taxonID	type	description	language	source
19120E4BB65D0A0345874BC6FDE3C4CC.taxon	description	(Figure 2) The following description refers to one voucher specimen in developmental stage 41 (ZSM 525 / 2004, field number LR 68, TL 42.5 mm, BL 14.0 mm), from locality 1. In dorsal view (Figure 2 a), body ovoid elongate, snout rounded. In lateral view (Figure 2 b), body depressed, BW 131 % of BH, snout rounded. Eyes moderately large, ED 14.0 % of BL, bulging, not visible in ventral view, positioned dorsolaterally but directed slightly more laterally than anterolaterally or dorsolaterally, situated at about the anterior quarter of the body. Nares elliptical, rimmed with a mediodorsal projection giving them a bean shape, moderately small, positioned dorsolaterally, directed anterodorsally and about equidistant between tip of snout and anterior edge of eyes, RN 106 % of NP; NN 53 % of PP. Spiracle sinistral, moderately sized, tubular, well visible from dorsal view, distal half free from body, spiracular opening orientated posterodorsally, closer to end of body than to snout, SS 65 % of BL, situated at the height of the lower part of tail musculature. Vent tube moderately sized, short, dextral, opening dextral at ventral edge of fin, its right wall displaced anterodorsally, directed posterolaterally, linked to ventral tail fin except its tip. Tail long and tapered. Caudal musculature strong, TMH 64 % of BH and 87 % of MTH, TMW 46 % of BW, at mid-length of tail its height is equal to half of total tail height, reaching tail tip. Caudal fins regular with straight edges, moderately weakly developed, their general appearance almost similar throughout tail musculature, point of maximum height attained in the second half of the tail, MTH 74 % of BH; dorsal fin originating at the dorsal tail – body junction and slightly taller than ventral fin at mid-length of tail; tail tip bluntly pointed, its state as reproduced in Figure 2 (caudal fins not reaching tail tip) probably being an artefact of preservation or damage during transport. Oral disc (Figure 2 c) of moderate size, ODW 21 % of BL and 38 % of BW, positioned and directed ventrally, emarginated. A single row of marginal papillae interrupted by a large median gap on the upper labium (DG 73 % of ODW) and by a small medial gap on the lower labium; total number of marginal papillae about 81 (43 on the left side and 38 on the right side). A few submarginal papillae positioned in the lateral parts of anterior and posterior labia. Papillae moderately large, cylindrical with rounded or pointed tips. No denticulate papillae. Keratodont row formula 1: 3 + 3 / 1 + 1: 2. The density of keratodonts on A 1 is about 55 per mm (a total of ca 144). The length of interrupted anterior keratodont rows (A 2, A 3, and A 4) decreases gradually towards the centre of the disc, P 1 and P 2 are of about similar size, slightly longer than P 3. Both jaw sheaths coarsely serrated; upper jaw completely black with a large medial serration surrounded by smaller serrations on each side forming a slight convexity; lower jaw partially pigmented, strong and V-shaped. Coloration in preservative. Dorsally: whole body and caudal musculature dark brown and covered by scattered black dots; dark pigments concentrated dorsally forming dark patches slightly visible on the interorbital area and beside the nares; a dark line slightly visible middorsally on body. Laterally: intestinal coils well visible; body and caudal musculature with scattered dark pigmentation; dorsal fin transparent with a few sparse black dots; ventral fin clear. Ventrally: branchial and cardial region opaque through ventral body wall; intestinal coils well visible. Variation. TL and BL of two tadpoles at stage 41 (ZSM 525 / 2004 – 526 / 2004) from locality 1 are 36.5 – 42.5 and 13.0 – 14.0 mm, respectively. The ratios vary in the following proportions: BW 124 – 131 % of BH; ED 13.1 – 13.6 % of BL; RN 106 – 121 % of NP; NN 53 – 57 % of PP; SS 63 – 65 % of BL; TMH 64 – 65 % of BH; TMH 64 – 71 % of MTH; TMW 46 – 51 % of BW; MTH 90 – 102 % of BH; ODW 20 – 21 % of BL; ODW 38 – 39 % of BW. KRF of the two tadpoles is 1: 3 + 3 / 1 + 1: 2.	en	Raharivololoniaina, Liliane, Grosjean, Stéphane, Raminosoa, Noromalala Rasoamampionona, Glaw, Frank, Vences, Miguel (2006): Molecular identification, description, and phylogenetic implications of the tadpoles of 11 species of Malagasy treefrogs, genus Boophis. Journal of Natural History 40 (23 - 24): 1449-1480, DOI: 10.1080/00222930600902399, URL: http://dx.doi.org/10.1080/00222930600902399
19120E4BB65F0A05459C4C01FBFBC40F.taxon	description	(Figure 3) The following description is based on one tadpole in developmental stage 35 (ZSM 519 / 2004, field number LR 5, TL 34.7 mm, BL 13.7 mm), from locality 5. In dorsal view (Figure 3 a), body ovoid elongate, snout rounded. In lateral view (Figure 3 b), body depressed, BW 117 % of BH, snout rounded. Eyes of moderate size, ED 11.0 % of BL, very slightly bulging, not visible in ventral view, positioned dorsolaterally but directed almost laterally, situated at about the anterior third of the body. Pineal ocellus in line with the anterior margin of eyes. Nares moderately sized, elliptical, rimmed with a flat mediodorsal projection giving them a bean shape, positioned dorsolaterally, directed anterolaterally and slightly dorsally, nearer to anterior edge of eyes than to snout, RN 109 % of NP; NN 55 % of PP. Spiracle sinistral, moderately sized, well visible from dorsal view, oval, its tip free from body; spiracular opening orientated posteriorly, closer to end of body than to snout, SS 66 % of BL, situated at the height of the lower part of tail musculature. Vent tube moderately sized, short, dextral, opening dextral at ventral edge of fin, its right wall displaced anterodorsally, directed posteriorly. Caudal musculature moderately strong, TMH 54 % of BH and 58 % of MTH, TMW 49 % of BW, at mid-length of tail its height is two-fifths of the total tail height, reaching tail tip. Caudal fins regular with straight edges, MTH 93 % of BH; dorsal fin low, originating on tail muscle and wider than ventral fin; tail tip pointed. Oral disc (Figure 3 c) of moderate size, ODW 26 % of BL and 43 % of BW, positioned and directed anteroventrally, emarginated. A single row of marginal papillae interrupted by a large median gap on the upper labium (DG 81 % of ODW) and without visible gap on the lower labium; total number of marginal papillae about 95. Submarginal papillae present, some of them clustered at the corners of the disc but most localized on posterior labium and distributed in two rows. Papillae of moderate size, elongate with rounded tip. No denticulate papillae. Keratodont row formula 1: 5 + 5 / 1 + 1: 2. The density of keratodonts on A 1 is about 60 per mm (a total of 164). The length of interrupted anterior keratodont rows (A 2 – A 6) decreases gradually towards the centre of disc, P 1 interrupted by a gap of less than 0.1 mm, keratodont rows of lower labium subequal in size. Both jaw sheaths completely dark pigmented; upper jaw a wide flattened, medially convex arch, finely serrated; lower jaw V-shaped, moderately finely serrated. Coloration in preservative. Dorsally: body grey with sparse dark markings, caudal musculature with scattered dark pigmentation. Laterally: intestinal coils visible; fins translucent with sparse black dots. Ventrally: branchial and cardial region not visible through ventral body wall; intestinal coils obscure. Variation. TL and BL of six tadpoles at stages 28 – 37, all from locality 5 (ZSM 517 – 520, and LR 4, LR 17) are 24.5 – 35.6 and 10.8 – 15.6 mm, respectively. The ratios vary in the following proportions: BW 117 – 157 % of BH; ED 10.0 – 12.2 % of BL; RN 109 – 207 % of NP; NN 54 – 63 % of PP; SS 61 – 69 % of BL; TMH 54 – 71 % of BH; TMH 58 – 84 % of MTH; TMW 43 – 58 % of BW; MTH 67 – 93 % of BH; ODW 26 – 29 % of BL; ODW 42 – 57 % of BW. KRF of the six tadpoles varies from 1: 4 + 4 / 1 + 1: 2 to 1: 5 + 5 / 1 + 1: 2.	en	Raharivololoniaina, Liliane, Grosjean, Stéphane, Raminosoa, Noromalala Rasoamampionona, Glaw, Frank, Vences, Miguel (2006): Molecular identification, description, and phylogenetic implications of the tadpoles of 11 species of Malagasy treefrogs, genus Boophis. Journal of Natural History 40 (23 - 24): 1449-1480, DOI: 10.1080/00222930600902399, URL: http://dx.doi.org/10.1080/00222930600902399
19120E4BB6590A1945B44BA7FB9BC665.taxon	description	(Figure 4) The following description refers to one voucher specimen in developmental stage 33 (ZSM 588 / 2004, field number LR 236, TL 44.3 mm, BL 16.1 mm), from locality 3. In dorsal view (Figure 4 a), body elliptical, snout rounded. In lateral view (Figure 4 b), body depressed, BW 116 % of BH, snout rounded. Eyes moderately large, ED 13.0 % of BL, bulging, not visible in ventral view, positioned dorsolaterally but directed anterolaterally and dorsolaterally, situated at about the anterior quarter of the body. Pineal ocellus placed anteriorly to margin of eyes. Nares round, moderately sized, rimmed, positioned dorsolaterally, directed anterolaterally and opening dorsolaterally, positioned nearer to tip of snout than to anterior edge of eyes, RN 86 % of NP; NN 44 % of PP. Spiracle sinistral, moderately small, slightly conical, slightly visible from dorsal view, free at tip, orientated posterodorsally; spiracular opening oval, situated closer to end of body than to tip of snout, SS 58 % of BL, situated below the longitudinal axis of tail musculature. Vent tube short, dextral, opening dextral, its right wall displaced anterodorsally, directed posterolaterally, entirely linked to ventral tail fin. Caudal musculature moderately strong, TMH 55 % of BH and 62 % of MTH, TMW 51 % of BW, at mid-length of tail its height about two-fifths of the total tail height, reaching tail tip. Caudal fins moderately developed, point of maximum height attained at the proximal third of the tail, MTH 89 % of BH; dorsal fin convex, originating at the dorsal tail – body junction and taller than ventral fin at mid-length of tail, ventral fin originating immediately behind body; tail tip pointed. Oral disc (Figure 4 c) moderately large, ODW 25 % of BL and 47 % of BW, positioned and directed ventrally, emarginated. Oral disc surrounded by a single row of 69 (34 on the left side and 35 on the right side) marginal papillae interrupted by a large median gap on the upper labium (DG 71 % of ODW), and by a small medial gap on the lower labium. A few submarginal papillae positioned on the lateral parts of the upper labium, and lateral and posterolateral parts of the lower labium. Papillae of moderate size, cylindrical with a rounded tip. No denticulate papillae. Keratodont row formula 1: 5 + 5 / 1 + 1: 2. Gap on P 1 shorter than 0.1 mm; about 188 keratodonts on A 1, the longest keratodont row of the oral disc (ca 61 per mm); P 1 and P 2 of about similar size, slightly longer than P 3. Upper jaw completely black, moderately serrated, a large arch with a large median convexity; lower jaw V-shaped, coarsely serrated, mostly coloured in black but its base less pigmented. Coloration in preservative. Dorsally: body and tail musculature mottled with black stippling; dark pigmentation concentrated dorsally forming dark patches between eyes and next to nares. Laterally: intestinal coils well distinct; dorsal fin with scattered black dots; ventral fin generally clear except near tail tip; at about mid-tail many small dark patches visible along the upper tail musculature. Ventrally: whole surface of ventral body transparent; branchial and cardial organs translucent but slightly visible through ventral body wall. Variation. TL and BL of two tadpoles at stages 33 and 41, from locality 3 (ZSM 588 / 2004 – 589 / 2004) are 43.5 – 44.3 and 15.5 – 16.1 mm, respectively. The ratios are: BW 116 – 140 % of BH; ED 13.0 – 14.8 % of BL; RN 86 – 113 % of NP; NN 44 – 49 % of PP; SS 58 – 70 % of BL; TMH 55 – 70 % of BH; TMH 62 – 63 % of MTH; MTH 89 – 112 % of BH; ODW 24 – 25 % of BL; ODW 44 – 47 % of BW. KRF of the two tadpoles varies from 1: 4 + 4 / 1 + 1: 2 to 1: 5 + 5 / 1 + 1: 2.	en	Raharivololoniaina, Liliane, Grosjean, Stéphane, Raminosoa, Noromalala Rasoamampionona, Glaw, Frank, Vences, Miguel (2006): Molecular identification, description, and phylogenetic implications of the tadpoles of 11 species of Malagasy treefrogs, genus Boophis. Journal of Natural History 40 (23 - 24): 1449-1480, DOI: 10.1080/00222930600902399, URL: http://dx.doi.org/10.1080/00222930600902399
19120E4BB6590A1945B44BA7FB9BC665.taxon	description	The following description refers to one specimen in developmental stage 31 (ZSM 580 / 2004, field number LR 231 a, TL 41.3 mm, BL 15.0 mm), from locality 3. In dorsal view (Figure 5 a), body ovoid elongate, snout rounded. In lateral view (Figure 5 b), body depressed, BW 117 % of BH, snout rounded. Eyes of moderate size, ED 11.3 % of BL, bulging, not visible in ventral view, positioned anterolaterally but directed anterolaterally and dorsolaterally. Nares round, moderately small, rimmed, positioned dorsally, directed anterolaterally, nearer to anterior edge of eyes than to tip of snout, RN 123 % of NP; NN 40 % of PP. Spiracle sinistral, moderately small, square, well visible from dorsal view, free at tip, orientated almost posteriorly; spiracular opening oval, situated closer to end of body than to tip of snout, SS 64 % of BL, situated at the height of the longitudinal axis of tail musculature. Vent tube short, dextral, opening dextral, its right wall displaced anterodorsally, directed posterolateroventrally. Caudal musculature moderately developed, TMH 50 % of BH and 60 % of MTH, TMW 40 % of BW, at mid-length of tail its height about two-fifths of the total tail height. Caudal fins moderately weakly developed, MTH 83 % of BH; dorsal fin originating at the dorsal tail – body junction, ventral fin originating immediately behind body; tail tip nearly rounded. Oral disc (Figure 5 c) of moderate size, ODW 20 % of BL and 37 % of BW, positioned and directed ventrally, emarginated. A single row of marginal papillae interrupted by a large median gap on the upper labium (DG 53 % of ODW), and by a very small medial gap on the lower labium. A few submarginal papillae positioned in the lateral parts of the upper and lower labia. Papillae stout, of moderate size, conical with a rounded tip. No denticulate papillae. Keratodont row formula 1: 3 + 3 / 1 + 1: 2. About 92 keratodonts on A 1 (ca 54 per mm). P 1 interrupted by a gap of less than 0.1 mm and slightly longer than P 2; P 3 about two-thirds of P 2. Upper jaw completely black, a large arch with a slight median convexity, moderately coarsely serrated; lower jaw strong, V-shaped, coarsely serrated. Coloration in preservative. Dorsally: the whole dorsum body and the caudal musculature with dark scattered pigmentation. Laterally: intestinal coils completely visible; lateral body and caudal musculature covered with sparse black dots; caudal fins moderately pigmented. Ventrally: the whole surface of ventral body transparent; branchial and cardial region visible through ventral body wall. Variation. TL and BL of 11 tadpoles at stages 31 – 41 (ZSM 580 / 2004 – 587 / 2004, and LR 231 c, LR 231 d, LR 231 i), all from locality 3, are 30.3 – 43.4 and 13.1 – 16.2 mm, respectively. The ratios vary in the following proportions: BW 111 – 133 % of BH; ED 10.7 – 12.6 % of BL; RN 113 – 155 % of NP; NN 40 – 47 % of PP; SS 59 – 67 % of BL; TMH 50 – 59 % of BH; TMH 46 – 58 % of MTH; TMW 35 – 44 % of BW; MTH 89 – 109 % of BH; ODW 17 – 21 % of BL; ODW 29 – 37 % of BW. KRF of the 11 tadpoles is 1: 3 + 3 / 1 + 1: 2.	en	Raharivololoniaina, Liliane, Grosjean, Stéphane, Raminosoa, Noromalala Rasoamampionona, Glaw, Frank, Vences, Miguel (2006): Molecular identification, description, and phylogenetic implications of the tadpoles of 11 species of Malagasy treefrogs, genus Boophis. Journal of Natural History 40 (23 - 24): 1449-1480, DOI: 10.1080/00222930600902399, URL: http://dx.doi.org/10.1080/00222930600902399
19120E4BB6450A1C4571495EFE27C1B2.taxon	description	(Figure 6) The following description is based on one tadpole in developmental stage 41 (ZSM 523 / 2004, field number LR 66, TL 26.5 mm, BL 10.0 mm), from locality 1. Hindlimbs of this specimen were largely used for tissue sampling and are therefore not shown in Figure 6. In dorsal view (Figure 6 a), body ovoid elongate, snout rounded. In lateral view (Figure 6 b), body greatly depressed, BW 145 % of BH, snout almost truncate. Eyes large, ED 16.0 % of BL, slightly bulging, not visible in ventral view, positioned dorsolaterally but directed almost laterally, situated at about the anterior third of the body. Nares moderately small, round, rimmed with a very small and flat mediodorsal projection, positioned dorsally, directed anterolaterally and dorsally, much closer to anterior edge of eyes than to snout, RN 169 % of NP; NN 44 % of PP. Spiracle sinistral, narrow and long, slightly visible from dorsal view, its tip free from body; spiracular opening orientated posterodorsally, much closer to end of body than to snout, SS 76 % of BL, situated at the height of the lower part of tail musculature. Vent tube small, short, medial, opening just before the beginning of ventral fin, opening medial, directed more posteriorly than posteroventrally, linked to caudal muscle. Caudal musculature strong, TMH 66 % of BH and 59 % of MTH, TMW 48 % of BW, especially in the anterior half of the tail; height of caudal musculature almost half of total tail height at mid-length of tail, almost reaching tail tip. Caudal fins regular with straight edges, very shallow, MTH 111 % of BH; dorsal fin originating next to body – tail junction, convex in its medial part; ventral fin beginning at the level of the ventral terminal of the body, following the caudal muscle; tail tip fine. Oral disc (Figure 6 c) enlarged, ODW 41 % of BL and 63 % of BW, positioned and directed ventrally, not emarginated. Several uninterrupted rows of marginal papillae around oral disc; no medial gaps in rows of marginal papillae on upper or lower labium. Papillae of internal row round and of moderate size, papillae of external rows small, elongate, and cylindrical. No denticulate papillae. Keratodont row formula 4: 3 + 3 / 3. About 203 keratodonts on A 3 (32 – 33 per mm). The length of interrupted anterior keratodont rows (A 5, A 6, and A 7) decreases gradually towards the centre of the disc, keratodont rows of lower labium subequal in length. Upper jaw not serrated and weakly developed, median part straight, a black halo on its distal part; lower jaw more developed, narrow V-shaped, ribbed, composed of two parts connected by a less keratinized median area, coarsely serrated. Coloration in preservative. Dorsally: part anterior to eyes covered with sparse black dots; dark pigmentations concentrated dorsally forming dark patches between eyes and beside nares; black colour on either side of lateral body obscuring intestine area; caudal musculature with scattered dark pigmentation. Laterally: intestinal coils invisible; caudal musculature with sparse black spots, showing some reticulations at its posterior end; dorsal fin with small black spots, ventral fin clear. Ventrally: branchial and cardial region translucent, well visible through ventral body wall; intestinal coils invisible. Variation. TL and BL of nine tadpoles at stages 29 – 41 (ZSM 521 / 2004 – 524 / 2004, and LR 61, LR 62, LR 64, LR 65, LR 67), all from locality 1, are 22.1 – 27.2 and 8.1 – 10.7 mm, respectively. The ratios vary in the following proportions: BW 118 – 145 % of BH; ED 14.9 – 16.8 % of BL; RN 143 – 196 % of NP; NN 42 – 49 % of PP; SS 70 – 81 % of BL; TMH 65 – 76 % of BH; TMH 59 – 84 % of MTH; TMW 48 – 67 % of BW; MTH 86 – 111 % of BH; ODW 36 – 47 % of BL; ODW 63 – 86 % of BW. KRF of the nine tadpoles is 4: 3 + 3 / 3. Boophis albipunctatus group Boophis sibilans Glaw and Thiesmeier, 1993 (Figure 7) The following description is based on one tadpole in developmental stage 25 (ZSM 557 / 2004, field number LR 269, TL 25.2 mm, BL 9.0 mm), from locality 4. In dorsal view (Figure 7 a), body elliptical, snout rounded. In lateral view (Figure 7 b), body depressed, BW 129 % of BH, snout rounded. Eyes moderately large, ED 12.2 % of BL, bulging, not visible in ventral view, positioned more dorsally than dorsolaterally but directed dorsolaterally, situated at about the anterior third of the body. Nares moderately sized, elliptical, rimmed with a flat mediodorsal projection, positioned dorsally, directed anterolaterally and opening almost dorsally, much closer to anterior edge of eyes than to snout, RN 148 % of NP; NN 50 % of PP. Spiracle sinistral, narrow and long, slightly visible from dorsal view, its tip free from body; spiracular opening orientated almost posteriorly, much closer to end of body than to snout, SS 72 % of BL, situated below the lower part of tail musculature. Vent tube small, medial, directed posteriorly, linked to caudal muscle, opening posterolateral. Caudal musculature strong, TMH 77 % of BH and 75 % of MTH, TMW 59 % of BW, at mid-length of tail height of caudal musculature almost half of total tail height, parallel in its proximal third then gradually tapering, almost reaching tail tip. Caudal fins moderately shallow, MTH 103 % of BH; dorsal fin originating next to body – tail junction, shallow in its anterior part then becoming convex towards mid-tail, ventral fin beginning just behind body and reaching its maximum height more posteriorly than dorsal fin; tail tip rounded. Oral disc (Figure 7 c) enlarged, ODW 44 % of BL and 89 % of BW, positioned and directed ventrally, not emarginated. Several rows of marginal papillae around oral disc, interrupted by a large median gap on the upper labium (DG 53 % of ODW); no gap on lower labium. Papillae small, conical with a more or less pointed tip. No denticulate papillae. Keratodont row formula 4: 3 + 3 / 3. Keratodonts on continuous rows A 1, A 2, and A 3 very small and difficult to count; estimation of keratodonts done on A 4 with a total of about 208 (ca 70 per mm). The length of interrupted upper keratodont rows (A 5, A 6, and A 7) decreases gradually towards the centre of the disc, keratodont rows of lower labium subequal in length. Upper jaw not serrated and weakly developed, black, median part straight; lower jaw more developed, V-shaped, ribbed, coarsely serrated, partially pigmented. Coloration in preservative. Dorsally: body and tail musculature brownish with scattered dark pigmentation; either side of dorsal body black coloured with white stripes (the blood vessels); some dark pigment concentrated dorsally forming dark patches between eyes and beside nares; tail muscle barred black and white. Laterally: intestinal coils not visible through lateral body wall; spiracle not pigmented; caudal fins clear; lower part of caudal musculature less pigmented anteriorly; musculature junctions very distinct anteriorly. Ventrally: opaque branchial and cardial organs slightly visible through ventral body wall; intestinal coils not visible. Variation. TL and BL of eight tadpoles at stage 25 (ZSM 556 / 2004 – 560 / 2004, and LR 269 a, LR 269 c, LR 269 f) from localities 2 and 4, are 19.1 – 25.2 and 6.9 – 9.0 mm, respectively. The ratios vary in the following proportions: BW 118 – 132 % of BH; ED 11.8 – 13.0 % of BL; RN 127 – 177 % of NP; NN 46 – 53 % of PP; SS 70 – 78 % of BL; TMH 59 – 77 % of BH; TMH 59 – 75 % of MTH; TMW 54 – 61 % of BW; MTH 97 – 113 % of BH; ODW 44 – 53 % of BL; ODW 88 – 96 % of BW. KRF of the eight tadpoles is 4: 3 + 3 / 3. Boophis luteus group Boophis luteus (Boulenger, 1882) (Figure 8) The following description is based on one tadpole in developmental stage 33 (ZSM uncatalogued, field number LR 218, TL 33.1 mm, BL 13.1 mm), from locality 3. In dorsal view (Figure 8 a), body ovoid, snout rounded. In lateral view (Figure 8 b), body depressed, BW 125 % of BH, snout rounded. Eyes moderately large, ED 14.5 % of BL, bulging, not visible in ventral view, positioned dorsolaterally but directed laterally, situated at about one-third of the body. Nares moderately sized, nearly oval, not rimmed, positioned and directed dorsally, closer to the anterior edge of eyes than to tip of snout, RN 133 % of NP; NN 64 % of PP. Spiracle sinistral, moderately small, slightly tubular, well visible from dorsal view, its tip free from body; spiracular opening oval, orientated posterodorsally, positioned at about three-quarters of body length, SS 76 % of BL, and situated well below the longitudinal axis of tail musculature. Vent tube short, dextral, opening at ventral edge of fin, opening directed posterolaterally, linked to caudal muscle, its right wall displaced anteriorly. Caudal musculature moderately strong, TMH 54 % of BH and 65 % of MTH, TMW 48 % of BW, height of caudal musculature about two-fifths of the total height at mid-tail, reaching tail tip. Caudal fins shallow anteriorly, deepest at about half of their length, their height decreasing progressively up to tail tip, MTH 82 % of BH; dorsal fin originating well posterior to the dorsal tail – body junction, ventral fin starting just behind the body; tail tip obtuse. Oral disc (Figure 8 c) large, ODW 37 % of BL and 63 % of BW, positioned anteroventrally and directed ventrally, not emarginated. Oral disc bordered by one or two rows of marginal papillae interrupted by a large median gap on the upper labium (DG 94 % of ODW), and by a small median gap on the lower labium. Submarginal papillae clustered in the corners of labia. Papillae of moderate size, cylindrical elongate with rounded tip. No denticulate papillae. Keratodont row formula 1: 5 + 5 / 1 + 1: 2. About 238 keratodonts on A 1 (ca 63 per mm). The length of the interrupted anterior rows A 2, A 3, and A 4 decreases towards centre of disc; keratodont rows of lower labium subequal in size. Both jaw sheaths serrated and fully black pigmented; upper jaw a wide flattened arch convex medially; lower jaw Vshaped. Coloration in preservative. Dorsally: body and tail musculature dark pigmented; dark pigmentation concentrated dorsally forming obvious dark patches between eyes and immediately next to nares. Laterally: intestinal coils well visible; dorsal fin pigmented; ventral fin almost clear, musculature mottled. Ventrally: branchial and cardial organs apparent through the transparent surface of body. Variation. TL and BL of 28 tadpoles at stages 25 – 41 (LR 160, LR 189, LR 227 f, LR 227 g, LR 227 h, LR 227 i, LR 227 j, LR 227 l, LR 227 m, LR 238, LR 238 a, LR 267, LR 267 a, LR 72, LR 73, LR 227 e, LR 227 k, LR 227 n, LR 227 q, LR 218, LR 227 b, LR 24, LR 227 o, LR 227 p, LR 227 a, LR 227 c, LR 227 d, LR 250), from localities 1 – 5, are 17.7 – 41.4 and 6.7 – 16.1 mm, respectively. The ratios vary in the following proportions: BW 121 – 145 % of BH; ED 10.0 – 16.7 % of BL; RN 100 – 182 % of NP; NN 40 – 64 % of PP; SS 58 – 88 % of BL; TMH 54 – 69 % of BH; TMH 58 – 76 % of MTH; TMW 42 – 65 % of BW; MTH 84 – 113 % of BH; ODW 26 – 44 % of BL; ODW 49 – 75 % of BW. KRF of the 28 tadpoles varies from 1: 3 + 3 / 1 + 1: 2 to 1: 5 + 5 / 1 + 1: 2.	en	Raharivololoniaina, Liliane, Grosjean, Stéphane, Raminosoa, Noromalala Rasoamampionona, Glaw, Frank, Vences, Miguel (2006): Molecular identification, description, and phylogenetic implications of the tadpoles of 11 species of Malagasy treefrogs, genus Boophis. Journal of Natural History 40 (23 - 24): 1449-1480, DOI: 10.1080/00222930600902399, URL: http://dx.doi.org/10.1080/00222930600902399
19120E4BB64C0A12458F4CE3FB74C3A4.taxon	description	(Figure 11) The following description refers to one voucher specimen in developmental stage 36 (ZSM 574 / 2004, field number LR 222 d, TL 30.5 mm, BL 11.5 mm), from locality 3. In dorsal view (Figure 11 a), body ovoid, snout rounded. In lateral view (Figure 11 b), body depressed, BW 118 % of BH, snout rounded. Eyes of moderate size, ED 10.0 % of BL, slightly bulging, not visible in ventral view, positioned almost dorsally but directed anterolaterally and dorsolaterally, situated at about the anterior quarter of the body. Nares oval, of moderate size, rimmed, positioned dorsolaterally, directed anterolaterally and dorsolaterally, nearer to anterior edge of eyes than to tip of snout, RN 133 % of NP; NN 49 % of PP. Spiracle sinistral, small, conical, inner wall free and formed such that aperture opens laterally instead of posteriorly; closer to end of body than to snout, SS 67 % of BL, situated a little below the longitudinal axis of tail musculature. Vent tube short, dextral, opening dextral, its right wall displaced anterodorsally, directed posterodorsally, linked to ventral tail fin except its tip. Tail relatively long. Caudal musculature moderately developed, TMH 50 % of BH and 67 % of MTH, TMW 37 % of BW, at mid-length of tail its height about half of the total tail height, reaching almost to tail tip. Caudal fins moderately weakly developed, MTH 75 % of BH; dorsal fin originating at the dorsal tail – body junction and slightly taller than ventral fin at mid-length of tail, ventral fin originating immediately behind body; tail tip pointed. Oral disc (Figure 11 c) moderately small, ODW 19 % of BL and 31 % of BW, positioned and directed anteroventrally, emarginated. Oral disc bordered by a single row of about 50 marginal papillae interrupted by a large median gap on the upper labium (DG 57 % of ODW), and by a small medial gap on the lower labium. A few submarginal papillae positioned in the lateral parts of the anterior and posterior labia. Papillae moderately large, stout, conical with a rounded tip. No denticulate papillae. Keratodont row formula 1: 2 + 2 / 1 + 1: 2. P 1 interrupted by less than 0.1 mm; P 2 the longest keratodont row of the oral disc. About 79 keratodonts on A 1 (ca 49 per mm); P 1 and P 2 of about similar length, P 3 about two-thirds of P 2. Upper jaw finely serrated, black with a larger medial serration surrounded by smaller serrations on each side forming a slight convexity; lower jaw black, moderately serrated, V-shaped. Coloration in preservative. Dorsally: whole dorsum body and caudal musculature uniformly mottled; eyes partially sunk into orbital sockets; nares mixed with the markings and difficult to see. Laterally: intestinal coils well visible; musculature junctions slightly distinct in the anterior part of tail musculature; dorsal fin moderately mottled; ventral fin almost clear. Ventrally: branchial and cardial region slightly visible through ventral body wall; intestinal coils dextral, visible ventrally with regular spiral shape. Variation. TL and BL of 26 tadpoles at stages 25 – 39 (ZSM 561 / 2004 – 574 / 2004, LR 221 a, LR 221 b, LR 221 b 2, LR 221 b 3, LR 222 c 1, LR 222 c 2, LR 222 c 3, LR 222 c 5, LR 222 c 7, LR 222 c 9, LR 222 c 11, LR 222 c 12), all from locality 3, are 18.3 – 33.5 and 7.3 – 12.5 mm, respectively. The ratios vary in the following proportions: BW 113 – 144 % of BH; ED 10.0 – 13.2 % of BL; RN 80 – 133 % of NP; NN 41 – 54 % of PP; SS 60 – 75 % of BL; TMH 40 – 56 % of BH; TMH 40 – 58 % of MTH; TMW 30 – 41 % of BW; MTH 75 – 118 % of BH; ODW 17 – 25 % of BL; ODW 27 – 42 % of BW. KRF of the 26 tadpoles varies from 1: 2 + 2 / 1 + 1: 2 to 1: 3 + 3 / 1 + 1: 2.	en	Raharivololoniaina, Liliane, Grosjean, Stéphane, Raminosoa, Noromalala Rasoamampionona, Glaw, Frank, Vences, Miguel (2006): Molecular identification, description, and phylogenetic implications of the tadpoles of 11 species of Malagasy treefrogs, genus Boophis. Journal of Natural History 40 (23 - 24): 1449-1480, DOI: 10.1080/00222930600902399, URL: http://dx.doi.org/10.1080/00222930600902399
