identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4FE5144D5B035D02D8497C058D42A752.text	4FE5144D5B035D02D8497C058D42A752.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmeticus Menge 1868	<div><p>Tmeticus Menge, 1868</p><p>Type species.</p><p>Tmeticus leptocaulis Menge, 1868 (= Tmeticus affinis (Blackwall, 1855)).</p><p>Diagnosis.</p><p>Males of this genus are easily recognized by possessing a mastidion (large tooth on frontal part of chelicera) and by their elongate palp with patella longer than cymbium, ventral terminal tooth on patella, and thin bulbus (as wide as terminal part of tibia). Females are recognized by their flat epigyne without a cavity. Males may be confused only with the trans-Palaearctic Hylyphantes graminicola (Sundevall, 1830) because it also has a mastidion and a patellar tooth. However, the males of Hylyphantes have shorter palp, undivided embolic division and screw-like embolus.</p><p>The females of Tmeticus may be confused with those of several genera, such as Oedothorax Bertkau, 1883 or with Donacochara speciosa (Thorell, 1875). However, Oedothorax females have a different colour pattern, and Donacochara speciosa is notably larger.</p><p>Description.</p><p>Small to medium-sized (2.5-4.1), light to dark-coloured erigonines. Male carapace unmodified and without sulci, it may be uniformly coloured or with a darker cephalic region. Abdomen unmodified, dark, of uniform colour. Male chelicera modified by possessing a mastidion (Ma, promarginal tooth). Maxilla with apical-retrolateral spine. Tibial spines 2-2-1-1. TmIV present. TmI 0.65-0.8. Male palp elongate. Femur, patella and tibia longer than wide. Patella with conical, ventral terminal tooth (Tt). Tibia with two apophyses (Ta). Paracymbium large, with or without ( Tmeticus affinis) distinct apical pocket. Tegulum with distinct sac (Ts) and large ( Tmeticus tolli) or small protegulum (Pt). Radix with straight apical process (Ap), tailpiece (Tp) without extension, embolus (Em) short and straight, or long and forming a semicircle; embolic membrane (Me) large. Epigyne without distinct fovea or openings. Median (=dorsal, sensu Hormiga 2002) plate plain or with ridges.</p><p>Composition.</p><p>According to Platnick’s (2010) catalogue eight species are listed in this genus: Tmeticus affinis (Blackwall, 1855) (Holarctic), Tmeticus bipunctis ( Bösenberg &amp; Strand, 1906) (Far East Asia), Tmeticus neserigonoides Saito &amp; Ono, 2001 (Japan), Tmeticus nigerrimus Saito &amp; Ono, 2001 (Japan), Tmeticus nigriceps ( Kulczyński, 1916) (Northern Siberia), Tmeticus ornatus (Emerton, 1914) (USA &amp; Canada), Tmeticus tolli Kulczyński, 1908 (Siberia) and Tmeticus vulcanicus Saito &amp; Ono, 2001 (Japan). In fact, there are three more species names within this genus: Tmeticus difficilis Kulczyński, 1926, Tmeticus dubius Kulczyński, 1926 and Gongylidium vile Kulczyński, 1885. Of these, the first two are listed under Centromerus, and the last one under Oedothorax . These three names were included in Tmeticus by Holm (1973) and Eskov (1994) but were considered as synonyms of Tmeticus tolli .</p><p>On the basis of the present study, we conclude that Tmeticus encompasses four species: Tmeticus affinis, Tmeticus nigriceps, Tmeticus ornatus and Tmeticus tolli . A new genus has been erected for Tmeticus bipunctis . Tmeticus neserigonoides might be correctly placed in this genus, but as we failed to re-examine its specimens, we treat it as incertae sedis (see below). Two other Japanese species belong elsewhere, but their correct assignments require further study.</p><p>Comments.</p><p>Tmeticus is unusual in the Erigoninae because all its species can be recognized by their carapace colour pattern. Three sibling species: Tmeticus nigriceps, Tmeticus ornatus and Tmeticus tolli cannot be recognized by their embolic division, but the females of these species have distinctly different epigynes.</p><p>Interrelationships.</p><p>Tmeticus affinis differs from the three other species by the shape of the paracymbium, the straight embolus and the high protegulum with papillae. It also possesses a different type of the tibial apophysis, not originating at the terminal edge of the tibia as in other Erigoninae and other Tmeticus, but slightly aside of the edge.</p><p>Relationships.</p><p>In general appearance, male palp structure and cheliceral dentition, the members of this genus are similar to Hylyphantes graminicola, but the latter has a different type of embolic division and epigyne. When Wiehle (1956) described Donacochara speciosa (Thorell, 1875) he compared it with Tmeticus affinis . Both species have a long palp, small bulbus, and the chelicera of the male has a mastidion. The embolic division in both species is rather similar, but the radical process and the embolus proper occupy different positions.</p><p>The embolic division of Tmeticus is similar to those in Phaulothrix hardyi (Blackwall, 1850) (cf. Millidge 1977: f. 18), Lophomma punctatum (Blackwall, 1841)(Fig. 58) or members of Oreoneta Chyzer &amp; Kulczyński, 1894 (Fig. 57). All these genera have a more or less straight, two-armed embolic division (embolus proper + anterior radical process), with the embolus proper situated more dorsally than the process. All three genera have a wide embolic membrane.</p><p>Millidge (1977) placed Tmeticus in a separate nominative group with Ostearius Hull, 1911, Donacochara Simon, 1884, Eboria Falconer, 1910 and Sciastes Bishop &amp; Crosby, 1938. Hormiga (2002) placed Tmeticus close to a very heterogeneous group of the higher Erigoninae that includes such unrelated genera (in terms of the structure of the embolic division) as Walckenaeria Blackwall, 1833 (twisted radix), Oedothorax, Entelecara Simon, 1884, Gonatium Menge, 1968 and others.</p><p>Judging from the drawings (Figs 35.110, 35.111 in Draney and Buckle 2005), Tmeticus can be related to the Nearctic Nanavia monticola Chamberlin &amp; Ivie, 1933. The latter species seems to have been mistakenly considered a synonym of Leptorhoptrum robustum (Westring, 1851) (see Platnick 2010). Both genera and species were synonymized by Eskov and Marusik (1994) on the basis of a comparison of Leptorhoptrum robustum and the poor figures of Nanavia monticola . Nanavia monticola has a very long palpal tibia, and the paracymbium and embolic division are very similar to those of Tmeticus affinis . The relationships of the two genera and taxonomic status of Nanavia Chamberlin &amp; Ivie, 1933 are outside the scope of this study and will be considered elsewhere.</p><p>Key to Tmeticus species</p><p>The males of Tmeticus nigriceps and Tmeticus ornatus cannot be distinguished.</p></div>	https://treatment.plazi.org/id/4FE5144D5B035D02D8497C058D42A752	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Marusik, Yuri M.;Koponen, Seppo	Marusik, Yuri M., Koponen, Seppo (2010): A review of the Holarctic genus Tmeticus Menge, 1868 (Araneae, Linyphiidae), with a description of a new genus. ZooKeys 59: 15-37, DOI: http://dx.doi.org/10.3897/zookeys.59.508, URL: http://dx.doi.org/10.3897/zookeys.59.508
D12CE4F829E320A4DE833741FB966C53.text	D12CE4F829E320A4DE833741FB966C53.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmeticus affinis (Blackwall 1855) Blackwall 1855	<div><p>Tmeticus affinis (Blackwall, 1855) Figs 111194961</p><p>Neriene affinis Blackwall 1855: 121 (D♂).</p><p>Tmeticus affinis: Wiehle 1960: 411, f. 751-756 (♂♀).</p><p>Tmeticus affinis: Merrett 1963: 410, f. 83A-C (♂).</p><p>Tmeticus affinis: Millidge 1977: 37, f. 150 (♂).</p><p>Tmeticus affinis: Roberts 1987: 44, f. 12e (♂♀).</p><p>Tmeticus affinis: Millidge 1993: 147, f. 32 (♀). For a complete set of references see Platnick (2010).</p><p>Material examined.</p><p>FINLAND: 2♀ (ZMUT), Turku Ruissalo, 14.11.1966 (M. Saaristo); 2♀ (ZMUT), Turku Ruissalo, sea shore litter, 27.10.1966 (M. Saaristo); 1♂ (ZMUT) Turku Hirvensalo Illoinen, 30.5.1966 (P.T. Lehtinen); 1♀ (ZMUT), Turku Kärsämöki Pomponrahka, 30.05.1967 (M. Saaristo); 1♀ (ZMUT) Pori Yyteri, 16.10.1966 (M. Saaristo); 1♀ (ZMUT), Pudasjärvi Hirvaskoski, 12.08.1959 (P.T. Lehtinen); 1♂ (ZMUT), Kuusamo Torankijärvi, 7.7.1966 (M. Saaristo); 1♀ (ZMUT), Kajaani, Koutaniemi, 16.07.1972 (P.T. Lehtinen); 1♂ (ZMUT) Inari Repojoki, 9.7.1961 (O.V. Lindqvist); 1♀ (ZMUT) Utsjoki Kevo, birch forest on lake shore, 20.06.-20.07.1970 (E.T. Linnaluoto). RUSSIA: Krasnoyarsk Province, 1♂ 2♀ (ZMMU), Mirnoye, Yenisei River left bank, 23.06.1978 (K.Yu.Eskov); 2♀ (ZMMU), Mirnoye, Yenisei River left bank, 27.07.1979 (K.Yu. Eskov). Yakutia, 2♂ 2♀ (ZMUT), El’gyay, big ”alas” pond, 24.07.1977 (S. Koponen); 1♂ (ZMMU), western Yakutia, Kempendyay River 80 km up stream from the mouth, riverside meadow, 1-15.08.1988 (K.Yu. Eskov). Kamchatka Peninsula, 1♂ 1♀ (IBPN), Talovskoye Lake, Kuyul River, 16.08.1990 (M.B. Skopets). Chukotka: 1♂ (ZMMU), Markovo, July 1986 (G. Chernova). CANADA, Alberta: 1♂ (only the photo provided by D.J. Buckle has been studied), Caribou Mountain Wildlands, Wentzel Lake, 50°02N; 114°28W, sweeping horsetail meadow, 16.07.2003 (T. Johnson).</p><p>Diagnosis.</p><p>This species is easily recognized by its brownish carapace with a darker cephalic region. Males are easily recognized by their palp, which has a characteristic tibial apophysis and embolic division with the anterior radical process equal in length to the embolus proper (embolus longer than anterior radial process in other species). Females are easily recognized by the shape of the epigyne.</p><p>Description.</p><p>For detailed description see Wiehle (1960). ♀ 2.5-3.0, ♂ 2.5-2.8. TmI 0.65-0.75. Carapace reddish brown, rather dark in males. Cephalic region slightly darker than thoracic, but there is no clear demarcation between the two. Abdomen black. Legs orange-brown. Palp as in Figs 1-3, 5-7, 49, 61; epigyne as in Figs 4, 8-9, 19.</p><p>Distribution.</p><p>This species is known all over Eurasia, from western Europe to Kamchatka. In the Nearctic Region, it has been reported from Alberta (Nordstrom and Buckle 2006).</p></div>	https://treatment.plazi.org/id/D12CE4F829E320A4DE833741FB966C53	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Marusik, Yuri M.;Koponen, Seppo	Marusik, Yuri M., Koponen, Seppo (2010): A review of the Holarctic genus Tmeticus Menge, 1868 (Araneae, Linyphiidae), with a description of a new genus. ZooKeys 59: 15-37, DOI: http://dx.doi.org/10.3897/zookeys.59.508, URL: http://dx.doi.org/10.3897/zookeys.59.508
615EB239A58CECDB74998403E776E9AE.text	615EB239A58CECDB74998403E776E9AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmeticus tolli Kulczynski 1908	<div><p>Tmeticus tolli Kulczynski, 1908 Figs 15-16, 2025345056</p><p>Gongylidium vile Kulczyński 1885: 37, pl. 10, f. 16 (♀), syn. n.</p><p>Tmeticus tolli Kulczyński 1908: 15, pl. 1, f. 3, 7-8, 22-23 (♂♀).</p><p>Tmeticus difficilis Kulczyński 1926: 50 (♀), syn. n.</p><p>Tmeticus dubius Kulczyński 1926: 49 (♀), syn. n.</p><p>Centromerus tolli: Sytshevskaja 1935: 90.</p><p>Tmeticus tolli: Holm 1973: 89.</p><p>Tmeticus tolli: Eskov 1994: 107.</p><p>Tmeticus tolli: Hormiga 2000: 56, f. 29A-I, pl. 67A-F, 68A-F (♂♀).</p><p>Tmeticus affinis: Marusik and Logunov 1999: 245 (misidentification).</p><p>Material examined.</p><p>RUSSIA: Krasnoyarsk Province: 1♂ 2♀ (ZMMU), Mirnoye, Yenisei River left bank, 23.06.1978 (K.Yu. Eskov). Evenkiya: 40♂♀ (ZMMU), Taimura River, Neptene River mouth, riparian spruce forest with alder, Summer 1982 (K.Yu. Eskov); 2♀ (ZMMU), Chambe River mouth, meteorological station “Kerbo”, floodplain willow stand, litter, 21.08.1982 (K.Yu. Eskov). Khabarovsk Province: 2♀ (IBPN), Okhotski Dist., Gyrbykan R. ( Ul’ya River basin), 20.08-15.09.1986 (I.D. Sukatcheva); 1♂ 3♀ (IBPN), Khetana River (tributary of Amka River, Ulya River basin), Agust 1985 (V.V. Zherikhin). Maritime Prov.: 3♂ 2♀ (IBPN), [05], Khanka Lake CW shore, Sosnovy Isl &amp; peninsula nearby, 44°52N; 132°07E, 17.07.1998 (Yu.M. Marusik). 1♀ (IBPN), [03], Khanka Lake, CE shore, 44°39N; 132°34E, 15-16.07.1998 (Yu.M. Marusik). Magadan Area: 3♂ 2♀ (IBPN), Motykley Bay, 59°30N; 148°50E, Summer 1994 (E. Izergina); 1♂ 1♀ (IBPN), 137th km of Kolyma Hwy, 60°25N; 151°30E, Ola River, valley forest, 28.09.1994 (Yu.M. Marusik); 1♂ 2♀ (IBPN), ca 50 km N of Magadan, Khasyn River, environs of Splavnaya Vil., 28.05.1988 (Yu.M. Marusik); 25♂♀ (IBPN), 30km N of Magadan, Snow Valley Vil., Dukcha River valley, 7.10.1984 (Yu.M. Marusik). Sakhalin Island: 1♂ 6♀ (IBPN), Okha Dist., Ten’ga River, May 1987 (A.M. Basarukin); 4♀ (IBPN), Tomari Dist., Ainskoye Lake, Ptichya river, 24.05.-10.06.1984 (A.M. Basarukin); 1♀ CE part, Leonidovka River, 8 km SE of Leonidovo Vil., 49°16.506N; 142°58.390E, 9.08.2001 (Yu.M. Marusik). Kamchatka Peninsula: 2♀ (ZMMU), 40 km from Ust’ -Kamchatsk, 09.1973 (A.S. Glikman); 1♂ 4♀ (IBPN), 10-12 km N of Paratunka Vil., Yelizovo Forestry, 53.050°N; 158.225°E, 15-28.07.2004 (A.S. Ryabukhin). MONGOLIA: Arkhangai Aimak: 2♂ 2♀ (IBPN) [12], Ondrer-Ulaan, Tsakhir, Chulut gorge 48°07N; 100°22E, 2100 m, 10-13.06.1997 (Yu.M. Marusik). Central (=Tov) Aimak: 1♀ (IBPN), Terelzh Mt., south exposed slope (about 80 km NE of Ulan-Bator, 1988 (S. Heimer).</p><p>Diagnosis.</p><p>Tmeticus tolli is easily distinguished from the similar Tmeticus ornatus and Tmeticus nigriceps by having a uniformly coloured carapace in both sexes.</p><p>Description.</p><p>Both sexes were described in detail by Hormiga (2000). ♀ 2.8-4.1, ♂ 2.5-3.1. TmI 0.68-0.8. Carapace from orange to pale yellow with black median stripe. Abdomen from light brown to blackish. Legs coloured as carapace. Palp as in Figs 27-30, 32, 50, 56, epigyne as in Figs 20, 31, 33-34.</p><p>Comments.</p><p>Tmeticus difficilis Kulczyński, 1926 was described on the basis of the female holotype from Lake “Kurarotschnoje” (=Kurazhechnoye, ca. 56°10N; 161°45E, collected 9.06.1909) and Tmeticus dubius Kulczyński, 1926 was described on the basis of two females from Lake “Klutschevskoje” (= maybe Klyuchi Vil., c. 60 km from Kurarochnoye Lake). In his descriptions Kulczyński (1926) compared both species with Tmeticus tolli . Both species were transferred to Centromerus (a member of the Micronetinae, a different subfamily) by Reimoser (1919) and this transfer was not contested by Charitonov (1932). Sytschevskaya (1935), who collected in the same places, suggested that both species, in addition to Gongylidium vile Kulczyński, 1885 (from Petropavlovsk-Kamchatski) maybe conspecific with Tmeticus tolli . Holm (1973) agreed with Sytschevskaya (1935). Eskov (1994) and Mikhailov (1997) listed the threespecies as synonyms of Tmeticus tolli, but formal synonymies were not proposed. In addition, Gongylidium vile (listed as Oedothorax vilus in Platnick’s (2010) catalogue has date priority over Tmeticus tolli .</p><p>Although Kulczyński (1885) compared Gongylidium vile with European Oedothorax (the epigyne of Tmeticus tolli is very similar to those in Oedothorax), he mentioned the colour of the carapace “flavido-rufo” (yellow-red = orange), which is typical for Tmeticus tolli and such coloration is absent from Oedothorax species. Furthermore, no Oedothorax species has been recorded from Kamchatka. In order to retain stability we suggest suppression of the name Gongylidium vile Kulczyński, 1885, because it does not appear in the literature (except for catalogues and nomenclatorial notes), whereas Tmeticus tolli has been used in more than 25 publications by more than 10 different authors during the past 50 years (Holm 1973; Eskov 1988, 1994; Koponen and Marusik 1992; Marusik et al. 1992, 1993, 2002; Marusik 1993, 2005a - b; Rybalov et al. 2002; Hormiga 2000; Tanasevitch 2006; Trilikauskas and Tanasevitch 2006; etc.). We agree with Eskov about the status of these species and here formally propose three new synonymies: Gongylidium vile Kulczyński, 1885, syn. n. = Tmeticus difficilis Kulczyński, 1926, syn. n. and Tmeticus dubius Kulczyński, 1926, syn. n. = Tmeticus tolli Kulczyński, 1908.</p><p>Distribution .</p><p>This species is distributed east of Yenisei (Eskov 1994) to Chukotka and southward to central Mongolia (Marusik and Logunov 1999: sub. Tmeticus affinis), Maritime Province of Russia (present data) and northern Sakhalin (Eskov 1994). Tmeticus tolli also occurs in northeastern China. YM saw one female specimen of this species in the collection of Baoding University (China).</p></div>	https://treatment.plazi.org/id/615EB239A58CECDB74998403E776E9AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Marusik, Yuri M.;Koponen, Seppo	Marusik, Yuri M., Koponen, Seppo (2010): A review of the Holarctic genus Tmeticus Menge, 1868 (Araneae, Linyphiidae), with a description of a new genus. ZooKeys 59: 15-37, DOI: http://dx.doi.org/10.3897/zookeys.59.508, URL: http://dx.doi.org/10.3897/zookeys.59.508
599334D130FEAC104975ADBF519C3F56.text	599334D130FEAC104975ADBF519C3F56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmeticus nigriceps (Kulczynski 1916) Kulczynski 1916	<div><p>Tmeticus nigriceps (Kulczynski, 1916) Figs 1823 –2435– 445255</p><p>Gongylidium nigriceps Kulczyński 1916: 8, pl. 1, f. 10 (♀).</p><p>Tmeticus nigriceps: Holm 1973: 89.</p><p>Tmeticus nigriceps: Eskov 1994: 107.</p><p>Misidentifications.</p><p>(all refer to Zornella cultrigera (L. Koch, 1879) see Holm (1973):</p><p>Gongylidium nigriceps: Tullgren 1955: 355, f. 56 (♀).</p><p>Gongylidium nigriceps: Hauge 1969: 6, f. 12 (♀).</p><p>Material examined.</p><p>RUSSIA: Arkhangel’sk Area: 1♂ (IBPN), Barents Sea, Dolgiy Ilsand, 69°12'N, Summer 2004 (O.L. Makarova). Polar Ural: 1♂ (ZMUT), Oktyabrskij, Ob River shore, Salix viminalis vegetation, 12.-13.7.1994 (S. Koponen); 1♀ (PSU-95), North Ural expedition by Fridolin, sample 36, Sob’ River right bank, 4.07.1925 (V. Fridolin). Yamal Peninsula: 1♂ 5♀ (ZMMU), Yorkugayakha River, environs of “Canary” trading station, riparian willow stand, 08.07.2002 (D. Osipov); 4♂ 1♀ (ZMMU), south Yamal, Shchuchye Vill, Shchuchya River (A.L. Tikhomirova); 1♂ (PSU-96), South Yamal, Khadyta-Yakha River, meadow valley, pitfall traps, 8.08.1982 (S.L. Esyunin); 1♂ 1♀ (PSU-97), same locality, river bank, drift, 26.07.1981 (S.L. Esyunin). Taimyr Peninsula: 1♂ 1♀ (ZMMU), Taimyr Reserve, Novaya River, Ary-Mas Site, 25.07.1992 (A.B. Ryvkin); 10♂ 1♀ (ZMMU), SW Taimyr, Nyapan’ Ridge, 70°09N; 87°47E, Carex-moss bog, pitfall trapping, 1-10.08.1999 (D. Osipov); 3♂ (ZMMU), NW shore of Pyasino Lake, 70°04'54N; 87°32'12E, Carex bog with sphagnum tussocks, 10-20.07.1999 (D. Osipov); 3♂ 2♀ NW shore of Pyasino Lake, 70°04'54N; 87°32'17E, Carex bog with sphagnum tussocks, Summer 1999 (D. Osipov); 2♀ (ZMMU), NW shore of Pyasino Lake, Lazannakh Lake, sandy beach, 70°05'47N; 87°26'28E, 1-10.07.1999 (D. Osipov). Yakutia: 2♀ (IBPN), Yana River down flow, environs of Kular Village, 70°35N; 134°34E, grass-herb- Arctagrostis meadow on the former open mine, Summer, 2000 (N.K.Potapova). Chukotka: 2♂ (IBPN), western Chukotka, Chaun River mouth part, 68.810°N; 170.432°E, Summer, 1986 (A.S. Ryabukhin); 1♂ (IBPN), western Chukotka, Chaun Bay, Pucheveyem River mouth, 25.07.1985 (A.S. Ryabukhin); 1♂ (IBPN) Lamutskoye Vil., 65°32'39N; 168°51'08E, along creek, 17.08.1968 (Novikova); 1♂ (IBPN), western Chukotka, Markovo Town, July 1986 (G. Chernova); 1♂ (IBPN), Vulvyveyem River upper flow, Gytlenumkuum Stand, 67°10N; 178°E, 8.06.1988 (Yu.M. Marusik).</p><p>Diagnosis.</p><p>Tmeticus nigriceps is easily distinguished from the other Palaearctic species by the dark cephalic region contrasting with the reddish thoracic area. Only the Nearctic Tmeticus ornatus has a similar colour pattern. The male palp is almost undistinguishable from the Siberian Tmeticus tolli and the Nearctic Tmeticus ornatus . In addition to the carapace pattern, females can be distinguished by the shape of their epigyne.</p><p>Description.</p><p>♀ 2.9-3.3, ♂ 2.3-2.7. TmI 0.69-0.72. Carapace orange with dark, blackish cephalic region (Figs 23-24, 38) and chelicera. Legs orange. Abdomen black. Palp as in Figs 35-37, 40-42, 52, epigyne as in Figs 18, 39, 43-44.</p><p>Comments.</p><p>Holm (1973) re-examined the specimens from Sweden identified and recorded as Gongylidiun nigriceps by Tullgren (1955) and concluded that they were misidentifications of Zornella cultrigera (L. Koch, 1879). The figure of the female epigyne from Norway in Hauge (1969) identified as Gongylidium nigriceps also refers to Zornella cultrigera .</p><p>Distribution .</p><p>This species is known from Dolgiy Island and the Polar Urals to Chukotka (Eskov 1994). It is restricted to the tundra zone.</p></div>	https://treatment.plazi.org/id/599334D130FEAC104975ADBF519C3F56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Marusik, Yuri M.;Koponen, Seppo	Marusik, Yuri M., Koponen, Seppo (2010): A review of the Holarctic genus Tmeticus Menge, 1868 (Araneae, Linyphiidae), with a description of a new genus. ZooKeys 59: 15-37, DOI: http://dx.doi.org/10.3897/zookeys.59.508, URL: http://dx.doi.org/10.3897/zookeys.59.508
DEBB7ED68B15BF02CB95BE7855538D0A.text	DEBB7ED68B15BF02CB95BE7855538D0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmeticus ornatus (Emerton 1914) Emerton 1914	<div><p>Tmeticus ornatus (Emerton, 1914) Figs 1721 –2245– 485154</p><p>Gongylidium ornatus Emerton 1914: 263, pl. 8, f. 3 (♂).</p><p>Tmeticus ornatus: Bishop and Crosby 1935: 227, pl. 18, f. 22-26 (♂♀).</p><p>Tmeticus ornatus: Paquin and Dupérré 2003: 122, f. 1284-1286 (♂♀).</p><p>Material examined.</p><p>CANADA: 4♂ 4♀ (ZMMU), Saskatchewan, Lady Lake, sedge tops - flooded marsh, 13-15.04.1971 (D.J. Buckle); 3♂ 3♀ (ZMUT), same locality, marsh, late April, 1978 (J.V. Buckle).</p><p>Diagnosis.</p><p>Differs from Tmeticus affinis, which also occurs in the Nearctic Region, by the carapace colour (black cephalic region and red-orange thoracic area in Tmeticus ornatus, carapace uniformly brown in Tmeticus affinis). The males are easily separated by their tibial apophyses (one apophysis with a claw-like processes in Tmeticus affinis and two separate apophyses in Tmeticus ornatus); the females have distinctly different epigynes.</p><p>Description .</p><p>♂ 2.5-3.3, ♀ 2.8-35. TmI 0.73-0.78. Carapace orange with darker cephalic region. Abdomen dark. Palp as in Figs 45-46, 51, 54. Epigyne as in Figs 17, 47-48.</p><p>Distribution.</p><p>This species has a trans-Nearctic distribution, recorded from British Columbia to Quebec and south to New York (Buckle et al. 2001). It does not occur north of 55°N and has a more southern distribution in comparison to the Palaearctic Tmeticus affinis, Tmeticus tolli and Tmeticus nigriceps .</p></div>	https://treatment.plazi.org/id/DEBB7ED68B15BF02CB95BE7855538D0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Marusik, Yuri M.;Koponen, Seppo	Marusik, Yuri M., Koponen, Seppo (2010): A review of the Holarctic genus Tmeticus Menge, 1868 (Araneae, Linyphiidae), with a description of a new genus. ZooKeys 59: 15-37, DOI: http://dx.doi.org/10.3897/zookeys.59.508, URL: http://dx.doi.org/10.3897/zookeys.59.508
1861121A7C9304DD73E201A3A7D6DEDA.text	1861121A7C9304DD73E201A3A7D6DEDA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmeticus neserigonoides Saito & Ono 2001	<div><p>Tmeticus neserigonoides Saito &amp; Ono, 2001</p><p>Tmeticus neserigonoides Saito and Ono 2001: 9, f. 15-20 (♂♀).</p><p>Tmeticus neserigonoides: Ono et al. 2009: 304, f. 638-642 (♂♀).</p><p>Comments.</p><p>Judging from the available figures, this species might belong in Tmeticus, The male has a long palp with a patellar tooth. However, the chelicera appears to lack a mastidion and the tibial apophyses are absent. Figures of the male palp are unclear, TmI index (0.59) is lower than in Tmeticus species (&gt;0.63).</p></div>	https://treatment.plazi.org/id/1861121A7C9304DD73E201A3A7D6DEDA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Marusik, Yuri M.;Koponen, Seppo	Marusik, Yuri M., Koponen, Seppo (2010): A review of the Holarctic genus Tmeticus Menge, 1868 (Araneae, Linyphiidae), with a description of a new genus. ZooKeys 59: 15-37, DOI: http://dx.doi.org/10.3897/zookeys.59.508, URL: http://dx.doi.org/10.3897/zookeys.59.508
64F280595756BA8206DC377E51480DC8.text	64F280595756BA8206DC377E51480DC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmeticus nigerrimus Saito & Ono 2001	<div><p>Tmeticus nigerrimus Saito &amp; Ono, 2001</p><p>Tmeticus nigerrimus Saito and Ono 2001: 13, f. 26-31 (♂♀).</p><p>Tmeticus nigerrimus: Ono et al. 2009: 304, f. 643-647 (♂♀).</p><p>Comments.</p><p>This species is clearly not related to Tmeticus affinis or other members of the genus due to the short palpal patella lacking a tooth in the male, embolic division of a different shape, the relatively long tibial apophysis, lack of a mastidion, epigyne with a septum and some other additional characters. The correct generic placement remains unclear.</p></div>	https://treatment.plazi.org/id/64F280595756BA8206DC377E51480DC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Marusik, Yuri M.;Koponen, Seppo	Marusik, Yuri M., Koponen, Seppo (2010): A review of the Holarctic genus Tmeticus Menge, 1868 (Araneae, Linyphiidae), with a description of a new genus. ZooKeys 59: 15-37, DOI: http://dx.doi.org/10.3897/zookeys.59.508, URL: http://dx.doi.org/10.3897/zookeys.59.508
17779557245D389688A66930B6FB26FE.text	17779557245D389688A66930B6FB26FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tmeticus vulcanicus Saito & Ono 2001	<div><p>Tmeticus vulcanicus Saito &amp; Ono, 2001</p><p>Tmeticus vulcanicus Saito and Ono 2001: 11, f. 21-25 (D♂♀).</p><p>Tmeticus vulcanicus: Ono et al. 2009: 304, f. 648-652 (♂♀).</p><p>Comments.</p><p>This species is clearly not related to Tmeticus affinis or other members of the genus due to the short palpal patella lacking a tooth in the male, embolic division of a different shape (anterior radical process absent), and some other characters. The correct generic placement remains unclear.</p></div>	https://treatment.plazi.org/id/17779557245D389688A66930B6FB26FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Marusik, Yuri M.;Koponen, Seppo	Marusik, Yuri M., Koponen, Seppo (2010): A review of the Holarctic genus Tmeticus Menge, 1868 (Araneae, Linyphiidae), with a description of a new genus. ZooKeys 59: 15-37, DOI: http://dx.doi.org/10.3897/zookeys.59.508, URL: http://dx.doi.org/10.3897/zookeys.59.508
256166292A651BB1DE91175C77057A1F.text	256166292A651BB1DE91175C77057A1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paratmeticus	<div><p>Paratmeticus gen. n.</p><p>Type species.</p><p>Oedothorax bipunctis Bösenberg and Strand, 1906.</p><p>Etymology.</p><p>Prefix “Para” - indicates the resemblance of this genus to Tmeticus The gender is masculine.</p><p>Diagnosis.</p><p>The new genus is easily distinguished from the similar Tmeticus by lacking distinct tibial apophyses, and in having the papillate tegular sac larger than the protegulum, a slightly twisted embolic division, a sharply pointed embolic membrane and a large distal suprategular apophysis, longer than the embolic division. In contrast to Tmeticus, themedian plate of the epigyne in the new genus is widest in the anterior region, rather than in the posterior region.</p><p>Description.</p><p>Medium-sized erigonine spiders. Uniformly coloured, male carapace without modifications, male chelicera with mastidion, inner row with 4 inner teeth and 5 outer teeth (all smaller than inner teeth). TmI 0.63-0.65. Male palp elongate, with patella as long as tibia, tibia lacks apophyses, distal suprategular apophysis longer than embolic division; embolic division slightly twisted with two arms: anterior radical process and embolus proper; embolus parallel to process with lamellate basal process; epigyne without cavity, median plate widest anteriorly.</p><p>Composition.</p><p>The type species only.</p></div>	https://treatment.plazi.org/id/256166292A651BB1DE91175C77057A1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Marusik, Yuri M.;Koponen, Seppo	Marusik, Yuri M., Koponen, Seppo (2010): A review of the Holarctic genus Tmeticus Menge, 1868 (Araneae, Linyphiidae), with a description of a new genus. ZooKeys 59: 15-37, DOI: http://dx.doi.org/10.3897/zookeys.59.508, URL: http://dx.doi.org/10.3897/zookeys.59.508
BE51D7C1EEA1D163F658FD47915756CA.text	BE51D7C1EEA1D163F658FD47915756CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paratmeticus bipunctis (Boesenberg & Strand 1906) Boesenberg & Strand 1906	<div><p>Paratmeticus bipunctis (Boesenberg &amp; Strand, 1906) comb. n. Figs 12 –145359– 606273</p><p>Oedothorax bipunctis Bösenberg and Strand 1906: 162, pl. 12, f. 258 (♀).</p><p>Tmeticus japonicus Oi 1960: 152, f. 50-51 (♂).</p><p>Tmeticus japonicus: Chikuni 1989: 58, f. 56 (♂♀).</p><p>Tmeticus japonicus: Eskov 1994: 107.</p><p>Tmeticus bipunctis: Saito and Ono 2001: 7, f. 10-14 (S, ♀).</p><p>Tmeticus bipunctis: Ono et al. 2009: 304, f. 634-637 (♂♀).</p><p>Material examined.</p><p>RUSSIA: Sakhalin Island: 1♂ 4♀ (IBPN), Okha Dist., 5-7 km N of Kolendo Vil., 22-23.08.1991 (A.M. Basarukin); 1♂ 2♀ (IBPN), Pil’tun Bay, 06.-0.7.1991 (A.M.Basarukin); 1♂ 1♀ (IBPN), Okha Dist., Ten’ga River, May 1987 (A.M. Basarukin); 10♂ 26♀ (IBPN), Korsakov Dist., Tunaiga Lake south shore, 26.09.1991 (A.M.Basarukin). Kuril Isles, 4♂ 3♀ (IBPN), Paramushir Isl. NE shore, environs of Severo-Kuril’sk, 50°40N; 156°06E, 10.08-15.09.1996 (Yu.M. Marusik); 2♂ 1♀ (ZMMU), Iturup Island, Dobroye Nachalo Bay, Lesozavodskoye, mixed forest, 14.08.1994 (K.Yu. Eskov). Kamchatka Peninsula, 6♂ 2♀ (IBPN), 10-12 km N of Paratunka Vil., Yelizovo Forestry, 53.050°N; 158.225°E, 15-28.07.2004 (A.S. Ryabukhin).</p><p>Description.</p><p>Well described by Saito and Ono (2001) and Ono et al. (2009). Total length: ♂ 2.5-3.2, ♀ 2.8-3.5. Carapace 1.29-1.57 long, 1.0-1.24 wide, slightly larger in males. Chelicera in male with mastidion. TmI 0.63-0.65. Carapace dirty brownish, sternum and chelicerae brown. Abdomen dark grey-blackish. Palp as in Figs 53, 59-60, 62-67, 69-71, epigyne as in Figs 14, 65, 72-73.</p><p>Distribution.</p><p>Kamchatka (south part), 8 islands in Kuril Archipelago (Shikotan, Kunashir, Iturup, Urup, Simushir, Ketoi, Shiashkotan, Paramushir, but seems to occur on all large islands); Sakhalin and Japan (Hokkaido, Honshu and Kyushu).</p></div>	https://treatment.plazi.org/id/BE51D7C1EEA1D163F658FD47915756CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Marusik, Yuri M.;Koponen, Seppo	Marusik, Yuri M., Koponen, Seppo (2010): A review of the Holarctic genus Tmeticus Menge, 1868 (Araneae, Linyphiidae), with a description of a new genus. ZooKeys 59: 15-37, DOI: http://dx.doi.org/10.3897/zookeys.59.508, URL: http://dx.doi.org/10.3897/zookeys.59.508
