taxonID	type	description	language	source
146C8794FFFFB936FF6FADD7FC20FFE1.taxon	diagnosis	Diagnosis. Body strongly convex, oval in shape. Antenna with 10 antennomeres, with 4 - antennomere club. Mandibles form semicircle when closed; right mandible with blunt tooth on anterior one-third. Ventral side of head with paired antennal grooves next to eyes. Mesosternum vertical between coxae, vertical surface not carinate. All tibiae slender; protibia with distinct tarsal groove, its outer margin strongly spinose; mesotibia slightly widened and its outer margin densely spinose; metatibia with few spines on ventral surface near outer margin. Tarsal formula 5 – 5 – 4 in both sexes. Males distinguished by expanded pro- and mesotarsi, curved mesotibiae bearing a process on the inner apical margin, and by a toothlike expansion of the lower margin of the metafemur.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFFFB936FF6FADD7FC20FFE1.taxon	distribution	Distribution. The distribution of the genus is Holarctic and it is also found in southern India; it is questionably Afrotropical and Neotropical (Newton 1998).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFFFB934FF6FA917FD98F848.taxon	description	(Fig. 1, 16)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFFFB934FF6FA917FD98F848.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.40 – 2.10 mm; greatest width = 1.00 – 1.50 mm. Reddish brown, shiny. Head with coarse, irregularly spaced punctures. Antennomere 2 longer and broader than 3; antennal club compact, antennomere 7 narrower than 8, apical antennomere slightly narrower than 9. Sides of pronotum rounded, posterior angles roundly obtuse. Pronotum coarsely, densely punctate laterally; punctation finer and more sparse medially. Elytral strial punctures coarse, finer anteriorly, separated by one diameter or more; interstrial punctures smaller, more widely spaced; anteriorly, strial punctures are not clearly distinguished from interstrial punctures. Metasternum strongly, densely punctate; medially, punctures finer and more sparse. Legs as in generic diagnosis. Abdominal sternites with dense microsculpture; sternites III – VII each with transverse row of small punctures in apical half. Median lobe of aedeagus (Fig. 1) broad, flat apically, weakly curved dorsoventrally; paired apices variable in length. Parameres broad, flat, each bearing two spines before thin, membranous apex that extends beyond apices of median lobe. Internal sac as in Fig. 1.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFFFB934FF6FA917FD98F848.taxon	discussion	Remarks. We have found the structure of the internal sac (Fig. 1) to be consistent throughout North America. However, it appears to differ somewhat from the internal sac of C. subtestacea as illustrated by Daffner (1983: 137, fig. 500; 1988: 293, fig. 77). We have seen no European specimens of C. subtestacea for comparison, and think that the placing C. picipennis (LeConte) into synonymy under C. subtestacea (Gyllenhal) should be reexamined.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFFFB934FF6FA917FD98F848.taxon	distribution	Distribution. North American distribution (Fig. 16): CANADA. BRITISH COLUMBIA, MANITOBA, NEW BRUNSWICK, NOVA SCOTIA, ONTARIO, QUEBEC. UNITED STATES. CALIFORNIA, COLORADO, DISTRICT OF COLUMBIA, IDAHO, IOWA, ILLINOIS, MASSACHUSETTS, MAINE, MICHIGAN, MINNESOTA, MONTANA, NEVADA, NEW MEXICO, NEW YORK, OREGON, UTAH, VERMONT, VIRGINIA, WASHINGTON. Extra-limital distribution: Europe; Siberia. The species is Holarctic. Previously recorded in North America (LeConte 1863, Horn 1880, Brown 1937 b, Hatch 1957, Daffner 1988) from: CANADA. BRITISH COLUMBIA. Creston (and 20 km NE). Kamloops (Lac du Bois). Oliver. Royal Oak. Summerland. Vancouver. ONTARIO. Brent (near Algonquin Park). Mer Bleue (near Ottawa). Hawthorne. Gogama (40 km NE, Mattagami River). Pelee Island. Simcoe. Arnprior. QUEBEC. Aylmer. Duparquet. Brome. UNITED STATES. DISTRICT OF COLUMBIA. IDAHO (northern). MAINE. Paris. MASSACHUSETTS. Elizabeth Island. Fall River. Framingham. Nashawena Island. Lincoln. Sherborn. MICHIGAN. Detroit. NEW MEXICO. Catron Co., 5 mi w Luna. Chavez Co., Bitter Lakes. WASHINGTON (southwestern). Seasonality. Adults have been collected in the months from May to November, and mostly from June to August. Bionomics. Adults have been collected mostly in forested habitats. They have been taken by flight intercept traps and evening car netting, but also in litter and blacklight traps. The species is seemingly absent from mid-continental grassland habitats. New material examined. We have seen 182 specimens, from the following localities. CANADA. MANITOBA. Ste. Rita, Agassiz Provincial Forest. Volstoi-Vita, Rt. 209. NEW BRUNSWICK. Hwy. 112, 26 km W Canaan Forks. 60 km NW Moncton, rt. 116. NOVA SCOTIA. 40 km SE Truro, Trafalgar, Liscomb Provincial Sanctuary. Westchester to Londonderry. N of upper Tantalon, Indian Lake road. Clyde River road. ONTARIO. 20 km SE Almonte, Middleville to White Lake. 2 km N Cloyne, Stoll Lake road. Alfred, Alfred Bog. Almonte. Gloucester. Iroquois Falls, nw Industrial Road. Kemptville Forest. Limoges, Larose Provincial Forest. Manitoulin Island, 2 mi S Maple Point. N Lake Abitibi, 80 km E Cochrane. Marlborough Provincial Forest, 15 km W North Gower. Stittsville. Rondeau Provincial Park. Shirleys Bay, 15 km W Ottawa. Constance Bay. Wheatley. L 3 C 6 Wolford Township 44 ° 52 ’ 03 ” N, 75 ° 43 ’ 50 ” W. 7 km se Westport. UNITED STATES. CALIFORNIA. Fresno Co., Fresno. Del Norte Co., Gasquet. Siskiyou Co., Etna. COLORADO. Huerfano Co., La Veta Pass. DISTRICT OF COLUMBIA. IOWA. Webster Co., DeKalb Research Farm, 3 mi W Dayton. ILLINOIS. Champaign Co., Mahomet, Nettie Hart Memorial Woods. 3 mi NW Homer. MASSACHUSETTS. Lincoln. MINNESOTA. Roseau Co., Roseau. Frontenac. MONTANA. Toston. NEW MEXICO. Las Vegas. Roswell. Catron Co., 5 mi W Luna. 8 mi SE Luna. Chaves Co., Bitter Lakes N. W. R. NEVADA. Esmeralda Co., Fish Lake Valley dunes. NEW YORK. Willard. OREGON. Harney Co., 17 mi E Frenchglen, Steens Mt. road. 3 mi E Frenchglen, Page Spring Camp. 62 mi SE Burns, Page Spring. UTAH. Benjamin. Callao. Provo. Richfield. Salt Lake, Salt Lake Airport. VERMONT. Franklin Co., Bakersfield. VIRGINIA. Fredericksburg. Bath Co., 9.6 km N Clifton Forge. Montgomery Co., Jefferson National Forest, Pandapas Pond. WASHINGTON. Benton Co., Hanford Site, West Lake.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFFDB93BFF6FA86DFBC9FCC1.taxon	description	(Fig. 16)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFFDB93BFF6FA86DFBC9FCC1.taxon	diagnosis	Diagnosis (adapted from Daffner 1988). Length 1.6 – 2 mm. Body oval, strongly convex, reddish brown, head and antennal club slightly darker. Antenna elongate with strong club. Head finely and densely punctate. Pronotum twice as wide as long, base at obtuse hind angles slightly sinuate, punctation fine and sparse. Elytra oval, acuminate posteriorly, very strongly and densely punctate, punctures separated by one diameter, strial punctures clearly distinct only in the posterior half, basally interstrial punctures as strong as strial punctures. Metasternum strongly and sparsely punctate throughout. All sternites with series of fine punctures on lower margin, first sternite finely, densely and rugosely punctate. Aedeagus with median lobe slender; parameres large, elongate, with apices extending beyond median lobe. For illustration of aedeagus see Daffner (1988: 293, Fig. 79 - 80).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFFDB93BFF6FA86DFBC9FCC1.taxon	distribution	Distribution. North American distribution (Fig. 16). UNITED STATES. CALIFORNIA, OREGON. Previously recorded (Daffner 1988) from: CALIFORNIA. Nevada Co., Sagehen Creek. Sacramento Co., Sacramento. Mendocino Co., Fort Bragg. OREGON. Umatilla Co., 16 mi E Ukiah, Frazier Camp. Seasonality. Adults are known only from the months of May to August. Bionomics. We assume that the species occurs in forested or wooded habitats. New material examined. We have seen no new specimens of this species.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF2B93BFF6FAF17FC21FEC1.taxon	diagnosis	Diagnosis. Body strongly convex. Antenna with 10 antennomeres, with strong 3 - antennomere club. Mandibles forming semicircle when closed together; left mandible with large tooth in basal one-third. Ventral side of head with distinct antennal grooves below eyes. Mesosternum vertical between the mesocoxae, without median carina. Tibiae strong, widened and shovel-like; protibiae with distinct tarsal grooves, their outer margins with strong spines; outer margins of meso- and metatibiae strongly spinose. Tarsal formula 5 - 5 - 4 in both sexes. Males are distinguished by large tooth-like expansion of lower margin of metafemur.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF2B93BFF6FAF17FC21FEC1.taxon	distribution	Distribution. The distribution of the genus is eastern Holarctic; it also occurs in Mexico, with undescribed species in the northern Neotropics (Newton 1998).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF2B93AFF6FA9F7FDE6F841.taxon	description	(Fig. 2, 17)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF2B93AFF6FA9F7FDE6F841.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.18 – 2.00 mm; greatest width = 0.78 – 1.39 mm. Reddish brown, shiny. Head finely, sparsely punctate. Antennomere 2 clearly longer and broader than 3; antennomere 7 thin, about half width of 8; apical antennomere distinctly narrower than 9. Sides of pronotum rounded, posterior angles obtuse. Pronotum finely, sparsely punctate. Elytral strial punctures coarse, finer anteriorly, separated by one diameter or less, except anteriorly; interstrial punctures minute, sparse. Metasternum strongly, densely punctate. Legs as in generic diagnosis. Abdominal sternites III – VIII each with transverse row of small punctures on apical half; sternite III rugose. Median lobe of aedeagus (Fig. 2) broad, angled about 90 degrees in lateral view; paired apices each divided at tip. Parameres (Fig. 2) extend beyond apex of median lobe. Armature of inverted internal sac as in Fig. 2.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF2B93AFF6FA9F7FDE6F841.taxon	distribution	Distribution. North American distribution (Fig. 17): CANADA. ONTARIO, QUEBEC. UNITED STATES. ALABAMA, ARKANSAS, ARIZONA, DISTRICT OF COLUMBIA, FLORIDA, GEORGIA, INDIANA, KENTUCKY, LOUISIANA, MICHIGAN, MISSISSIPPI, NORTH CAROLINA, OKLAHOMA, SOUTH CAROLINA, TEXAS, VIRGINIA. Extra-limital distribution: Mexico (doubtful, see Remarks). Previously recorded in North America (Horn 1880, Brown 1937 b, Daffner 1988) from: CANADA. ONTARIO. Arnprior. QUEBEC. Aylmer. UNITED STATES. MICHIGAN. Detroit. DISTRICT OF COLUMBIA. Seasonality. Adults have been collected from February to November, and mostly in June to August, with the early and late season records from southerly locations in Florida and Texas. Bionomics. Adults have been collected in habitats ranging from open pine forest and juniper thicket to mature mixed and bottomland beech-magnolia forest. Taken mostly by flight intercept traps but also by sweeping and ultraviolet light traps.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF2B93AFF6FA9F7FDE6F841.taxon	discussion	Remarks. We question the above synonymy and think it needs re-examination. We suspect that I. uncus is actually a valid species, and we have seen several undescribed species of Isoplastus from Mexico and southward. New material examined. We have seen 434 specimens from the following localities. CANADA. ONTARIO. L 3 C 6 Wolford Township, 44 ° 52 ’ 03 ” N, 75 ° 43 ’ 50 ” W. Tilbury. UNITED STATES. ALABAMA. Dale Co. Ft. Rucker Military Reservation. Jefferson Co., Hoover. ARKANSAS. Little Rock Co., Little Rock. Logan Co., Magazine Mountain. ARIZONA. Cochise Co., Chiricahua Mts., Turkey Creek. FLORIDA. Alachua Co., Gainesville. 10 km SW Gainesville. Jackson Co., Marianna, Florida Caverns State Park. Sneads, Three Rivers State Park. Leon Co., 20 mi N Tallahassee, Tall Timbers Station. Liberty Co. 14 km N Bristol, Appalachicola Bluffs and Ravines Preserve. Suwanee Co., Suwanee River State Park. GEORGIA. Greene Co., Oconee National Forest. INDIANA. Tippecanoe Co. KENTUCKY. Edmonson Co., Mammoth Cave National Park. Metcalfe Co., 0.5 mi NE East Fork. LOUISIANA. Catahoula Parish, Walters, Catahoula National Wildlife Refuge. Grant Parish, 18 km N Alexandria, Stuart Lake Camp. Natchitoches Parish, Kisahatchie Bayou. West Feliciana Parish, Feliciana Preserve. MISSISSIPPI. Pontotoc Co., 32 km SW Tupelo, Tockshish. NORTH CAROLINA. Northampton Co., 7 km S Jackson. OKLAHOMA. Latimer Co., Red Oak. Wagoner Co., 3 mi W Wagoner. SOUTH CAROLINA Anderson Co., Pendleton. Oconee Co., 12 mi NE Walhalla. Pickens Co., Calhoun. TEXAS. Bell Co., 1 km W Youngsport. Brazos Co., Lick Creek Park, College Station. Ellis Co., 3.6 mi NW Italy. Hays Co., 6 mi NW Dripping Springs. 6.5 mi SW Hunt. Kerr Co., Kerrville. Montgomery Co., 4.5 mi N Montgomery. Polk Co., 20 km S Coldspring. Sabine Co., 9 mi E Hemphill, Beech Bottom. San Augustine Co., Piney Woods Conservation Center. San Jacinto Co., 5 km S Coldspring. Shelby Co., 20 km SE Shelby. Tyler Co., 5 mi E Spurger. Wood. Co., Godwin Woods, 3.5 mi SW Hainesville. VIRGINIA. Fredericksburg. Warren Co., 4 km NNW Linden.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF3B939FF6FA897FEDBFB21.taxon	description	(Fig. 3, 17) Description. Length (pronotum + elytra) = 1.36 – 1.80 mm; greatest width = 0.96 – 1.36 mm. Yellowish to reddish brown, shiny. Head moderately coarsely, irregularly punctate. Antennomere 2 clearly longer and broader than 3; antennomere 7 thin, less than half as wide as 8; apical antennomere distinctly narrower than 9. Sides of pronotum rounded, posterior angles obtuse. Pronotum finely, moderately sparsely punctate. Elytral strial punctures coarse, closely spaced; strial rows reach base on inner half of elytra. Interstrial punctures fine, moderately sparse. Metasternum moderately coarsely, irregularly punctate. Legs as in generic diagnosis. Abdominal sternites III – VIII weakly rugose, each with transverse row of small punctures on apical half. Median lobe of aedeagus (Fig. 3) evenly curved in lateral view; acute tips of paired apices inwardly curved. Parameres not extending beyond apex of median lobe. Armature of inverted internal sac as in Fig. 3.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF3B939FF6FA897FEDBFB21.taxon	materials_examined	Type material. Holotype, male, in SBPC, with label data: “ Archbold Biol. Sta. / Lk. Placid / Highlands Co. FLA. / 2 July 1986 / M. Deyrup // Malaise Trap / Trail 1 SSo ”. Paratypes, 41, as follows: “ FLORIDA: Collier Co. / Naples / V- 16 1984 / R. A. Belmont, colr. / u. v. blacklight trap ” (1, FSCA); same data except “ VII- 16 1984 ” (2, FSCA); “ FLORIDA: Gilchrist Co. / W. High Springs, nr Poe / Springs; 14 - VI- 1994 / C. Hernandez, pitfall ” (2, FSCA); same data as holotype except “ 3 Sept. 1986 // Trail 2 SSo ” (1, ABSC); same data except “ 30 Aug. 1983 ” (1, ABSC); same data except “ 8 Aug 1987 // SSo ” (1, ABSC); same data except: “ 5 May 1985 // Trail 2 SSo ” (1, ABSC); same data except “ 29 Aug 1987 // SSo ” (2, ABSC); “ Archbold Biol. Sta. / Lk. Placid / Highlands Co. FLA / 19 Aug 1993 / M. Deyrup ” (1, ABSC); FL: Highlands Co. / Archbold Biol. Sta. / 18 June 1998; Mark Deyrup // in malaise trap in / scrubby flatwoods / near weather / station ” (1, ABSC); FL: Highlands Co. / Archbold Biol. Sta. / 28 August 2009 / M. Deyrup, H. Otte // flight trap w. pan / Ceratiola habitat / Site 2. N 27.14512 / W 081.35766 ” (1, ABSC); “ FL: Highlands Co. / Carter Ck. North / 13 April 2009 / M. Deyrup, A. May, H. Otte // Yellow sand scrub / N 27.533, W 081.412 / bowl traps / fringe tree area ” (2, SBPC); same data except: “ 14 April 2009 // Townes trap / N 27.53362 / W 081.40720 ” (4, SBPC); “ FL: Highlands Co. / Flamingo Villas / 22 May 2009 / M. Deyrup, A. May, H. Otte // scrub habitat / Townes trap / N 27.44228 / W 081.38276 ” (1, ABSC); “ FL: Highlands Co. / Flamingo Villas Pres / 28 May 2009. H. Otte / A. May, M. Deyrup // flight trap w. pan / scrub habitat / Site 4. N 27.44231 / W 081.37822 ” (1, SBPC); same data except: “ 26 May 2009 // Malaise trap / scrub habitat ” (1, SBPC); “ FL: Highlands Co. / Highlands Hammock / State Park / 26 May 2009. H. Otte / A. May, M. Deyrup // flight trap w. pan / scrub habitat / Site 4. N 27.43213 / W 081.51671 ” (1, ABSC); same data except: “ 27 May 2009 // scrub habitat / flight trap pan / N 27.43378 / W 081.51613 ” (1, ABSC); same data except: “ 4 Aug. 2009 // scrub habitat / bowl trap / N 27.48192 / W 081.52861 ” (1, ABSC); same data except: “ 6 Aug. 2009 // yellow bowl trap / scrub habitat / Site 8. N 27.48376 / W 081.52372 ” (1, ABSC); “ FL: Highlands Co. / Highlands Park / Estates. 14 Aug. 2009 / M. Deyrup, H. Otte / A. May // scrub habitat / flight trap / N 27.53864 / W 081.35071 ” (1, ABSC); FL: Highlands Co. / Lake June State Pk. / 28 May 2009 / M. Deyrup, A. May / H. Otte // Townes trap / scrub habitat / N 27.29982 / W 081.42441 ” (1, SBPC); same data except: “ 2 June 2009 // flight trap w. pan / scrub habitat / N 27.30001 / W 081.42403 ” (2, SBPC); “ FLORIDA: Levy Co. / 4.0 mi SW Archer / 9 - VI- 1992; P. E. / Skelley, b-light ” (1, FSCA); “ FLORIDA: Levy Co. / 4.0 mi. SW Archer on / Rt- 24; 15 - VI- 1996 / P. E. Skelley. Uv light ” (2, FSCA); “ FL: Polk Co. / A. D. Broussard / Catfish Ck. State Pk. / 5 Jun. 2009, M. Deyrup / H. Otte, A. May // flight trap with pan / scrub habitat / N 27.98598 / W 081.49663 ” (1, SBPC); “ FL: Polk Co. / Lake Marion Crk. / Estates / 28 - V- 1998 / R. Morris / FIT ” (1, FSCA); FL: Polk Co. Lake / Wales Ridge Forest / 14 July 2009, H. Otte / A. May, M. Deyrup // flight trap w. pan / scrub habitat / Site 5. N 27.68668 / W 081.49616 ” (1, ABSC); “ FL: Polk Co., TNC / Tiger Creek Pres. / 6 July 2009 / M. Deyrup, H. Otte / A. May // scrub habitat / flight trap / N 27.82158 / W 081.47717 ” (1, SBPC); “ UNITED STATES: FL: Sarasota Co., Myakka / River St. Pk. 16 & 17. IV. 08 / N 27 ° 16.5 ’ W 82 ° 16. 2.8 km / Liveoak-cabbage palm hammock / Eve car netting. S. Peck 08 – 45 ” (1, SBPC).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF3B939FF6FA897FEDBFB21.taxon	distribution	Distribution. The species is known only from peninsular Florida, at the localities given above (Fig. 17). Seasonality. Adults have been collected from the months of April to September, and mostly in June to August. Bionomics. Adults have been collected mostly in forest and scrub vegetation on sandy soils. They have been taken by malaise, flight intercept, blacklight, car net, and yellow bowl traps.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF3B939FF6FA897FEDBFB21.taxon	etymology	Etymology. This species is named for the state of Florida, U. S. A., where all know specimens have been collected.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF0B938FF6FAB76FED4FCA1.taxon	diagnosis	Diagnosis. Body moderately convex. Antenna with 11 antennomeres, with 5 - antennomere club interrupted at antennomere 8, which is narrow and disc-shaped but clearly visible. Mandibles forming semicircle when closed together, and without prominent teeth. Ventral side of head without antennal grooves. Mesosternum vertical between the mesocoxae, not carinate. Tibiae uniformly widened apically; protibia with distinct tarsal groove, its outer margin strongly spinose; mesotibia outer margin with two dense rows of spines; outer margin of metatibia with short, strong spines. Tarsal formula 5 - 5 - 4 in both sexes. Males distinguished by strongly curved mesotibiae bearing process on inner apical margin and by strong tooth-like expansion of lower margin of metafemur.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF0B938FF6FAB76FED4FCA1.taxon	distribution	Distribution. The genus is Holarctic in distribution; it also occurs in India, Thailand, and China (Newton 1998).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF1B93FFF6FAE97FEC8FFA1.taxon	description	(Fig. 4, 18)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF1B93FFF6FAE97FEC8FFA1.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.49 – 2.16 mm; greatest width = 0.98 – 1.42 mm. Reddish brown, moderately shiny. Fine, irregular microsculpture scattered on head and pronotum, more prominent on elytra. Head with fine, moderately spaced punctures anteriorly; punctures larger and more closely spaced on vertex. Antennal club variable in width; antennomere 8 thin and about half width of antennomere 7; apical antennomere clearly narrower than 9 and 10. Sides of pronotum rounded, posterior angles obtuse. Pronotum moderately finely, sparsely punctate medially, punctures more closely spaced laterally. Elytral strial punctures coarse, separated by less than one diameter, visible to base of pronotum; interstrial punctures smaller and more widely spaced. Metasternum coarsely punctate, punctures more densely spaced laterally, with impunctate area medially. Metatibia broad, with apex about three times width at base. Male mesotibial process triangular. Abdominal sternite III rugose, sternites IV – VII each with row of fine punctures before apex. Median lobe of aedeagus broad; apical lobes short with rounded apices (Fig. 4). Parameres (Fig. 4) broad, with membranous apices extending beyond apices of median lobe. Internal sac as in Fig. 4.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF1B93FFF6FAE97FEC8FFA1.taxon	distribution	Distribution. North American distribution (Fig. 18): CANADA. ALBERTA, BRITISH COLUMBIA. MANITOBA, ONTARIO, QUEBEC, SASKATCHEWAN. UNITED STATES. ARIZONA, CALIFORNIA, IDAHO. MINNESOTA, UTAH, WASHINGTON. Previously recorded in North America (Hatch 1927, Brown 1937 b, Daffner 1985, 1988) from: CANADA. BRITISH COLUMBIA. Creston. 23 km NE Creston. Mabel Lake at Squaw Valley. 13 km N Nelway. Royal Oak. Summerland. MANITOBA. Aweme. ONTARIO. Arnprior. Constance Bay. Leamington. Mer Bleue, near Ottawa. QUEBEC. Aylmer. SASKATCHEWAN. Saskatoon. UNITED STATES. ARIZONA. Flagstaff. CALIFORNIA. Nevada Co., Sagehen Creek. WASHINGTON. Okanogan Co., near Buzzard Lake. Seasonality. Adults are known from the months of May to October, with most from June to August. Bionomics. Adults have been collected in habitats ranging from tall-grass prairie and riparian forest, to spruce-aspen forest, and mostly from mixed hardwood forest. Taken by flight intercept traps, but also by evening car netting, sweeping, and ultraviolet light traps. New material examined. We have seen 53 specimens from the following localities. CANADA ALBERTA. 15 km SW Beaver Mines on road 774. Cypress Hills Provincial park. MANITOBA. Richer, Sandilands Provincial Forest, forest road 19. Pinawa, route 433. 10 - 12 km E. Marchand. 17 km N Woodridge, highway 210. St. Rita, Agassiz Provincial Forest. Tolstoi to Vita. ONTARIO. Arnprior. Kemptville Forest. Constance Bay. Thunder Bay. L 3 C 6 Wolford Township, 44 ° 52 ’ 03 ” N, 75 ° 43 ’ 50 ” W. UNITED STATES. ARIZONA. Flagstaff, Oak Creek Canyon at Sterling Canyon. Pinery Creek, Chiricahua Mountains. Yavapai Co., Village Oak Creek. IDAHO. Bonner Co., Sagie. MINNESOTA. Lyon Co., Camden State Park. Pipestone Co., Pipestone National Monument. UTAH. Daggett Co., nr. Red Canyon, Flaming Gorge National Recreation Area. Summit Co., Henrys Fork Camp. WASHINGTON. Walla Walla.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF6B93EFF6FAFF4FE6EFFC1.taxon	description	(Fig. 5, 19)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF6B93EFF6FAFF4FE6EFFC1.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.78 – 2.40 mm; greatest width = 1.22 – 1.70 mm. Pale to dark reddish brown, mostly shiny, antennal club often darker; some individuals with microsculpture on elytra. Head moderately coarsely and densely punctate. Antennal club variable in width, more robust in larger individuals. Antennomere 8 disc-like, about half width of 9. Apical antennomere clearly narrower than 9 and 10. Sides of pronotum rounded, posterior angles obtuse. Pronotum moderately finely punctate; punctures irregularly separated, more dense laterally. Elytral strial punctures coarse, separated by less than one diameter; striae cannot be separated apically from coarse interstrial punctures. Metasternum coarsely punctate; punctation dense laterally, sparse medially. Metatibia narrow, width at apex twice width at base. Male mesotibial process short, triangular. Abdominal sternite III coarsely punctate, sternites IV – VII each with transverse row of fine punctures before apex. Median lobe of aedeagus broad, apical lobes short with rounded apices (Fig. 5). Parameres (Fig. 5) broad, with membranous apices extending well beyond apices of median lobe. Internal sac as in Fig. 5.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF6B93EFF6FAFF4FE6EFFC1.taxon	distribution	Distribution. The species is Holarctic. North American distribution (Fig. 19): CANADA. ALBERTA, BRITISH COLUMBIA. MANITOBA, NUNAVUT TERRITORY, ONTARIO, QUEBEC, SASKATCHEWAN, YUKON TERRITORY. UNITED STATES. ALASKA, IDAHO, INDIANA, NORTH CAROLINA, NEVADA, UTAH, OREGON. VIRGINIA, WASHINGTON, WYOMING. Extra-limital distribution: Eastern Europe, Mongolia, Siberia. Previously recorded in North America (Daffner 1983, 1988) from: CANADA. BRITISH COLUMBIA. 25 km W Creston. 10 km W Summerland. 13 km N Nelway. ONTARIO. Chaffeys Locks. Stittsville. QUEBEC. Gatineau. UNITED STATES. VIRGINIA. Pulaski Co., 7 mi SE Mechanicsburg. WYOMING. 34 mi E Lovell. OREGON. Clackamas Co., Salmon River near Zigzag. WASHINGTON. Okanogan Co., near Buzzard Lake. Seasonality. Adults have been collected in the months from June to October, with most in July to September. Bionomics. In North America, the species is mostly northern or at upper elevations in boreal forest or in tundra habitats. Adults have been collected in habitats ranging from subarctic tundra to spruce-willow and pine and pine-aspen forests to mixed hardwood forest. They have been taken mostly by flight intercept traps, but also by evening car netting, and unbaited pit-traps. New material examined. We have seen 232 specimens from the following localities. CANADA ALBERTA. Opal. MANITOBA. 100 road km SE Flin Flon. 17 km N Woodridge, Sandilands Provincial Forest. NUNAVUT TERRITORY. Kugluktuk, N 67.78463, W 115.20987. ONTARIO. 20 km SE Almonte, Middleville to White Lake. Chaffeys Locks, Queens University Biology Station. Gloucester. 15 km NW Renfrew. Hamilton. Manitoulin Island. Heckston, 20 km SE Kempville. 15 km W Ottawa, Shirleys Bay. Stittsville. L 3 C 6 Wolford Township, 44 ° 52 ’ 03 ” N, 75 ° 43 ’ 50 ” W. SASKATCHEWAN. Moose Mountain Provincial Park, Kenosee Lake. YUKON TERRITORY. Champagne, Alaska Highway km 1580. Dempster Jct., 40 km E Dawson. EMAN plot, site LMK 333 Y. Haines Jct, 10 km S. Long Lake Road, site LMK 25 Moose Creek, 14 km NW Stewart Crossing. Ross River. UNITED STATES. ALASKA. 11 mi S Anderson Jct, Rt. 3, mi 270, Alaska highway. Big Delta. Nenana, 13 mi NE, mile 318, Alaska Highway. Tok. IDAHO. Clark Co., 2.8 km W rt. 115, Stoddard Campground. INDIANA. Tippecanoe Co. NORTH CAROLINA. Haywood Co., Balsam Mt., Blue Ridge Parkway. NEVADA White Pine Co., Snake Range, Wheeler Peak trail. UTAH. Summit Co., Bear River Camp. WYOMING. 34 mi E Lovell, Big Horn Mountains.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF7B93EFF6FAE17FD47FBA1.taxon	description	(Fig. 6, 19)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF7B93EFF6FAE17FD47FBA1.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.55 – 1.68 mm; greatest width = 1.16 – 1.24 mm. Reddish brown, shiny. Head finely, sparsely punctate. Antennal club variable in width; antennomere 8 saucer-shaped, about half width of antennomere 9; apical antennomere longer than and almost as wide as 10. Sides of pronotum weakly rounded, posterior angles obtuse. Pronotum minutely, sparsely punctate. Elytral strial punctures coarse and closely spaced, striae distinct to base of pronotum; interstrial punctures minute, sparse. Metasternum coarsely, densely punctate laterally, finely punctate medially. Metatibia narrow, width at apex about twice width at base. Male mesotibial process moderately long, crossing over long tibial spine. Abdominal sternites III – VII each with a row of coarse punctures at base. Median lobe of aedeagus (Fig. 6) broad with moderately long apical lobes. Parameres (Fig. 6) narrow, not reaching apices of median lobe. Internal sac as in Fig. 6.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF7B93EFF6FAE17FD47FBA1.taxon	distribution	Distribution. North American distribution (Fig. 19): UNITED STATES. ALABAMA. NEBRASKA, OKLAHOMA Previously recorded only from Latimer County, Oklahoma. Seasonality. Adults are known from the months of March, May, and September. Bionomics. The few collecting records suggest that adults occur in open and mixed forests. One specimen is from beach drift. New material examined. We have examined three specimens with the following data: UNITED STATES. ALABAMA. Wilcox County, Camden, 24. III. 1959, J. I. Cowger (1, USNM). NEBRASKA. Keith County, Arthur Bay, Lake McConaughy, 22. V. 2008, R. Turnbow, beach drift (1, SBPC). OKLAHOMA. Latimer Co., IX. 1990, K. Stephan (1, TAMU).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF4B93DFF6FADD7FA08F881.taxon	diagnosis	Diagnosis. Body convex. Antenna with 11 antennomeres with 5 - antennomere club interrupted at antennomere 8, which is narrow and disc-shaped but clearly visible. Mandibles variable, left mandible usually with one or two small blunt teeth on anterior one-third. Ventral side of head with or without well defined antennal grooves. Mesosternum vertical between mesocoxae, not carinate. Tibiae uniformly widened to apex; protibiae with tarsal grooves; outer margins of all tibiae with strong spines. Tarsal formula 5 - 5 - 4 in both sexes. Abdominal sternites III – VII each with basal transverse row of large punctures. Males are distinguished by larger, often sinuate spur on inner apical margin of usually straight mesotibia; inner apical margin of mesotibia not produced; and apical tooth-like expansion of the lower margin of the metafemur.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF4B93DFF6FADD7FA08F881.taxon	distribution	Distribution. The distribution of the genus is Nearctic and northern Neotropical (Mexico, Colombia) (Newton 1998).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF4B93CFF6FA8D7FBE5F8C2.taxon	description	(Fig. 7, 20)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF4B93CFF6FA8D7FBE5F8C2.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.50 – 2.00 mm; greatest width = 1.10 – 1.60 mm. Reddish brown, shiny. Head punctures fine anteriorly, coarser and denser posteriorly and laterally. Head with antennal grooves ventrally. Mandibles short, forming semicircle when closed; minute tooth near middle of left mandible. Antennal club slender; antennomere 8 disc-like, nearly as wide as apex of 7; apical antennomere longer than and nearly as wide as 10. Eyes of normal size. Pronotum broad, weakly rounded laterally, posterior angles obtuse, basal margin strongly rounded. Pronotum finely punctate. Elytral strial punctures deep, separated by ± 1 diameter. Interstrial punctures minute and sparse. Metasternum with fine punctures medially; coarse, unevenly spaced punctures laterally. Male mesofemur with acute toothlike expansion at apex of inner margin. Male mesotibia sinuate on inner margin. Abdominal sternites III – VII each with basal transverse row of large punctures. Median lobe of aedeagus (Fig. 7) cylindrical, strongly curved dorsoventrally at basal one-third; in lateral view, evenly dorsoventrally flattened in apical one-fourth; paired apices moderately broad. Parameres narrow, not reaching apex of median lobe. Internal sac as in Fig. 7.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF4B93CFF6FA8D7FBE5F8C2.taxon	distribution	Distribution. North American distribution (Fig. 20): UNITED STATES. DISTRICT OF COLUMBIA, FLORIDA, GEORGIA, MARYLAND, MISSISSIPPI, OKLAHOMA, TEXAS, VIRGINIA. Previously recorded in North America (Brown 1937 b, Daffner 1988) from: UNITED STATES. DISTRICT OF COLUMBIA. FLORIDA. Dade Co., Chekika State Recreation Area, 50 km SW Miami. Deering Estate Park, S. Miami. Long Pine Key, Everglades National Park. Monroe Co., 1.5 km NW Royal Palm and Royal Palm Hammock. OKLAHOMA. Latimer Co., Red Oak. VIRGINIA. Luray, Shenandoah National Park. Daffner (1988) includes a single specimen record from Arizona: Santa Cruz Co., Patagonia, Sonoita Creek Sanctuary. We have not seen this specimen to confirm the identity and have not included the record on our map. Seasonality. Adults have been collected from April to November, with most in the months of June to August. Bionomics. Adults have been collected in various forest types ranging from cypress swamp, subtropical hammock, pinelands, oak-palm hammock, bottomland forest, redbay-live oak forest, and post oak savannah, mostly on sandy soils, They have been taken mostly by flight intercept traps but some from malaise and black light traps. New material examined. We have seen 242 specimens from the following localities: UNITED STATES. DISTRICT OF COLUMBIA. Washington. FLORIDA. Collier Co., Ochopee. Dade Co., Chekika State Recreation Area. Long Pine Key. Deering Estate Park. Highlands Co., Archbold Biology Station, Lake Placid. Highlands Hammock State Park. Hillsborough Co., Hillsborough State Park. Jackson Co., Florida Caverns State Park. Levy Co., 6 mi W Otter Creek. Monroe Co., Royal Palm Hammock. 1.5 km NW Royal Palm. Putnam Co., Welaka, University of Florida Station. GEORGIA. Clarke Co., Athens. MARYLAND. Talbot Co., Wittman. MISSISSIPPI. Pontotoc Co., 32 km SW Tupelo, Tockshish. OKLAHOMA. Latimer Co., Red Oak. TEXAS. Aransas Co., 10 km SE Austwell. Brazos Co., Lick Creek Park, College Station. El Paso Co., Wilderness Park Museum. Montgomery Co., 4.5 mi N Montgomery. Wood Co., 3.5 mi SW Hainesville, Godwin Woods. VIRGINIA. Ft. Monroe.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF5B922FF6FAB17FD05FF22.taxon	description	(Fig. 8, 21)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF5B922FF6FAB17FD05FF22.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.90 – 2.40 mm; greatest width = 1.26 – 1.70 mm. Reddish brown, shiny. Head punctures fine and sparse anteriorly, coarser and more dense posteriorly and laterally. Head with antennal grooves ventrally. Mandibles short, forming semicircle when closed; small tooth at apical one-third of left mandible. Antennal club slender; antennomere 8 disc-like, narrower than apex of 7; apical antennomere about as long as and narrower than antennomere 10. Eyes of normal size. Pronotum broad, rounded laterally, posterior angles obtuse, basal margin rounded. Pronotum finely punctate; punctures sparse medially, more dense laterally. Elytral strial punctures deep, separated by ± 1 diameter. Interstrial punctures minute and sparse. Metasternum finely punctate medially; with coarse, irregularly spaced punctures laterally. Male mesofemur unmodified. Male mesotibia straight on inner margin. Abdominal sternites III – VII each with basal transverse row of large punctures. Median lobe of aedeagus (Fig. 8) cylindrical, evenly curved in lateral view; paired apices short, moderately slender. Parameres narrow, not reaching apex of median lobe. Internal sac as in Fig. 8.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFF5B922FF6FAB17FD05FF22.taxon	distribution	Distribution. North American distribution (Fig. 21): CANADA. ONTARIO, QUEBEC. UNITED STATES. ARKANSAS, FLORIDA, GEORGIA, ILLINOIS, INDIANA, KENTUCKY, LOUISIANA, MASSACHUSETTS, MICHIGAN, MINNESOTA, MISSISSIPPI, NORTH CAROLINA, NEW JERSEY, NEW YORK, OKLAHOMA, SOUTH CAROLINA, TENNESSEE, TEXAS, VIRGINIA. Extra-limital distribution: Mexico. Previously recorded in North America (Daffner 1988) from: CANADA. ONTARIO. 7 km SW Carleton Place. Rondeau Provincial Park. UNITED STATES. MASSACHUSETTS. Middlesex Co., Lincoln. NEW JERSEY. Cape May Co., Eldora. INDIANA. Parke Co., 4 mi W Rockville. FLORIDA. Putnam Co., Welaka. Lake Co., Ocala National Forest, 5.6 mi S Astor Park, Alexander Springs Creek. Alachua Co., Gainesville. OKLAHOMA. Latimer Co., Red Oak. Seasonality. Adults have been collected from February to October, with most in the months of June to August. Bionomics. Adults have been collected mostly in upland and lowland mixed forests, and a few in juniper and other scrub vegetation. They were taken mostly by flight intercept traps, but a few in malt traps, malaise traps, by Berlese extraction from litter, sweeping, evening car netting, and blacklight traps. New material examined. We have seen 855 specimens from the following localities: CANADA. ONTARIO. Chaffeys Locks, Queens University Biology Station. Heckston, 20 km SE Kemptville. Rondeau Provincial Park. Shirleys Bay, 15 km W Ottawa. L 3 C 6 Wolford Township, 44 ° 52 ’ 03 ” N, 75 ° 43 ’ 50 ” W. QUEBEC. Hull, Gatineau Park. UNITED STATES. ALABAMA. Dale Co., Fort Rucker Military Reservation. Jackson Co., 10 mi N Scottsboro. Marshall Co., 1 mi NW Guntersville. ARKANSAS. Pulaski Co., Little Rock. Washington Co., 3 mi S Devils Den State Park. FLORIDA. Alachua Co., Gainesville. S. side Bivens Arm Lake. Calhoun Co., Scotts Ferry, highway 71. Clay Co., Gold Head Branch State Park. Columbia Co., O’Leno State Park. Dade Co., Chekika State Recreation Area. Gadsden Co., 1 km E Chattahoochee. Hernando Co., Bayport. Hillsboro Co., Hillsboro River State Park. Jackson Co., Florida Caverns State Park. Three Rivers State Park. Jefferson Co., Rt. 98 at W. Aucilla Sloughs. Lake Co., 5.6 mi S Astor Park, Alexander Springs Cree. Leon Co., 20 km N Tallahassee, Tall Timbers Research Station. Levy Co., 6 mi W. Otter Creek. 16 km NE Cedar Key. Liberty Co;., 14 km N Bristol, Appalachicola River Bluffs Preserve. Torreya State Park. Madison Co., 5 km S Ellaville. Marion Co., Ocala National Forest. Silver Springs Road. Okaloosa Co., Milligan. Osceola Co., Kenansville. Putnam Co., Welaka, University Florida Station. Sarasota Co., Myakka River State Park. St. Johns Co., Faver-Dykes State Park. Suwanee Co., Suwanee River State Park. Walton Co., 18 km SW Freeport. Topsail Hill State Park, Campbell Lake. GEORGIA. Macon, Ocmulgee National Monument. Brantley Co., Hickox. Clarke Co., Whitehall Forest. Greene Co., Oconee National Forest, Oconee River Recreation Area. McIntosh Co., Sapelo Island. ILLINOIS. Coles Co., Charleston. Jackson Co., 7 mi W Carbondale. Mason Co., Sand Ridge State Forest. INDIANA. Parke Co., 4 mi W Rockville. Tippecanoe Co. KENTUCKY. Edmonson Co., Mammoth Cave National Park. Rowan Co., 24 km SW Morehead, Cave Run Lake. Whitley Co., D. Boone National Forest. LOUISIANA. Calcasieu Parish, Sam Houston Jones State Park. Feliciana Parish, Feliciana Preserve. Grant Parish, 18 km N Alexandria, Stuart Lake Camp. 3 KM SW Pollack. MICHIGAN. Kalamazoo Co., Hickory Corners. MINNESOTA. Lyon Co., Camden State Park. Itasca Co., Plot 2, T 58, R 24, S 35. Pine Co., Little Sand Creek. MISSISSIPPI. Hind Co., 19 km NE Gibson, Owens Creek. Pontotoc Co., 32 km SW Tupelo. NORTH CAROLINA. Northampton Co., 5 km S Jackson. Wilkes Co., Jeffress Park, Blue Ridge Parkway. NEW YORK. Elbridge. OKLAHOMA. Latimer Co. Red Oak. SOUTH CAROLINA. Anderson Co., Pendleton. TENNESSEE. Sevier Co., 2.5 mi W Gatlinburg. TEXAS. Angelina Co., 2.5 mi N Rockland. Bandera Co., Lost Maple State Natural Area. Bell Co., 1 km W Youngsport. Brazos Co., Lick Creek Park, College Station. Brewster Co., Big Bend National Park, N side Emory Peak, Maple Canyon. Ellis Co., 3.6 mi NE Italy. Hays Co., 6 mi NW Dripping Springs. Jeff Davis Co., Davis Mountains Resort, Upper Lympia Creek Canyon. Kerr Co., 6.5 mi SW Hunt. Polk Co., 20 km S Coldspring. Sabine Co., 8 mi E Hemphill. 9 mi E Hemphill, “ Beech Bottom. ” San Jacinto Co., 5 km S Coldspring. Big Creek Scenic Area. Shelby Co., 20 km SE Shelbyville. Travis Co., Bull Creek Arm of Lake Austin. Trinity Co., 9.6 km S. Apple Springs. Tyler Co., L Kirby State Forest. Wood Co., 3.5 mi SW Hainesville. VIRGINIA. Bath Co., 9.6 km N Clifton Forge. Bedford Co., Blue Ridge Parkway, mile 55, White Oak Flat. Essex Co., 1 mi SE Dunnsville. Fairfax Co., Burling Park. Franklin Co., Blue Ridge Parkway, mile 154, Smart View. Shenandoah Co., 16 km S Strasburg, Elizabeth Furnace. Note. One male, tentatively identified as this species, is from BRITISH COLUMBIA. 5 mi. NW Oliver, 12. VI. 1958, H. & A Howden (CNCI). This outlier record needs to be confirmed with additional material. We do not show it on the map.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEBB921FF6FAF77FA2FFDC1.taxon	description	(Fig. 9, 22) Description. Length (pronotum + elytra) = 1.80 – 2.08 mm; greatest width = 1.16 – 1.44 mm. Reddish brown, shiny. Head punctures fine medially, moderately coarse on vertex, irregularly spaced. Antennal grooves not well defined. Mandibles short, forming semicircle when closed; two teeth at apical one-third of left mandible. Antennal club slender; antennomere 8 disc-like, distinctly narrower than apex of 7; apical antennomere slightly narrower than 9 and 10. Eyes of normal size. Pronotum broad, sides rounded, posterior angles roundly obtuse, basal margin rounded. Pronotal punctures minute and sparse. Elytral strial punctures round and deep, smaller near base, variably separated; interstrial punctures minute and sparse. Metasternum with fine punctures medially; coarse, unevenly spaced punctures laterally. Male mesofemur unmodified. Male mesotibia weakly concave on inner margin. Abdominal sternites III – VII each with basal transverse row of large punctures. Median lobe of aedeagus (Fig. 9) cylindrical, strongly curved dorsoventrally; apical one-third progressively flattened to apex; paired apices broad and flat. Parameres narrow, each bearing two apical setae, not reaching apex of median lobe. Internal sac as in Fig. 9.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEBB921FF6FAF77FA2FFDC1.taxon	materials_examined	Type material. Holotype, male, in TAMU, with label data: “ UNITED STATES: TEXAS: Bell Co. / 1 km W Youngsport, FIT / Bowmer Rch., Lampasas Riv. / 30.96197 ° N. 97.72903 ° W / IV- 10 – 21 - 2010, T. Robbins ”. Paratypes, 71, as follows: 20, same data as holotype (10, TAMU; 10, SBPC); “ UNITED STATES Texas Brewster Co. / Big Bend National Park / Maple Cyn. 5200 ’ / 9. VII. 82 G. A. P. Gibson ” (2, SBPC); same data except: “ 30. VI – 9. VII. 82 / pan trap ” (2, SBPC); “ UNITED STATES: TEXAS: Hays Co. / 6 mi NW Dripping Springs / 30 ° 13.589 ’ N, 98 ° 11.096 ’ W / X- 3 – 30 - 2005, FIT-ground / 1,340 ’, E. G. Riley et al. - 119 / Juniperus unmanaged plot ” (2, TAMU); same data except: “ VI- 3 – 30 - 2005 / 418 / Juniperus managed plot ” (1, TAMU); same data except: “ III- 31 – IV- 26 - 2006 / 340 ” (2, TAMU); same data except: “ IV- 28 – VI- 2 - 2006 / 379 / Juniperus managed plot ” (7, TAMU); same data except: “ 30 ° 13.554 ’ N, 98 ° 11.039 ’ W / III- 31 – IV- 27 - 2006 / 353 ” (5, TAMU); same data except: “ IV- 28 – VI- 2 - 2006 / 392 /. Juniperus unmanaged plot ” (5, TAMU); “ UNITED STATES: TEXAS: Kerr Co. / 6.5 mi SW Hunt, 1960 ’ / 29 ° 59.409 ’ N, 99 ° 23.244 ’ W / IV- 28 – VI- 2.2006, FIT-grd. / E. G. Riley, et al. 405 / upland deciduous forest ” (7, TAMU); same data except: “ IX- 2 – X- 5 - 2006 / 561 ” (TAMU); 5, same data except: “ VI- 3 – 30 - 2006 / 444 ” (9, TAMU); same data except: “ X- 27 – XI- 11 - 2005 / 171 ” (1, TAMU); same data except “ III- 31 – IV- 27 - 2006 / 366 ” (TAMU); 1, “ TEXAS: Wood Co. / Godwin Woods, 3.5 mi. SW / Hainsville, V- 29 – VI- 27 - 2000 / 32 ° 42 ’ 30 ” N, 95 ° 24 ’ 36 ” W / Coll. Wm. Godwin, FIT ” (2, TAMU);, “ OKLAHOMA: / Latimer Co. / XI. 2002 / K. Stephan ” (1, TAMU); same data except “ XII. 2002 ” (1, TAMU).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEBB921FF6FAF77FA2FFDC1.taxon	distribution	Distribution. The species is known only from Oklahoma and Texas in the United States (Fig. 22). Seasonality. Adults have been collected from April to December, mostly in the months of April and May. Bionomics. Adults have been collected mostly in scrub and juniper thickets, and in upland mixed forest.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEBB921FF6FAF77FA2FFDC1.taxon	etymology	Etymology. The epithet “ bidentatus ” refers to the double teeth on the left mandible of this species.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFE8B921FF6FAC17FBFBF8A1.taxon	description	(Fig. 10, 22) Description. Length (pronotum + elytra) = 1.34 – 1.44 mm; greatest width = 1.00 – 1.04 mm. Reddish brown, shiny. Head punctation variable in size and spacing. Antennal grooves not well defined. Mandibles moderately long, left mandible with blunt tooth near middle. Antennal club slender; antennomere 8 disc-like, narrower than apex of 7; apical antennomere longer than and as wide as 10. Eyes of normal size. Pronotum broad, sides rounded; posterior angles rounded, barely evident; basal margin rounded. Pronotal punctures minute and sparse. Elytral strial punctures round and deep, variable spaced; interstrial punctures minute and sparse. Metasternum finely punctate medially, coarsely punctate laterally; lateral punctures separated by less than one diameter. Male mesofemur unmodified. Male mesotibia straight. Abdominal sternites III – VII each with basal transverse row of large punctures. Median lobe of aedeagus (Fig. 10) cylindrical and curved in basal half, with flat paired apices. Parameres narrow, not reaching apex of median lobe. Internal sac as in Fig. 10.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFE8B921FF6FAC17FBFBF8A1.taxon	materials_examined	Type material. Holotype, male, in SBPC, with label data: “ UNITED STATES: TX: Cameron Co., 3 m / 15 km se Brownsville, TNC / Southmost Preserve, N 25 ° 50.6 ’ / W 97 ° 22.9 ’, palm forest FITs / 1. III – 4. IV. 04, S. & J. Peck 04 – 54 ”. Paratypes, 7, as follows: same data as holotype (1, SBPC); “ TX: Cameron Co. / Brownsville, Audubon / Sabal Palm Grove / 31. V – 10. VIII. 83 / S. & J. Peck, FIT ” (1, SBPC); “ UNITED STATES: TX: ne Cameron Co. / Laguna Atascosa NWR / N 26 ° 14.0 W 97 ° 21.0. 2 m / Thorn thicket FIT, 1. III – 6. IV. 04 / S. & J. Peck, 04 – 50 ” (3, SBPC); “ FLA: Columbia Co. / O’Leno State Park / 16. VI – 8. VIII. 1981 / S. Peck, sand-oak- / pine, intercept tp. ” (1, SBPC); “ FLORIDA: Suwanee / Suwanee R. St. Pk. / 15. VI – 8. VIII. 1981 / S. Peck, mixed / forest intercept ” (1, SBPC).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFE8B921FF6FAC17FBFBF8A1.taxon	distribution	Distribution. The species is known only from Florida and Texas in the United States (Fig. 22). Its distribution in the extreme south of Texas suggests that the species may also occur in Mexico. The Florida populations seem to be disjunct from those in Texas. They may have once been connected across the Gulf Coastal Plain at times of Pleistocene low sea levels. This was a route from a western source for faunas moving into Florida, and is recognized for the movement of both vertebrates (Webb 1990) and arthropods (Deyrup 1989). Seasonality. Adults are known only from the months of March to August. Bionomics. Adults are known from mesic forests, thorn thicket, and palm forest, often on sandy soils. All were collected in flight intercept traps.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFE8B921FF6FAC17FBFBF8A1.taxon	etymology	Etymology. Latin “ exiguus ”, small, refers to the small size of this species.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFE8B920FF6FA8F7FAA0FE81.taxon	description	(Fig. 11, 22) Description. Length (pronotum + elytra) = 2.40 – 2.60 mm; greatest width = 1.48 – 1.72 mm. Reddish brown, shiny. Head punctures of moderate size, irregularly spaced, more dense laterally. Antennal grooves not well defined. Mandibles elongate, not forming semicircle when closed; both mandibles edentate, Antennal club slender; antennomere 8 disc-like, narrower than apex of 7; apical antennomere longer and narrower than 10. Eyes reduced in size, length about one-third length of head. Pronotum broad, sides weakly rounded, posterior angles obtuse, basal margin rounded. Pronotal punctures moderately small and dense. Elytral strial punctures round and deep, smaller toward base, separated by ± 1 diameter. Interstrial punctures fine and sparse; few large punctures between striae VI and VII. Metasternum finely punctate medially; coarsely, densely punctate laterally. Male mesofemur unmodified. Male mesotibia weakly concave on inner margin. Abdominal sternites III – VII each with basal transverse row of large punctures. Median lobe of aedeagus (Fig. 11) cylindrical, strongly curved dorsoventrally, flattening to paired apices in apical one-fourth. Parameres slender, sinuate apically, not reaching apex of median lobe. Internal sac as in Fig. 11.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFE8B920FF6FA8F7FAA0FE81.taxon	materials_examined	Type material. Holotype, male, in SBPC, with label data: “ N. MEX: Lincoln Co. / Sierra Blanca, 8000 ’ / 15 mi NW Ruidoso / 20. VIII. 75, S. Peck // Oak Flat Camp / Oak litter Ber 317 ”. Paratypes, 20, as follows: same data as holotype (5, SBPC); “ NMEX: Santa Fe Co. / 14 mi NE Santa Fe / 18. VI – 3. VII. 79, / S & J Peck, 9600 ’ / aspen w / stream ” (5, SBPC); “ ARIZ: 10 miNW Flagstaff / San Francisco Mts. 9500 ’ / 18 – 24. VII. 79, S & J Peck / spruce-fir-aspen / meadow, malaise ” (5, SBPC); “ ARIZ: Flagstaff / Oak Ck. Can. 5900 ’ / 17 – 25. VII. 79, at / Sterling Can. / riparian woods / malaise, S & J Peck ” (3, SBPC); “ ARIZ: Cochise Co. / Rustler Park, 9000 ’ / VIII- 31 - 1976 / Fred G. Andrews // Berlese soil / around Amanita / muscaria ” (1, CSCA); “ Chir. Mts. Ariz. / VIII- 16 - 68 7500 - / 9500 ’ Sweeping / V. Roth ” (1, CSCA).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFE8B920FF6FA8F7FAA0FE81.taxon	distribution	Distribution. The species is known only from Arizona and New Mexico in the United States (Fig. 22). Seasonality. Adults have been collected in the months of June to August. Bionomics. Adults have been collected in riparian forest, mixed forest, oak woodland, and upper elevation spruce-fir-aspen forest. They were taken mostly by flight intercept and malaise traps, and also in litter and by sweeping.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFE8B920FF6FA8F7FAA0FE81.taxon	etymology	Etymology. Latin “ parvoculus ”, small eyes, refers to the reduced eye size of this species.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFE9B920FF6FAED7FED4F881.taxon	diagnosis	Diagnosis. Body strongly convex. Antenna with 10 antennomeres with 4 - antennomere club. Mandibles together forming semicircle; left mandible with large tooth on basal one-third. Ventral side of head with distinct antennal grooves below the eyes. Mesosternum vertical between the mesocoxae, not carinate. Tibiae expanded, shovel-like, their outer margins with strong spines. Tarsal formula 5 - 5 - 4 in both sexes. Males distinguished by large tooth-like expansion on lower margin of metafemur.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFE9B920FF6FAED7FED4F881.taxon	distribution	Distribution. The distribution of the genus is worldwide except for polar and the Afrotropical regions (Newton 1998).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEEB926FF6FADD7FB9CFDE1.taxon	description	(Fig. 12, 23)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEEB926FF6FADD7FB9CFDE1.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.18 – 1.74 mm; greatest width = 0.86 – 1.28 mm. Yellowish to reddish brown, shiny. Head moderately coarsely, densely punctate. Antennal club darker than funicle; apical antennomere longer than and as wide as penultimate antennomere. Sides of pronotum rounded, posterior angles rounded; base rounded medially, weakly concave before posterior angles. Pronotal punctation fine and widely spaced. Elytral strial punctures deep and closely spaced, not clearly evident in basal one-fourth. Interstriae finely punctate. Metasternum coarsely, densely punctate; punctures finer medially. Metatibia widest from middle to apex, three times as wide as at base. Abdominal sternites III – VII each with transverse row of small punctures before apex. Median lobe of aedeagus (Fig. 12) broad, paired apices inwardly curved. Parameres extend to or slightly beyond apex of median lobe. Internal sac as in Fig. 12.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEEB926FF6FADD7FB9CFDE1.taxon	distribution	Distribution. North America (Fig. 23): CANADA. ONTARIO, QUEBEC. UNITED STATES. ALABAMA, DISTRICT OF COLUMBIA, FLORIDA, GEORGIA, ILLINOIS, INDIANA, KENTUCKY, MASSACHUSETTS, MARYLAND, MICHIGAN, NORTH CAROLINA, SOUTH CAROLINA, TEXAS, VIRGINIA, VERMONT. Previously recorded in North America (LeConte 1853, Zimmerman 1869, Brown 1937 b, Daffner 1988) from: CANADA. ONTARIO. Arnprior. Blackburn Hamlet. Leamington. Mer Bleue, near Ottawa. Ottawa. Port Rowan. Simcoe. QUEBEC. Aylmer. UNITED STATES: DISTRICT OF COLUMBIA. FLORIDA. Alachua Co., Gainesville. Dade Co., Everglades National Park, Long Pine Key. Sarasota Co., Myakka River State Park. Suwanee Co., Suwanee River State Park. GEORGIA. Habersham Co. KENTUCKY. 4 mi E Horse Cave. MASSACHUSETTS. Lincoln. Tyngsboro. MICHIGAN. Detroit. SOUTH CAROLINA. Oconee (not Greene) Co., Oconee National Forest. Seasonality. Adults have been collected from the months of April to October, with most in June to August, and early and late records are from more southern localities. Bionomics. Adults have been collected in wooded habitats ranging from mixed deciduous, open pineland, and warm temperate bottomlands to mangrove-hammock transition and oak-palm hammock. They have been taken most often by flight intercept traps, evening car netting and ultraviolet light traps. New material examined. We have seen 232 specimens from the following localities: CANADA. ONTARIO. Almonte. Chaffeys Locks, QUBS. Guelph. Kemptville Forest. Mer Bleue. Limoges, 40 km E Ottawa. Marlborough Forest, 15 km W North Gower. Pakenham Mountain road. Rideau Ferry to Portland. Rondeau Provincial Park. L 3 C 6 Wolford Township 44 ° 52 ’ 03 ” N, 75 ° 43 ’ 50 ” W. Tilbury. UNITED STATES. ALABAMA. Blount Co., 7 mi S Cleveland. Jefferson Co., Hoover. Vestavia. Morgan Co., 3.5 mi SW Fayette. FLORIDA. Dade Co., Long Pine Key. Highlands Co., Archbold Biology Station. Monroe Co., Big Pine Key. Sugarloaf Key, Kitchings Hammock. Sarasota Co., Myakka River State Park. ILLINOIS. Champaign Co., Allerton Park, Monticello. Mahomet, Nettie Hart Memorial Woods. 3 mi NW Homer. Cook Co., Chicago Heights. INDIANA. Monroe Co., Bloomington. Tippecanoe Co., MARYLAND. Montgomery Co., Potomac. Rockville. Talbot Co., Wittman. Worchester Co., 8 km WNW Snow Hill. 6 km SW Snow Hill. NORTH CAROLINA. Transylvania Co., Mt. Pisgah, Blue Ridge Parkway mile 414. SOUTH CAROLINA. Greenville Co., Greenville. Oconee Co., Oconee State Park. TEXAS. Aransas Co., 10 km SE Austwell. Brazos Co., College Station, Lick Creek Park. Fort Bend Co., Brazos Bend State Park. Montgomery Co., 4.5 mi N Montgomery. VIRGINIA. Fort Monroe. Fairfax Co., Burling Park. Warren Co., 4 km NNW Linden. VERMONT. Franklin Co., Bakersfield.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEFB926FF6FAC37FB62F9E1.taxon	description	(Fig. 13, 24)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEFB926FF6FAC37FB62F9E1.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.42 – 1.90 mm; greatest width = 1.00 – 1.42 mm. Light reddish brown, shiny. Head punctures variable in size and spacing. Antennal club usually not darker than funicle; antennomere 9 longer and wider than 8; apical antennomere narrower than 9, subequal in length. Sides of pronotum rounded, posterior angles rounded; base rounded medially, concave before posterior angles. Pronotal punctation fine and widely spaced. Elytral strial punctures deep and closely spaced; clearly visible almost to base. Interstriae finely punctate. Metasternum densely punctate with coarse punctures laterally, finer punctures medially. Metatibia evenly and strongly widened, widest at apex, there 3.5 times as wide as at the base. In males, abdominal sternite III with a pair of foveae at anterior margin; foveae usually bearing white setae; sternites IV – VII each with transverse row of small setose punctures near middle. Median lobe of aedeagus (Fig. 13) broad; paired apices curved, meeting apically. Parameres extend beyond apex of median lobe. Internal sac as in Fig. 13	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEFB926FF6FAC37FB62F9E1.taxon	distribution	Distribution. North America (Fig. 24). UNITED STATES. FLORIDA; only in the extreme south, in Dade and Monroe counties. Previously recorded (Daffner 1988) from: FLORIDA. Dade Co., Everglades National Park, Royal Palm Hammock. 1.5 km NW Royal Palm. S Miami, Deering Estate Park. Old Cutler Hammock. Monroe Co., Big Pine Key, Watsons Hammock. Big Torch Key. Cudjoe Key. Lignum Vitae Key. No Name Key. Sugar Loaf Key, Kitchings Hammock. Key Largo, Pennekamp State Park. Fat Deer Key. This distribution suggests that the species may also occur in Cuba or the Bahamas. Seasonality. Adults are known from the cooler months of October to March. Bionomics. All adults have been collected in subtropical hammock forests. They have been taken in flight intercept and malaise traps, except one in an ultraviolet light trap. New material examined. We have seen 152 specimens. There are no new records.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEFB925FF6FA837FEC3FF81.taxon	description	(Fig. 14, 24)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEFB925FF6FA837FEC3FF81.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.22 – 1.54 mm; greatest width = 0.90 – 1.20 mm. Reddish brown, shiny. Head punctures moderately coarse, irregularly spaced. Antennal club not or slightly darker than funicle; apical antennomere longer and narrower than penultimate antennomere. Sides of pronotum rounded, posterior angles rounded; base rounded medially, weakly concave before posterior angles. Pronotal punctation fine, irregularly spaced. Elytral strial punctures deep and closely spaced, discernible nearly to base. Interstriae finely punctate. Metasternum coarsely, densely punctate laterally, more finely punctate medially. Metatibia widest in apical half, 2.5 times as wide as at the base. Abdominal sternites III – VII each with transverse row of fine punctures near middle. Median lobe of aedeagus (Fig. 14) broad, paired apices inwardly curved. Parameres extend slightly beyond apex of median lobe. Internal sac as in Fig. 14.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEFB925FF6FA837FEC3FF81.taxon	distribution	Distribution. North America (Fig. 24). UNITED STATES. ALABAMA, ARKANSAS, FLORIDA, OKLAHOMA, TEXAS, VIRGINIA. Previously recorded in North America (Daffner 1988) from: UNITED STATES. OKLAHOMA. Latimer Co., 5 mi W Red Oak. Seasonality. Adults have been collected from April to November, with most being in the months of June to September. Bionomics. Adults have been collected mostly in mixed hardwood forest. They have been taken by flight intercept or ultraviolet light or yellow pan traps. New material examined. We have seen 48 specimens from the following new localities: UNITED STATES. ALABAMA. Jefferson Co., Westavia. ARKANSAS. Little Rock. FLORIDA. Alachua Co., Gainesville. Monroe Co., Middle Torch Key. OKLAHOMA. Marshall Co., Willis, Lake Texoma. TEXAS. Angelina Co., 4 mi SE Zavalla, Angelina National Forest. Bastrop Co., Bastrop State Park. Tyler Co., Kirby State Forest. Wood Co., 3.5 mi SW Hainesville, Godwin Woods. VIRGINIA. Bath Co., 9.6 mi N Clifton Forge.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFECB925FF6FAFD7FD1AFA01.taxon	description	(Fig. 15, 24)	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFECB925FF6FAFD7FD1AFA01.taxon	diagnosis	Diagnosis. Length (pronotum + elytra) = 1.40 – 2.02 mm; greatest width = 1.00 – 1.68 mm. Reddish brown, shiny. Head punctures moderately fine, variably spaced. Antennal club of same color as funicle; apical antennomere longer and narrower than penultimate antennomere. Sides of pronotum rounded, posterior angles rounded; base rounded medially, weakly concave before posterior angles. Pronotal punctation fine, widely spaced. Elytral strial punctures deep and closely spaced, not discernible near base. Interstriae finely, sparsely punctate. Metasternum finely, sparsely punctate medially; coarsely, irregularly punctate laterally. Metatibia widest just before apex, 3 to 4 times as wide as at base. Abdominal sternites III – VII each with transverse row of fine punctures before apex. Median lobe of aedeagus (Fig. 15) broad, paired apices short and broad. Parameres broad, not extending beyond apex of median lobe. Internal sac as in Fig. 15.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFECB925FF6FAFD7FD1AFA01.taxon	distribution	Distribution. North American distribution (Fig. 24): UNITED STATES. FLORIDA; only in the extreme south in Dade and Monroe counties. Previously recorded in North America (Daffner 1988) from: UNITED STATES. FLORIDA. Dade Co., S Miami, Deering Estate Park. Everglades National Park, Royal Palm Hammock. Monroe Co., Big Pine Key, Watson Hammock. Cudjoe Key. Fat Deer Key. Key Largo, Pennekamp State Park. Lignum Vitae Key, near Islamorada. Sugarloaf Key, Kitchings Hammock. Seasonality. Adults have been collected throughout the year, but mostly in the cooler months from October to February. Bionomics. All collections are from subtropical hammock forest in extreme southern Florida. All have been taken by flight intercept and ultraviolet light traps. New material examined. We have seen 132 specimens, including the following new localities: UNITED STATES. FLORIDA. Dade Co. Old Cutler Hammock, 7900 SW 176 th St. Everglades National Park, Long Pine Key. Monroe Co. Big Pine Key, No Name Road. Big Torch Key. Key Vaca. Middle Torch Key. No Name Key. Stock Island. Sugarloaf Key, SE ¼, Section 23. This distribution suggests that the species may also occur in Cuba or the Bahamas.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEDB924FF6FA83CFB10FA6C.taxon	description	(Fig. 21) A nisotoma ecarinata Horn 1880: 292. “ Lectotype ”, female, in MCZC, type number 3012, (We do not know of a publication validating this specimen as the lectotype and we do not so designate it here.), seen; MCZC cotype 33560, seen. Type locality: “ western Nevada, ” probably from woodland or forest habitat in the vicinity of Reno or Lake Tahoe.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEDB924FF6FA83CFB10FA6C.taxon	diagnosis	Diagnosis. As for the genus.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEDB924FF6FA83CFB10FA6C.taxon	distribution	Distribution. Previously recorded in North America (Horn 1880, Hatch 1929, 1957) from: CANADA. BRITISH COLUMBIA. Penticton. UNITED STATES. NEVADA (unspecified locality). CALIFORNIA. See Fig. 21. Seasonality. The few known specimens were collected from May to July. Bionomics. The few known adults with habitat data were collected in mixed forest.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEDB924FF6FAC8AFDFDF818.taxon	diagnosis	Diagnosis. Body moderately flattened, oblong. Antenna with 11 antennomeres, with 5 - antennomere club interrupted by small antennomere 8. Mandibles moderately long, without distinct teeth. Ventral side of head without antennal grooves. Pronotum transverse, about as wide as elytra. Elytra longer than wide. Prosternum short in front of procoxae. Mesosternum oblique, without median longitudinal carina. Metasternum longer than first two visible abdominal sternites combined. Abdomen with 6 visible sternites (sternites III – VIII) in both sexes. Legs moderately slender; all tibiae spinose on outer margin. Tarsal formula 5 - 5 - 4 in both sexes. Males distinguished by weakly expanded protarsi and mesotarsi, and by toothlike expansion of lower margin of metafemur. The distribution of the genus is western Nearctic (Newton 1998).	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEDB924FF6FAC8AFDFDF818.taxon	discussion	Discussion. We have examined five previously unreported specimens of Ecarinosphaerula. The label data of the specimens are as follows: “ Pasadena / Cal. // H. C. Fall Collection ” (1 female, MCZC); “ UNITED STATES: Cal., Tulare Co. / Sequoia Nat. Park, 7000 ’ / Halstead Creek / 29. V. 1984 / R. Baranowski ” (1 male, SBPC); “ CA: Monterey Co. / 36 ˚ 04.33 ’ N, 121 ˚ 34.95 ’ W / UC Big Creek Reserve / vi. 29 - vii. 6.2002 / M. Caterino, FIT (11) ” (2 males, SBMN); “ UNITED STATES: CA: Marin Co., 2 kmS / Olema, N 38 ˚ 00.5 ’ W 122 ˚ 46 ’ / Mixed ravine forest FIT, 20 m / 9. V- 11. VI. 03, S. Peck, 03 - 20 ” (2 males, SBPC). There are at least two distinct species among these specimens, based on examination of male genitalia. The original description of the species E. ecarinata (Horn 1880: 292) was based on two specimens from western Nevada, with a label with the word “ Morrison ”, whom we assume was the collector, and which is not a locality. One of these specimens appears to be a female and bears a label as being the (unpublished?) lectotype (MCZC type 3012). An additional specimen, not seen by us, was reported by Hatch (1957) as E. ecarinata from Penticton, British Columbia. At this time, with the paucity of material of this genus, we cannot provide more information.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
146C8794FFEDB924FF6FADD7FE06FC6D.taxon	discussion	This group is characterized by having the mesosternum oblique between the mesocoxae. In North America it is known to contain only the genera Leiodes Latreille and Ecarinosphaerula Hatch. Baranowski (1993) revised the North American species of Leiodes. We here contribute new data on the genus Ecarinosphaerula Hatch.	en	Peck, Stewart B., Cook, Joyce (2013): Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini). Insecta Mundi 2013 (310): 1-32, DOI: 10.5281/zenodo.5193714
