taxonID	type	description	language	source
E7C605360146577F82EF8C054A7497E5.taxon	description	Fig. 4 A – F	en	Chen, Wei-Cai, Bei, Yong-Jian, Li, Peng (2025): A new species of the genus Leptobrachella (Amphibia, Anura, Megophryidae) from the Darongshan Nature Reserve, Guangxi, China. ZooKeys 1260: 171-194, DOI: 10.3897/zookeys.1260.161514
E7C605360146577F82EF8C054A7497E5.taxon	diagnosis	Diagnosis. Leptobrachella darongshanensis sp. nov. is assigned to the genus Leptobrachella based on molecular phylogenetic results and the following generic morphological characters: small size, presence of an inner metacarpal tubercle, absence of vomerine teeth, distinct tympanum, and the presence of macro-glands (pectoral and femoral glands, often including supra-axillary and ventrolateral glands). Leptobrachella darongshanensis sp. nov. can be distinguished from its congeners by the following combination of characters: (1) medium body size (SVL 24.9 – 27.7 mm in males; 32.0 – 35.4 mm in females); (2) dorsal skin rough with small, raised tubercles and ridges; (3) ventral surface creamy white with minute irregular textures and tiny pale brown spots laterally on the belly; (4) flanks bearing irregular black spots; (5) distinct black supratympanic line extending from posterior corner of eye to supra-axillary gland; (6) rudimentary toe webbing present between toes I – IV, lateral dermal fringes absent on toes; (7) ventrolateral glandular line distinct and continuous; (8) iris bicolored, upper half tangerine, lower half silver with black reticulations; pupil black with tangerine edges; (9) tibiotarsal articulation reaching the posterior margin of the eye when hindlimb is adpressed along body; (10) advertisement call dominant frequency 6.1 – 6.7 kHz at 20.0 ° C.	en	Chen, Wei-Cai, Bei, Yong-Jian, Li, Peng (2025): A new species of the genus Leptobrachella (Amphibia, Anura, Megophryidae) from the Darongshan Nature Reserve, Guangxi, China. ZooKeys 1260: 171-194, DOI: 10.3897/zookeys.1260.161514
E7C605360146577F82EF8C054A7497E5.taxon	description	Description of holotype. Adult male (SVL 27.7 mm). Head width subequal to length (HW / HL = 0.98). Snout projecting, extending beyond lower jaw; nostril oval, closer to snout tip than to eye; canthus rostralis rounded; loreal region concave, sloping towards lips; interorbital region flat; pupil vertical; eye diameter larger than snout length (ED / SNT = 1.16); internarial distance subequal to interorbital distance (IN / IOD = 0.98); tympanum distinct, round, concave, diameter much smaller than eye diameter (TD / ED = 0.44); distinct, raised supratympanic fold extending from posterior corner of eye to supra-axillary gland. Vomerine teeth absent. Tongue pyriform, notched posteriorly. Fingers lacking webbing and lateral fringes; tips slightly swollen; relative finger lengths I <II <IV <III; nuptial pads absent; subarticular tubercles absent; inner palmar tubercle prominent, oval; outer palmar tubercle prominent, rounded; inner and outer palmar tubercles separated. Toes lacking lateral fringes, tips rounded, not swollen; relative toe lengths I <II <V <III <IV; subarticular tubercles absent, replaced by low, indistinct dermal ridges; inner metatarsal tubercle large, elongated; outer metatarsal tubercle absent; webbing rudimentary between toes I – IV. Tibia length ~ 47 % of SVL (TIB / SVL = 0.47); tibiotarsal articulation reaching posterior margin of eye when hindlimb is adpressed anteriad; heels not meeting when thighs are held at right angles to body axis (Fig. 4 A – F). Dorsal skin texture rough with numerous small, raised tubercles and short ridges; ventral skin smooth; pectoral glands oval, creamy white, ~ 1.4 mm in diameter; femoral glands oval, ~ 1.1 mm in diameter, positioned closer to knee than to vent; supra-axillary glands small, round, ~ 1.0 mm in diameter; ventrolateral glandular line distinct and continuous; limbs with sparse tubercles; a pair of small tubercles present above the cloaca (Fig. 4 A – F). Color of holotype in life. Dorsum brown with distinct chocolate markings, including a dark brown triangular marking between eyes and a dark brown ‘ W’ - shaped marking on the scapular region, interspersed with irregular dark spots; tympanum pale brown; distinct black supratympanic line from posterior corner of eye to supra-axillary gland; three dark brown spots on upper lip; flanks with five irregular black spots and numerous small creamy yellow tubercles; dorsal surfaces of hindlimbs with three distinct transverse dark brown bars; dorsal surfaces of forelimbs with three transverse dark brown bars; elbows, upper arms, and tibiotarsal regions pale yellow; ventral surface creamy white with minute irregular textures and tiny pale brown spots laterally on belly; throat and chest pale pinkish; pectoral and femoral glands creamy white; supra-axillary glands pale yellow; iris bicolored, upper half tangerine, lower half silver with black reticulations; pupil black, surrounded by a tangerine ring (Fig. 4 A – F). Color of holotype in preservative. Dorsum dark brown; transverse dark bars on limbs distinct; ventral surfaces creamy white; pectoral, femoral, supra-axillary, and ventrolateral glands creamy white. Variation. Measurements of the type series are presented in Table 3. Coloration and pattern in paratypes are generally consistent with the holotype (Fig. 5). Variation is observed primarily in the intensity and distribution of dark dorsal spots and flank spots. For example, specimen NNU 001394 exhibits more pronounced brown dorsal spotting (Fig. 5 D). Measurements of holotype (in mm): SVL 27.7, HL 9.5, HW 9.3, SNT 3.4, ED 4.0, IOD 2.6, TD 1.8, IN 2.5, TIB 12.9, FLL 13.5, TFL 19.1, ML 7.1, HLL 39.1, FG-knee 4.1.	en	Chen, Wei-Cai, Bei, Yong-Jian, Li, Peng (2025): A new species of the genus Leptobrachella (Amphibia, Anura, Megophryidae) from the Darongshan Nature Reserve, Guangxi, China. ZooKeys 1260: 171-194, DOI: 10.3897/zookeys.1260.161514
E7C605360146577F82EF8C054A7497E5.taxon	etymology	Etymology. The specific epithet darongshanensis is a toponym derived from the type locality, Mt. Darongshan. The suggested English common name is Darongshan Leaf Litter Toad. The suggested Chinese common name is Da Rong Shan Zhang Tu Chan (大容山掌突蟾).	en	Chen, Wei-Cai, Bei, Yong-Jian, Li, Peng (2025): A new species of the genus Leptobrachella (Amphibia, Anura, Megophryidae) from the Darongshan Nature Reserve, Guangxi, China. ZooKeys 1260: 171-194, DOI: 10.3897/zookeys.1260.161514
58F36D1B57E6510DA31150A686F2E9DC.taxon	description	Fig. 6, Suppl. material 1	en	Chen, Wei-Cai, Bei, Yong-Jian, Li, Peng (2025): A new species of the genus Leptobrachella (Amphibia, Anura, Megophryidae) from the Darongshan Nature Reserve, Guangxi, China. ZooKeys 1260: 171-194, DOI: 10.3897/zookeys.1260.161514
58F36D1B57E6510DA31150A686F2E9DC.taxon	description	Description of Guangxi population. The morphological characters of the Darongshan specimens align closely with the original description of L. yunkaiensis by Wang et al. (2018). Measurements are summarized in Table 4. Males SVL 24.6 – 27.2 mm; head length subequal to width (HL 9.3 – 10.5 mm; HW 9.4 – 10.5 mm); snout projecting beyond lower jaw; nostril oval, closer to snout tip than to eye; canthus rostralis rounded; loreal region concave, sloping; interorbital region flat; pupil vertical; ED 3.5 – 4.2 mm; IN 2.5 – 3.1 mm; tympanum distinct, round, concave; distinct, raised supratympanic fold from posterior corner of eye to supra-axillary gland; vomerine teeth absent; tongue pyriform, notched posteriorly. Fingers lacking webbing and fringes; tips slightly swollen; relative finger lengths I <II ≈ IV <III; nuptial pads absent; subarticular tubercles absent; inner and outer palmar tubercles prominent, separated. Toes lacking webbing except rudimentary between II and III; lateral dermal fringes narrow to wide; tips oval, not swollen; relative toe lengths I <II <V <III <IV; subarticular tubercles absent, replaced by indistinct dermal ridges; inner metatarsal tubercle elongated; outer metatarsal tubercle absent; tibiotarsal articulation reaching mid-eye level when leg adpressed; heels meeting when thighs held at right angles to body. Dorsal skin rough with small tubercles; ventral skin smooth; pectoral glands rounded, creamy white, ~ 1.3 mm diameter; femoral glands oval, ~ 1.6 mm diameter, closer to knee than vent; supra-axillary glands small, round, ~ 0.5 mm diameter; ventrolateral glandular line distinct and continuous; limbs with sparse tubercles. Dorsum brown with sparse pale yellow markings; dark brown interorbital triangle; tympanum brown; brown supratympanic line; dark brown spots on upper lip; flanks with irregular black spots and creamy yellow tubercles; dorsal thigh with 3 – 5 distinct transverse dark brown bars; dorsal forearms with three distinct transverse dark brown bars; elbows and upper arms tangerine; ventral surface pink or creamy white with minute irregular creamy textures; ventral limbs with tiny irregular creamy yellow spots; pectoral, femoral, supra-axillary glands creamy yellow; pupil black; iris bicolored, upper half tangerine, lower half silver (Fig. 6, Suppl. material 1).	en	Chen, Wei-Cai, Bei, Yong-Jian, Li, Peng (2025): A new species of the genus Leptobrachella (Amphibia, Anura, Megophryidae) from the Darongshan Nature Reserve, Guangxi, China. ZooKeys 1260: 171-194, DOI: 10.3897/zookeys.1260.161514
58F36D1B57E6510DA31150A686F2E9DC.taxon	distribution	Distribution. The type locality of L. yunkaiensis is Dawuling Forest Station, Maoming City, Guangdong, China (Wang et al. 2018). Previous records include Ehuangzhang Nature Reserve, Guangdong and Lidong, Bobai City, Guangxi, China (Chen et al. 2018; Yu et al. 2024). The Darongshan Nature Reserve, Yulin City, Guangxi, reported herein, represents the fourth known locality for this species (Fig. 1).	en	Chen, Wei-Cai, Bei, Yong-Jian, Li, Peng (2025): A new species of the genus Leptobrachella (Amphibia, Anura, Megophryidae) from the Darongshan Nature Reserve, Guangxi, China. ZooKeys 1260: 171-194, DOI: 10.3897/zookeys.1260.161514
