taxonID	type	description	language	source
183FDF25FFDBA562FF6DF896FB3BFBC5.taxon	materials_examined	Type material examined: Syntype, female. Red ring “ Type ” label; [Ghana] “ Gold Coast. Bibianaha. 700 ft. II. 1912. H. G. F. Spurrell. 1912 - 275. ”; with handwritten: “ Roeselia cretacea type ♀ Hmpsn ”; QR code label with unique number: NHMUK 010598832 (NHMUK). Additional material examined: Liberia. 2 males, 750 m, Nimba County, Nimba Mts., ENNR, Cellcom road, 7 ° 33 ' 3.78 " N, 8 ° 31 ' 46.49 " W, 16 - 28. xii. 2018, Cold Cathode UV Light Trap (8 W), Sáfián, Sz., Simonics, G. leg., ANHRT: 2018.43, slide Nos: LGNA 953, LGNA 954; 1 male, 611 m, Lofa County, Wologizi Mts, base camp forest, 8 ° 07 ' 17 " N, 9 ° 57 ' 42 " W, 20. xi. - 01. xii. 2017, MV Light Trap (125 W), Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L. leg., ANHRT: 2017.33, slide No.: LGNA 595; 1 male, 530 m, Lofa County, Foya Proposed Protected Area, 7 ° 56 ' 36 " N, 10 ° 16 ' 36 " W, 10 - 19. xi. 2017, MV Light Trap (125 W), Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L. leg., ANHRT: 2017.33; 1 male, 883 m, Lofa County, Wologizi Mts, Ridge Camp 2, 8 ° 07 ' 20.79 " N, 9 ° 56 ' 50.75 " W, 22 - 31. xi. 2018, Light Trap (blended bulb 250 W), Sáfián, Sz., Simonics, G. leg. ANHRT: 2018.43; 1 male, 494 m, Nimba county, Nimba Mts., ENNR, Grassfield FDA Field Station, 7 ° 29 ' 32.45 " N, 8 ° 34 ' 54.05 " W, 22. xi. 2018 Light Trap, Blended Bulb (250 W), Sáfián, Sz., Simonics, G. leg. ANHRT: 2018.43; 1 male, 700 m, ENNR, Nimba Mts, Cellcom Rd, 7 ° 32 ’ 47.5 ” N, 8 ° 32 ’ 1.33 ” W, 10 - 24. III. 2017, Light Trap (250 W blended bulb) & cold cathode UV light bucket trap (8 W), Sáfián, Sz., Simonics, G. leg., ANHRT: 2017.36; 2 males, 701 m, Nimba Mountains, Mount Gangra western slope, 7 ° 33 ' 29.73 " N, 8 ° 38 ' 16.4 " W, 16 - 17. III. 2017, Light Trap (250 W blended bulb) & cold cathode UV light bucket trap (8 W), Sáfián, Sz., Simonics, G. leg., ANHRT: 2017.36 (ANHRT). Ivory Coast. 1 male, 1171 m, Mt. Tonkoui Peak N 07 ° 27 ' 15.2 ", W 07 ° 38 ' 12.5 " 3 - 7. V. 2017, MV light, Aristophanous, A., Aristophanous, Geiser, M., Moretto, P. leg., ANHRT: 2017.25, slide No.: LGNA 594; 2 males, same site, 9 - 16. IV. 2016, Light Trap, Aristophanous, M., Moretto, P. leg., ANHRT: 2017.17. Guinea. 1 male, 1500 m, Nimba Mts, SMFG concession area (Societe des Mines de Fer de Guinee) 600 Forest, 7 ° 39 ' 49.93 " N, 8 ° 22 ' 22.19 " W, 21 - 30. viii. 2017, Cold Cathode UV Light Trap (8 W), Sáfián, Sz., Simonics, G. leg., ANHRT: 2017.36, slide No.: LGNA 835; 1 female, 700 m, Nimba Mts, SMFG concession area, (Societe des Mines de Fer de Guinee), Cite 1, 7 ° 42 ' 2.83 " N, 8 ° 23 ’ 58.60 " W, 16 - 25. VII. 2017, General coll. at light, Sáfián Sz. leg., ANHRT: 2017.36, slide No.: LGNA 998 (ANHRT). Ghana. 1 female, Bunso Arboretum, 6 ° 15 ' 58.03 " N, 0 ° 27 ' 45.72 " W, 13 - 14. IX. 2010, leg. Sz. Sáfián, slide No.: LGNA 990 (HNHM).	en	László, Gyula M. (2020): On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina 30: 82-92, DOI: 10.37828/em.2020.30.8, URL: http://dx.doi.org/10.37828/em.2020.30.8
183FDF25FFDEA568FF6DFE1FFDFFFBA0.taxon	materials_examined	Holotype. Female. “ Zambia, 1147 m, Lukwakwa, West Lunga NP (Cryptosepalum forest / Dambo), S 12 ° 39 ' 40 ", E 24 ° 26 ' 13 ", 9 - 15. XI. 2018, MV Light Trap, Aristophanous, M., Dérozier, V., László, G., Oram, D. leg., ANHRT: 2018.40 ”, unique number: ANHRTUK 00056751, slide No.: LGNA 692 (ANHRT). Paratypes. Zambia. 15 males, 1 female, with the same data as the holotype, unique numbers: ANHRTUK 00055289, 00055290, 00055291, 00055293, 00055331, 00055332, 00055333, 00055435, 00055436, 00058751, 00058754, 00061221, 00061223, 00061249, 00066639, 00072250; 2 males, same data but collected by Lepi LED Light Trap, unique numbers: ANHRTUK 00070756, 00074154; 1 male, same data but collected by Actinic Light Trap, unique number: ANHRTUK 00056895; 2 males, same locality but collected at 4 - 8. XI. 2013 by Smith, R., Takano, H. & Oram, D., ANHRT: 2017.10, unique numbers: ANHRTUK 00073451, 00073452, slide Nos: LGNA 60, LGNA 93; 1 male, 1400 m, Hillwood, Ikelenge, S 11 ° 16 ' 02 ", E 24 ° 18 ' 59 ', 30. IV. - 11. V. 2014, Light Trap, leg. Smith, R., Takano, H., Chmurova, L. & Smith, L., ANHRT: 2017.11, unique number: ANHRTUK 00073453, slide No.: LGNA 198; 3 males, same locality, but collected at 25 - 27. XI. 2014 by Smith, R. & Takano, H., ANHRT: 2017.12, unique numbers: ANHRTUK 00001290, 00001289, 00073454, slide Nos: LGNA 415, LGNA 392, LGNA 292; 1 female, 1365 m, Jan Fisher’ s Farm, Chingola (Miombo / Riverine forest mosaic, 12 ° 36 ’ 10 ” S, 27 ° 55 ’ 48 ” E, 14 - 17. XI. 2019, MV Light Trap, Bashford, M., Miles, W., Mulvaney, L. leg., ANHRT: 2019.25, unique number: ANHRTUK 00132526 (ANHRT).	en	László, Gyula M. (2020): On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina 30: 82-92, DOI: 10.37828/em.2020.30.8, URL: http://dx.doi.org/10.37828/em.2020.30.8
183FDF25FFDEA568FF6DFE1FFDFFFBA0.taxon	diagnosis	Diagnosis. E. inopinata is an allopatric sister species of E. cretacea distinguished by the following characters: E. inopinata has a considerably darker forewing ground colour with more sharply defined, dark greyish transverse lines and with much wider blackish irregular patches in the postmedial area compared to those of E. cretacea; in addition, the postmedial line of the new species is continuous, whereas that of the sister species consists of a row of dots. In the male genitalia, the differences between the two taxa are subtle, the new species has a somewhat shorter uncus, a narrower, medially less dilated valvae, and slightly shorter and narrower harpe compared to E. cretacea. The differences between the two species are well expressed in the configuration of the female genitalia: E. inopinata has slightly shorter papillae anales, much shorter and thinner apophyses, somewhat wider and shorter goblet-shaped ostium bursae, a somewhat thicker ductus bursae with considerably shorter medial sclerotization, a conspicuously shorter and thicker tubular distal part and much more elongated proximal part of corpus bursae compared to those of E. cretacea; however, the most conspicuous difference between the two species is expressed by the complete lack of signum bursae in E. inopinata, while its West African sister species has a pair of heavily sclerotized semi-circular signa bursae.	en	László, Gyula M. (2020): On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina 30: 82-92, DOI: 10.37828/em.2020.30.8, URL: http://dx.doi.org/10.37828/em.2020.30.8
183FDF25FFDEA568FF6DFE1FFDFFFBA0.taxon	description	Description (Figs 9 - 12). Forewing length 9.5 - 11 mm in males, 11 - 12 mm in females. Antenna filiform in both sexes with somewhat longer and denser ciliation in males compared to that of females. Head relatively large, labial palps medium long, curved dorsad, inner surface greyish white, lateral and ventral surface of first and second segments dark blackish grey, third segment pale greyish white; frons and vertex pale greyish white; compound eyes moderately large, globular. Thorax whitish grey, basal twothird of collar darker greyish, abdomen creamy white – brownish white striped. Intraspecific variability limited, all known specimens are rather similar in coloration without noticeable alterations in wing pattern. Sexual dimorphism is moderately expressed in size (females slightly larger) and in coloration, namely females have slightly darker forewing and considerably darker hindwing ground colour compared to those of males. Forewing relatively long and broad, rather quadrangular, apically rounded. Forewing ground colour pale graphite grey, with blackish grey irregular patches in the postmedian area. Sub-basal and basal lines diffuse, shadow-like, represented by groups of dark grey hair scales; antemedial line fine, dark grey, interrupted, almost straight, running angularly towards costa where it joins the medial line, forming a small but conspicuous dark grey quadrangular costal patch; medial line rather thick, slightly diffuse and shadow-like, medially angled, with lower section reaching ventral margin in right angle. Orbicular stigma rather large, rounded, consisting of raised pale whitish grey scales. Postmedial line relatively thick, very sharply defined by blackish scales, costal section broadly arcuate, ventral section straight, running parallel with medial line. Subterminal line diffuse, interrupted, shadow-like, consisted of dark graphite grey patches of different size; terminal line very fine, undulating, nearly continuous, blackish; terminal area with dark grey suffusion; cilia pale brownish grey chequered with dark grey. Hindwing pale whitish grey in basal half, gradually darkened towards termen in males, darker grey in females; cilia whitish grey in both sexes. Underside of forewing uniformly dark graphite grey, that of hindwing pale graphite grey, darkened along the margin in males, similar but darker in females, without traces of pattern. Male genitalia (Figs 21, 22). Uncus relatively short, rather narrow, distally slightly dilated, apically with a short, fine, pointed spine; subscaphium rather long and narrow poorly sclerotized; tegumen short and broad, rather triangular; valvae weakly sclerotized, medium long, medially moderately dilated, apically broadly rounded, tongue-shaped; harpe spike-like, rather long, relatively thick, slightly arcuate, apically pointed, situated in the central plate of valva; transtillae well-developed, semi-circular, medially divided; sacculus long and broad, basal third more strongly sclerotized than apical one; fultura inferior (juxta) very small, with a pair of ear-like rounded lobes; vinculum rather broad and long, V-shaped, saccus short, apically acutely pointed. Aedeagus short and narrow, caecum penis rather long, evenly rounded; carina without sclerotization; vesica membranous in full length, without cornuti. Female genitalia (Fig. 16). Ovipositor short, trapezoidal, papillae anales conical; apophyses posteriores and anteriores rather short, conspicuously thin, posterior ones slightly longer than anteriors; 8 th tergite conspicuously short, weakly sclerotized, ribbon-like; ostium bursae moderately sclerotized, goblet-shaped, relatively short and narrow; ductus bursae very short, relatively narrow, medial quarter more strongly sclerotized than distal and proximal quarters, slightly dilated, cervix bursae membranous, rather thick, sack-like; corpus bursae membranous, distal third tubular, rather thick, dilated anteriorly, proximal half elongate ovoidal; signum bursae absent.	en	László, Gyula M. (2020): On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina 30: 82-92, DOI: 10.37828/em.2020.30.8, URL: http://dx.doi.org/10.37828/em.2020.30.8
183FDF25FFDEA568FF6DFE1FFDFFFBA0.taxon	etymology	Etymology. The latin adjective inopinata means unexpected, referring to the surprising occurrence of a new Ezishnola species in NW Zambia.	en	László, Gyula M. (2020): On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina 30: 82-92, DOI: 10.37828/em.2020.30.8, URL: http://dx.doi.org/10.37828/em.2020.30.8
183FDF25FFD2A56AFF6DF91CFB2FFECD.taxon	materials_examined	Holotype. female, “ Kalinzu Forest Ankole Uganda Nov. 1961 R. H. Carcasson ”, “ Coryndon Museum B. M. 1963 - 322 ”, QR code label with unique number: NHMUK 014046240, slide No.: NHMUK 010316559 (NHMUK). Paratype. Uganda. 1 male, Katera Sango Bay, Masaka, Oct. 1960, R. H. Carcasson, QR code label with unique number: NHMUK 014046239, slide No.: NHMUK 010316558 (NHMUK).	en	László, Gyula M. (2020): On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina 30: 82-92, DOI: 10.37828/em.2020.30.8, URL: http://dx.doi.org/10.37828/em.2020.30.8
183FDF25FFD2A56AFF6DF91CFB2FFECD.taxon	diagnosis	Diagnosis. E. carcassoni sp. n. is an East African allopatric sister species of E. creatacea and E. inopinata known so far only from Uganda. The new species differ externally from both sister species by its conspicuously interrupted transverse lines consisting of a dense row of sharply defined blackish dots, whereas the crosslines are more continuous in the other species, although similar wing pattern is not unknown in E. cretacea. In the male genitalia, E. carcassoni differs from both sister species by its somewhat shorter uncus and valva, in addition more robust, almost straight harpe (those of the other two species are thinner and slightly arcuate) and considerably narrower, more elongate transtillae. In the female genitalia, the new species has somewhat shorter apophyses than those of E. cretacea, ovoid corpus bursae without tubular section and without sack-like cervix bursae (corpus bursae with conspicuously long tubular section and cervix bursae is sack-like in E. cretacea), in addition E. carcassoni lacks signum bursae. The differences between the female genitalia of the new species and E. inopinata are as follows: E. carcassoni has somewhat longer apophyses, narrower ductus bursae with longer sclerotized medial section compared to those of E. inopinata, in addition less elongate, more rounded corpus bursae with simple cervix bursae, while the corpus bursae of E. inopinata is more elongated with anteriorly dilated distal tubular section and its cervix bursae with a well-developed, membranous sack.	en	László, Gyula M. (2020): On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina 30: 82-92, DOI: 10.37828/em.2020.30.8, URL: http://dx.doi.org/10.37828/em.2020.30.8
183FDF25FFD2A56AFF6DF91CFB2FFECD.taxon	description	Description (Figs 13, 14). Forewing length 12 mm in male, 13 mm in female. Antenna filiform in both sexes with somewhat longer and denser ciliation in males compared to that of females. Head relatively large, labial palps medium long, curved dorsad, inner surface brownish white, lateral and ventral surface of first and second segments brownish grey, third segment pale creamy white; frons and vertex pale brownish white; compound eyes moderately large, globular. Thorax brownish grey, basal twothird of collar darker greyish, abdomen brownish white. Intraspecific variability limited. Sexual dimorphism is moderate, expressed in size (female slightly larger) and in coloration, (female hindwing somewhat darker than that of male). Forewing relatively long and broad, rather quadrangular, apically rounded. Forewing ground colour pale brownish grey, with some darker brown irregular patches in the postmedian area. Sub-basal and basal lines diffuse, shadow-like, represented by groups of brownish hair scales; antemedial line fine, brownish grey, interrupted, almost straight, running angularly towards costa; medial line rather thick, interrupted, diffuse, present only in the ventral third; Orbicular stigma rather large, rounded, consisting of raised pale whitish grey scales. Postmedial line relatively thick, interrupted, very sharply defined by dense row of dark brown scales, upper section broadly arched, lower section running parallel with medial line. Subterminal line rather undulating, interrupted, consisting of dark brown triangular patches of different size; terminal line thick, interrupted, consisting of a row of dark brown streaks; terminal area with some red-brown suffusion; cilia pale brownish grey chequered with darker brown. Hindwing pale greyish white, with some darker brownish suffusion terminally in males, darker grey in females, turning to brownish grey in the terminal area; cilia whitish grey in both sexes. Underside of forewing uniformly dark brownish grey, that of hindwing paler brownish grey, darkened along the margin in males, similar but darker in females, without traces of pattern. Male genitalia (Fig. 23). Uncus relatively short, rather narrow, gradually tapered distally, apically with a short, fine, pointed hook; subscaphium rather long and narrow poorly sclerotized; tegumen short and broad, rather triangular; valvae weakly sclerotized, relatively short, medially moderately dilated, apically broadly rounded, tongue-shaped; harpe thorn-like, rather long and robust, almost straight, apically curved and pointed, situated in the central plate of valva; transtillae well-developed, elongate-ovoid, medially divided; sacculus long and broad, basal third more strongly sclerotized than apical one; fultura inferior (juxta) very small, with a pair of ear-like rounded lobes; vinculum rather broad and long, broad V-shaped, saccus short, apically acutely pointed. Aedeagus short and narrow, caecum penis rather long, evenly rounded; carina without sclerotization; vesica membranous in full length, without cornuti. Female genitalia (Fig. 17). Ovipositor short, trapezoidal, papillae anales conical; apophyses posteriores and anteriores short, relatively thin, of same length; 8 th tergite conspicuously short, weakly sclerotized, ribbon-like; ostium bursae moderately sclerotized, goblet-shaped, relatively short and narrow; ductus bursae very short, relatively narrow, medial half more strongly sclerotized than distal and proximal quarters, cervix bursae membranous, rather thick, without sack-like protrusion; corpus bursae membranous, elongate-ovoid without distal tubular part; signum bursae absent.	en	László, Gyula M. (2020): On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina 30: 82-92, DOI: 10.37828/em.2020.30.8, URL: http://dx.doi.org/10.37828/em.2020.30.8
183FDF25FFD2A56AFF6DF91CFB2FFECD.taxon	etymology	Etymology. The new species is named after the British entomologist Robert Herbert Carcasson, collector of the type specimens.	en	László, Gyula M. (2020): On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina 30: 82-92, DOI: 10.37828/em.2020.30.8, URL: http://dx.doi.org/10.37828/em.2020.30.8
183FDF25FFD2A56AFF6DF91CFB2FFECD.taxon	discussion	Remark. An unidentified specimen is illustrated in Hacker et al. (2012) on p. 529 from Budongo Forest, Uganda. Based on its external habitus, this specimen almost certainly belongs to E. carcassoni.	en	László, Gyula M. (2020): On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina 30: 82-92, DOI: 10.37828/em.2020.30.8, URL: http://dx.doi.org/10.37828/em.2020.30.8
