxon	description	urn: lsid: zoobank. org: act: 134 C 55 D 0 - CBC 2 - 4788 - B 199 - 2197 C 7344475 Fig. 9	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6544344FDF7FC7BFEDDC185.taxon	diagnosis	Diagnosis Bright yellow overall with isolated dorsal black markings; wings hyaline, slightly infuscate apically; face rugulose-punctate, distinctly protruding dorsally; clypeus shallowly longitudinally strigose with a strong subapical median tubercle, ventral margin hardly produced laterally; frons smooth, with a weak mid-longitudinal carina reaching median ocellus; mesopleuron with epicnemial carina reaching level of mesopleural pit; subtegular ridge convex; fore wing with 2 m – cu distinctly apical to rs – m; tergite 1 stout in female (male unknown); tergite 1 about smooth, following tergites more or less densely punctate; apical margins of all tergites straight. CT 2.8; ML 0.7; POL 1.3; OOL 1.3; Fl 1 4.0; Fl 15 2.2; T 1 1.2; OT 4.0. Differential diagnosis Bright yellow species with wings hyaline, otherwise characterized by the combination of the entirely punctate tergite 2, the weak mid-longitudinal carina on the frons and the stout tergite 1. It is the only known yellow Epirhyssa species with both a distinctly sculptured clypeus and a stout tergite 1.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6544344FDF7FC7BFEDDC185.taxon	etymology	Etymology Refers to Mrs Jackson’s ambiguous marital status as reported by label data. In the Duala language, “ tómbea ó dibá ” means “ to get married ” for a woman.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6544344FDF7FC7BFEDDC185.taxon	materials_examined	Type material Holotype CAMEROON: ♀, Nkoemvon [02 ° 48 ’ N 11 ° 08 ’ E], 16 Mar. – 4 May 1980, Ms. D. Jackson (BMNH). Paratypes 1 ♀, same label data (BMNH); 1 ♀, same label data except: 30 Mar. – 19 Apr. 1980 (BMNH); 1 ♀, same label data except: Korup 1981, Mrs D. Jackson (BMNH).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6544344FDF7FC7BFEDDC185.taxon	description	Description Female (4 specimens) B 7.2 – 8.9; A 9.5; F 9.0 – 10.5. (holotype B: 8.0; A NA; F 9.9). COLOR. Bright yellow overall with black parts: mandible, ocellar area (macula connected to eye margin), a median spot on each mesoscutal lobe, scuto-scutellar groove, a mid-longitudinal macula of variable length on tergites 4 – 7; flagellum dark brown; flagellum and ovipositor sheath reddish-brown; wings hyaline, slightly infuscate apically, venation brown. HEAD. Face subquadrate, rugulose-punctate, distinctly protruding dorsally; clypeus longitudinally strigose, with a strong subapical median lobe, ventral margin hardly produced laterally; mandible stout; frons, vertex and temple sparsely and shallowly punctate; inner margins of toruli expanded backwards into two converging weak carinae, carinae then fused into a mid-longitudinal carina reaching median ocellus; occipital carina mid-dorsally interrupted; antenna with at least 22 flagellomeres (no specimen with complete flagellum). MESOSOMA. Mesosoma shallowly and moderately densely punctate, but mesonotum deeply transversely striate and anterior half of pronotum smooth; epicnemial carina reaching level of mesopleural pit; apex of subtegular ridge convex, not flanged laterally; submetapleural carina slightly enlarged anteriorly, then evenly narrowed toward apex, strongly notched at anterior third in one paratype. WINGS. Fore wing with 2 m – cu distinctly apical to rs – m, cu – a barely apical to Rs & M, and Rs moderately bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a shortly below junction with M. METASOMA. Tergite 1 moderately stout, nearly smooth; following tergites shallowly and more or less densely punctate, punctation progressively finer toward apex; apical margins of all tergites straight.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C6544344FDF7FC7BFEDDC185.taxon	distribution	Distribution Cameroon.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C656437AFDF2FAB9FCB4C386.taxon	description	Fig. 10	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C656437AFDF2FAB9FCB4C386.taxon	diagnosis	Diagnosis Bright yellow overall with isolated dorsal black markings; wings hyaline, slightly infuscate apically; face rugulose-punctate and moderately protruding dorsally; clypeus nearly smooth, with median subapical tubercle strong, ventral margin produced laterally; frons nearly smooth, with a mid-longitudinal carina reaching median ocellus and lateral curved carina toward lateral ocellus; antenna with 32 – 34 flagellomeres; mesopleuron with epicnemial carina reaching level of mesopleural pit; subtegular ridge concave, laterally flanged; fore wing with 2 m – cu slightly apical to rs – m; tergite 1 stout in female, very slender in male; tergites 1 – 3 smooth, following tergites shallowly punctate with basal transverse striations; apical margins of tergites 3 – 6 moderately concave. CT 2.9; ML 0.6; POL 1.2; OOL 1.3; Fl 1 3.8; Fl 15 2.2; FL 33 0.9; T 1 1.2 – 1.3; OT 4.1. Differential diagnosis A mostly bright yellow species with hyaline wings, otherwise characterized by the combination of the concave subtegular ridge, the mid-longitudinal carina on the frons and the stout tergite 1 of the female. The only other species with a concave and flanged subtegular ridge is E. gavinbroadi, in which the clypeus is distinctly punctate.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C656437AFDF2FAB9FCB4C386.taxon	materials_examined	Material examined Holotype DEMOCRATIC REPUBLIC OF CONGO: ♀, Haut-Uele, Paulis [Isiro, 03 ° 28 ’ N 25 ° 43 ’ E], xii- 1947, P. L. G. Benoit (MRAC). Paratype DEMOCRATIC REPUBLIC OF CONGO: 1 ♂, Bambesa, Jul. 1933, H. J. Bredo (MRAC). Other material CAMEROON: 1 ♀, Nkoemvon, 30 Mar. – 19 Apr. 1980, Ms. D. Jackson (BMNH); 2 ♀♀, same label data except 13 Jul. – 4 Aug. 1980 (BMNH); 1 ♀, same label data except Oct. – Nov. 1980 (BMNH).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C656437AFDF2FAB9FCB4C386.taxon	description	Description Female (5 specimens) B 10.4 – 12.8; A 9.1 – 12.0; F 8.6 – 11.5 (holotype B: 10.4; A 9.1; F 8.6). COLOR. Bright yellow overall, sometimes with infuscate to black mid-longitudinal stripes on tergites 5 – 7, and black markings: mandible, ocellar area (macula connected to eye margin), upper occiput and mesoscutum but lateral margins and a large central macula, sometimes apex of scutellum, basal margin and apex of propodeum; ovipositor sheath reddish testaceous; wings hyaline, weakly infuscate apically, venation brown. HEAD. Face rugulose-punctate and moderately protruding dorsally; clypeus smoothly sculptured, with a strong subapical median tubercle, ventral margin moderately produced laterally and medially; mandible stout; frons nearly smooth with sparse fine punctures; inner margins of toruli expanded backwards into two converging weak carinae, carinae then fused into a mid-longitudinal carina reaching median ocellus; frons also with an additional lateral curved carina from posterior margin of torulus to next to lateral ocellus; vertex and temples sparsely punctate, inter-ocellar area somewhat rugose; occipital carina mid-dorsally evanescent; hypostomal carina strongly raised above mandibular base; antenna with 32 – 34 flagellomeres. MESOSOMA. Mesosoma shallowly and moderately densely punctate, but mesonotum deeply transversely striate and anterior half of pronotum smooth; epomia moderate; dorsal faint apex of epicnemial carina nearly reaching level of mesopleural pit; apex of subtegular ridge concave, weakly to distinctly flanged laterally; submetapleural carina slightly and evenly narrowed toward apex. WINGS. Fore wing with 2 m – cu slightly apical to rs – m, cu – a slightly apical to Rs + M, Rs strongly bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a close to junction with M. METASOMA. Tergite 1 stout and smooth, tergites 2 – 3 smooth, tergites 3 – 6 shallowly punctate with weak transverse striations basally; apical margins of tergites 3 – 6 moderately concave, others straight. Male (1 specimen) B 8.5; A 8.2; F 7.9. Similar to female but metasoma much slenderer (T 1 2.0).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C656437AFDF2FAB9FCB4C386.taxon	distribution	Distribution Democratic Republic of Congo. New record: Cameroon.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66B437FFDC8FEACFEFFC7D7.taxon	description	urn: lsid: zoobank. org: act: C 701 D 367 - 2 EF 0 - 4 A 73 - 8 D 41 - F 8 F 5 EDADAA 21 Fig. 11	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66B437FFDC8FEACFEFFC7D7.taxon	diagnosis	Diagnosis Head and pronotum yellowish-orange; remainder of mesosoma black; metasoma orange; wings blackish with a pale subapical patch; face sparsely punctate and moderately protruding dorsally; clypeus smooth, with median subapical tubercle moderate, ventral margin truncate; frons laterally smooth, medially transversely striate, with two submedian carinae diverging toward lateral ocelli; antenna with 36 flagellomeres; mesopleuron with epicnemial carina reaching level of mesopleural pit; subtegular ridge convex; fore wing with 2 m – cu slightly apical to rs – m; tergite 1 stout in female (male unknown); tergites 1 – 3 smooth, following tergites moderately punctate with basal transverse striations; apical margins of all tergites straight. CT 2.0; ML 0.6; POL 1.3; OOL 1.6; Fl 1 3.6; Fl 15 2.1; FL 33 0.9; T 1 1.3; OT 4.4. Differential diagnosis A species with unique color pattern, with most of mesosoma black, metasoma orange and wings blackish with a subapical light patch; otherwise characterized by the sparsely punctate face, the truncate ventral margin of the clypeus and the stout tergite 1 of the female.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66B437FFDC8FEACFEFFC7D7.taxon	etymology	Etymology Dedicated to Claire Villemant, MNHN curator, hymenopterologist and ichneumonomaniac, adviser, teacher, supporter, coach and friend.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66B437FFDC8FEACFEFFC7D7.taxon	materials_examined	Type material Holotype NIGERIA: ♀, Ilorin Prov. [06 ° 48 ’ N 05 ° 18 ’ E], 18 Jun. 192 [sic], De. G. W. S. Macfie, Pres. by Imp. Bur. Ent. 1921 – 129 (BMNH).	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66B437FFDC8FEACFEFFC7D7.taxon	description	Description Female (holotype) B 17.1; A 13.1; F 13.1. COLOR. Head and pronotum yellowish-orange with mandible and ocellar area black; remainder of mesosoma blackish testaceous; metasoma orange; fore leg and mid leg from tibia yellowish orange, remainder of legs blackish testaceous; antenna and ovipositor sheath brown; wings strongly infuscate but with large subapical and small mid-dorsal pale yellowish patches on fore wing. HEAD. Face sparsely punctate, almost smooth laterally, and moderately protruding dorsally; clypeus smooth, with a moderate subapical median tubercle, ventral margin truncate; mandible stout; frons laterally smooth, medially transversely striate; inner margin of toruli expanded backwards into two weak sinuate submedian carinae, carinae then diverging and reaching lateral ocelli; frons also with an additional lateral curved carina from posterior margin of torulus to next to lateral ocellus; vertex and temples almost smooth, inter-ocellar area somewhat rugose; occipital carina mid-dorsally evanescent; hypostomal carina strongly raised above mandibular base; antenna with 36 flagellomeres. MESOSOMA. Mesosoma finely, sparsely and shallowly punctate, but mesonotum transversely striate and pronotum, speculum area and propodeum medially smooth; epomia very weak; dorsal faint apex of epicnemial carina reaching level of mesopleural pit; apex of subtegular ridge convex, not flanged laterally; submetapleural carina moderate and even all along. WINGS. Fore wing with 2 m – cu slightly apical to rs – m, cu – a slightly apical to Rs + M, Rs moderately bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a at its anterior third, remote from junction with M. METASOMA. Tergite 1 moderately stout; tergites 1 – 3 smooth, following tergites moderately punctate with weak transverse basal striations; apical margins of all tergites straight. Male Unknown.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66B437FFDC8FEACFEFFC7D7.taxon	distribution	Distribution Nigeria.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66D437FFDECFD6AFADBC087.taxon	diagnosis	Diagnosis (updated from Townes 1969) Large to very large insects (fore wing length 10 – 30 mm); clypeus small, transversely rectangular, ventral margin laterally and sometimes medially bluntly produced; occipital and hypostomal carinae joining above mandible base; mandibular teeth subequal, lower tooth pointed and upper tooth more or less chisel shaped; propodeum of moderate length, without carina dorsally but pleural carina distinct; fore wing with areolet closed except in occasional dwarf males, receiving 2 m – cu within its apical half, pterostigma about 5.5 x longer than wide; hind wing with distal abscissa of Cu present, joining Cu & cu – a near or at junction with M; mid trochantellus with a ventral longitudinal ridge; tarsal claws simple, large; tergite 1 fused with its sternite and without glymma; tergites 3 – 6 almost smooth to finely and sparsely punctate and with isolated aciculate areas; female with sternites 2 – 4 each with a pair of tubercles near anterior margin, and with an apical truncate horn-like process on last tergite; male with metasoma strongly depressed, gonosquama lanceolate, strongly depressed, with a sharp piliferous groove along inner lower margin and a short subapical piliferous groove on outer face.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66D437DFDE7F9BBFDD6C32B.taxon	description	Fig. 12	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66D437DFDE7F9BBFDD6C32B.taxon	diagnosis	Diagnosis Head orange with a black part of variable extent; mesosoma orange, posterior third sometimes dark brown; metasoma mostly black with pale yellow or orange markings of variable extent; flagellum twocolored, basally black and apically orange; wings blackish with lighter patch of variable extent; face moderately punctate to smooth, flat; clypeus smooth, with median subapical tubercle weak, ventral margin strongly produced laterally; frons smooth with a pair of submedian carinae diverging toward lateral ocelli; epicnemial carina laterally absent; subtegular ridge convex; fore wing with areolet strongly petiolate, receiving 2 m – cu at its distal apex, opposite 3 rs – m; tergite 1 very stout; tergites 1 – 4 smooth or mostly so, following tergites finely and densely punctate; apical margins of tergites 3 – 5 strongly concave. CT 3.4; ML 0.6; POL 1.2; OOL 1.7; Fl 1 4.4; Fl 15 2.3; Fl 39 1.0; T 1 1.1; OT 4.8.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66D437DFDE7F9BBFDD6C32B.taxon	materials_examined	Material examined Holotype DEMOCRATIC REPUBLIC OF CONGO: ♀, reg. Lac Kivu, Kadjudju [Kajuju, 02 ° 09 ’ S 28 ° 54 ’ E], 1932, G. Babault, MNHN EY 8831.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66D437DFDE7F9BBFDD6C32B.taxon	description	Description Female (holotype) B 22.1; A 15.5; F 18. COLOR. Head orange with isolated black parts to almost entirely black; antenna basally black, fading to orange from flagellomere 12; mesosoma orange, posterior third varying from orange to entirely black; legs orange with mid and fore legs variably orange to dark brown; metasoma black with tergites 1 – 3 varying from orange to black, tergite 4 and following largely pale yellow laterally, sternites pale yellow; ovipositor sheath black, apical half fading to reddish-orange; wings blackish with lighter yellowish parts of variable extension (color variation: Seyrig 1937; Benoit 1951). HEAD. Face flat, laterally smooth, medially moderately punctate; clypeus smooth, with a weak median subapical tubercle, ventral margin strongly produced laterally; upper head almost smooth; frons with a pair of weak submedian carinae, carinae diverging below median ocellus, inner margins of toruli hardly produced backwards; occipital carina complete, mid-dorsally weak; antenna with 40 flagellomeres. MESOSOMA. Mesosoma finely and moderately densely punctate, but mesonotum deeply transversely striate and anterior half of pronotum smooth; epomia indistinct; epicnemial carina absent except for a faint ventral remnant; apex of subtegular ridge convex, not flanged laterally; submetapleural carina barely expanded anteriorly and evenly narrowed toward apex. WINGS. Fore wing with areolet strongly petiolate, receiving 2 m – cu at its apex, cu – a strongly apical to Rs & M, Rs strongly bowed forwards; hind wing with distal abscissa of Cu joining Cu & cu – a shortly above middle of Cu & cu – a. METASOMA. Tergite 1 stout, nearly as long as apically wide; tergites 1 – 3 quite smooth, tergite 4 smooth with some fine punctures basally, following tergites finely and densely punctate; apical margins of tergites 3 – 5 strongly concave. Male Unknown.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
1509C719C66D437DFDE7F9BBFDD6C32B.taxon	distribution	Distribution Democratic Republic of Congo.	en	Rousse, Pascal, Noort, Simon Van (2014): A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91 (91): 1-42, DOI: 10.5852/ejt.2014.91
