identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
BDC8EB8A7B6E5C4F914A3DE028DCE6E9.text	BDC8EB8A7B6E5C4F914A3DE028DCE6E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Retiboletus ater Yan C. Li & T. Bau 2020	<div><p>Retiboletus ater Yan C. Li &amp; T. Bau sp. nov. Figures 2a-c, 3</p><p>Etymology.</p><p>ater referring to the color of the basidiomata.</p><p>Type.</p><p>China. Yunnan Province: Jingdong County, Ailaoshan National Nature Reserve, alt. 2500 m, 14 July 2008, Y.C. Li 1215 (holotype: KUN-HKAS 56069!).</p><p>Description.</p><p>Basidiomata small to medium-sized. Pileus 3-5 cm in diameter, hemispherical to applanate, surface dry, densely subtomentose, black (4F3) to blackish (4E2) in the center and gray (3D1) or yellowish-gray (3C2-3) towards margin, context 2.5 cm thick in the center of the pileus, pallid gray (2D1) to cream (2C3-4), unchanging when bruised. Hymenophore adnate or slightly depressed around apex of stipe; pores angular, tubes up to 11 mm long, 0.3-1 mm wide, white (2B1) when young and yellowish (2A2) in age, becoming brownish-yellow (5C7-8) when injured. Stipe 4-6 × 0.8-1.2 cm, clavate to flexuous, solid; surface dry, blackish to gray, prominently and coarsely reticulate over the upper 1/3; context white (2A1) in the upper part and yellowish to cream yellow downwards, unchanging when injured; basal mycelium white (2A1). Taste and odor indistinct.</p><p>Basidia 26-38 × 6-10 μm, clavate, thin-walled, 4-spored, hyaline to yellowish in KOH. Basidiospores [60/3/2] (7)8-10.5(11) × 3-4.5(5) μm [Q = (1.89) 2-3.33 (3.67), Qm = 2.52 ± 0.42], subfusiform and inequilateral in side view with shallow suprahilar depression, elongate fusoid or narrowly oblong in ventral view, slightly thick-walled (up to 0.5 μm), brownish to yellowish-brown in KOH, olive-brown to brown in Melzer’s reagent, smooth. Hymenophoral trama boletoid; hyphae cylindrical, 3.5-9 μm wide, hyaline to yellowish in KOH, yellowish to brownish-yellow in Melzer’s reagent. Cheilo- and pleurocystidia 26-55 × 6-10 μm, abundant, subfusiform to fusiform, thin-walled, with yellowish-brown contents, surface without encrustations. Caulocystidia forming the reticulum over the stipe surface, similar to cheilo- and pleurocystidia. Pileipellis a trichoderm about 280 μm thick, composed of more or less vertically arranged, slightly interwoven, brown to dark brown hyphae, 5-15 μm wide; terminal cells 45-111 × 9-15 μm, narrowly clavate to subcylindrical or subfusiform, sometimes narrowly mucronate, rostrate, slightly thick-walled (up to 0.5 μm), hyaline to yellowish in KOH, yellowish to brownish-yellow in Melzer’s reagent. Pileal trama composed of thin- to slightly thick-walled (up to 0.5 μm) hyphae, 5-11 μm wide, hyaline to yellowish in KOH, yellowish to brownish-yellow in Melzer’s reagent. Clamp connections absent in all tissues.</p><p>Habitat, ecology and distribution.</p><p>Solitary on the ground in forests dominated by plants in the family Fagaceae; currently known from southwestern China.</p><p>Additional specimens examined.</p><p>China. Yunnan Province: Jingdong County, Ailaoshan National Nature Reserve, alt. 2500 m, 14 July 2008, Y.C. Li 1224 (KUN-HKAS 56078).</p><p>Discussion.</p><p>Retiboletus ater is characterized by the black to blackish or gray to yellowish-gray pileus, the white to yellowish hymenophore, the gray to brownish-gray stipe, the prominent and coarse reticulum over the upper 1/3 of the stipe and the trichoderm pileipellis with hyphae 9-15 μm wide. It generally shares the same colored pileus and hymenophore with R. fuscus (Hongo) N.K. Zeng &amp; Zhu L. Yang, R. griseus (Frost) Manfr. Binder &amp; Bresinsky, R. nigrogriseus N.K. Zeng, S. Jiang &amp; Zhi Q. Liang, and R. pseudogriseus N.K. Zeng &amp; Zhu L. Yang. However, R. fuscus is characterized by an overall reticulate stipe, slight longer basidiospores (9-12 × 3.5-4.5 μm) and narrower pileipellis hyphae (4-8 μm wide) (Zeng et al. 2016). Retiboletus griseus has a reticulum over the upper 2/3 of the stipe, a cream or grayish-brown stipe often with orange-yellow stains when hurt, and a distribution in North/Central America (Smith and Thiers 1971; Ortiz-Santana et al. 2007). Retiboletus nigrogriseus is characterized by the white to olivaceous contex in the stipe, the entirely reticulate stipe and the cutis pileipellis with hyphae 4-10 μm wide. Retiboletus pseudogriseus has a grayish white pileus which is covered with brown to blackish brown squamules, white context becoming brown when injured, and a slender and completely reticulate stipe.</p><p>In the phylogenetic analysis (Fig. 1), R. ater forms an independent lineage within Retiboletus, future studies would require more molecular sequence data to help fully resolve its evolutionary relationships to the other species.</p></div>	https://treatment.plazi.org/id/BDC8EB8A7B6E5C4F914A3DE028DCE6E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Hai-Ying;Li, Yan-Chun;Bau, Tolgor	Liu, Hai-Ying, Li, Yan-Chun, Bau, Tolgor (2020): New species of Retiboletus (Boletales, Boletaceae) from China based on morphological and molecular data. MycoKeys 67: 33-44, DOI: http://dx.doi.org/10.3897/mycokeys.67.51020, URL: http://dx.doi.org/10.3897/mycokeys.67.51020
E4227468BD4D562F9F6D8E71FFF13E81.text	E4227468BD4D562F9F6D8E71FFF13E81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Retiboletus sinogriseus Yan C. Li & T. Bau 2020	<div><p>Retiboletus sinogriseus Yan C. Li &amp; T. Bau sp. nov. Figures 2d-f, 4</p><p>Etymology.</p><p>sino (Latin) = China, reflecting that the basidiomata were collected in China + griseus for similarity of the basidiomata of this species to Retiboletus griseus .</p><p>Type.</p><p>China. Liaoning Province: Anshan City, Qianshan, alt. 400 m, 25 Aug 2015, J. Li 260 (holotype: KUN-HKAS 91288!).</p><p>Description.</p><p>Basidiomata medium-sized. Pileus 6-7.2 cm in diameter, subhemispherical to applanate, sometimes convex; surface tomentose, grayish-brown (5D2-3) to brown (5E4), rimose when dry, always cracked into small squamules on grayish (4B1) to whitish (2A1) background; context 1-2 cm thick in the center of pileus, white (1A1), unchanging when injured. Hymenophore adnate, sometimes slightly depressed around apex of stipe; pores angular, 0.3-1 mm wide, tubes up to 14 mm long, yellow (4C3-4) to grayish-yellow (4B2-3), unchanging when injured. Stipe 6-8 × 1.1-1.5 cm, subcylindrical, solid; surface dry, pale yellow at apex, blackish-yellow towards the base, entirely covered with moderately developed reticulum; context white to cream in the upper part and yellowish to yellow downwards, unchanging when injured; basal mycelium yellow. Taste and odor indistinct.</p><p>Basidia 21-27 × 9-11 μm, clavate, thin-walled, four-spored; sterigmata 4-5 μm long. Basidiospores [40/2/2] (9) 10.0-13.5 (-14.0) × (3) 4.0-5.0 (-5.5) μm, Q = (2.25-) 2.5-3.25 (-3.42), Qm = 2.88 ± 0.32, subfusiform to ellipsoid, slightly thick-walled (up to 0.5 μm), hyaline to yellowish in KOH, olive-brown to yellowish-brown in Melzer’s reagent, smooth. Hymenophoral trama boletoid. Cheilo- and pleurocystidia 35-56 × 7-12 μm, abundant, subfusiform to fusiform, thin-walled, with yellowish-brown to brown contents, without encrustations. Pileipellis a subcutis, 100-120 μm thick, composed of thin-walled filamentous hyphae 4-7 μm wide, with subcylindrical to clavate terminal cells 33-72 × 4-6 μm, sometimes with subacute apex, colorless to pale yellowish-brown in KOH, yellow-brown to brownish in Melzer’s reagent. Pileal trama composed of thin-walled hyphae 4-9 μm wide, colorless to pale yellowish-brown in KOH, yellow-brown to brownish in Melzer’s reagent. Clamp connections absent in all tissues.</p><p>Habitat, ecology and distribution.</p><p>Solitary on the ground in mixed forests dominated by plants in the families Fagaceae and Pinaceae; currently known from northeastern China.</p><p>Additional specimens examined.</p><p>China. Liaoning Province: Anshan City, Qianshan, alt. 400 m, 25 Aug 2015, J. Li 258 (KUN-HKAS 91286).</p><p>Discussion.</p><p>Retiboletus sinogriseus has a grayish-brown to brown pileus, a pale yellow to blackish-yellow stipe. Such traits are very similar to those of R. griseus . Interestingly, R. sinogriseus clusters with R. griseus with strong statistical support (Fig. 1). However, R. griseus, originally described from North America but not found in China yet, has a distinctly pallid hymenohore and broad pileipellis hyphae which are up to 17 μm wide (Singer 1947; Smith and Thiers 1971; Ortiz-Santana et al. 2007). Additionally, the R. sinogriseus / R. griseus clade is clustered with R. zhangfeii N.K. Zeng &amp; Zhu L. Yang, R. nigrogriseus and R. nigerrimus ( R. Heim) Manfr. Binder &amp; Bresinsky (however without bootstrap support). In this assemblage, R. zhangfeii differs significantly from R. sinogriseus by its differently colored pileus, hymenophore, stipe and context (Zeng et al. 2016). Retiboletus nigrogriseus has a black to gray pileus, white to grayish white hymenophore, white to olivaceous contex in the stipe and much smaller basidiospores 8-10.5 × 3.5-4.5 μm . Retiboletus nigerrimus, originally described from Papua New Guinea, has a pileus with a distinctive blue tinge, a context lemon yellow in pileus and orange in the base of stipe and longer and narrower basidiospores 11.5-14.5 × 3.6-4.6 μm (Heim 1963).</p><p>Nine species of Retiboletus were recorded from China, including two new species described herein. For the convenience of identification, a key to the species in China is given below.</p></div>	https://treatment.plazi.org/id/E4227468BD4D562F9F6D8E71FFF13E81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Hai-Ying;Li, Yan-Chun;Bau, Tolgor	Liu, Hai-Ying, Li, Yan-Chun, Bau, Tolgor (2020): New species of Retiboletus (Boletales, Boletaceae) from China based on morphological and molecular data. MycoKeys 67: 33-44, DOI: http://dx.doi.org/10.3897/mycokeys.67.51020, URL: http://dx.doi.org/10.3897/mycokeys.67.51020
