taxonID	type	description	language	source
1F5C2C08FF88F82EFF62FB97FEB925BA.taxon	description	Description. MALE (holotype, range measurements for 2 paratypes in parantheses, Figs. 1 – 2). Idiosoma 590 long (575 – 580), 350 wide (310 – 335). Prodorsal shield length 165 (160 – 165), width 130 (125 – 140), not fused with scapular shields. Distance between bases of setae se 115 (115 – 120), posterior margin concave, lateral margins with incisions near posterior angles. Setae c 2 short, filiform, about 30 (24 – 26) in length, situated on medial margins of humeral shields. Setae c 3 enlarged in basal part and filiform in apical part, 150 (160 – 175) in length. Hysteronotal shield with straight anterior margin, anterior angles rounded, with strongly sclerotized median band stretching from level of trochanters IV to supranal concavity; length from anterior margin to bases of setae ps 1 330 (325 – 330), width at anterior margin 105 (105 – 110). Area between prodorsal and hysteronotal shields wide, distance along midline between these shields 80 (80 – 95). Width of opisthosoma at level of setae f 2 90 (85 – 90). Opisthosomal lobes small, nearly rectangular, with slightly acute inner angle and blunt lateral angle. Terminal cleft U-shaped, length 25 (20 – 25), supranal concavity opens posteriorly. Setae e 1 and openings gl approximately at the same transverse level. Lengths of dorsal setae: c 2 - d 2 175 (170 – 175), d 2 - e 2 180 (170 – 175), d 2 - gl 95 (70 – 90), e 2 - h 2 70 (62 – 70), h 2 - h 2 60 (50 – 60), h 3 - h 3 45 (45 – 46), ps 1 - h 3 8 (6 – 7). Transventral sclerite shaped as a thin Y, length along midline 10 (8 – 12), width of neck part 8 (6 – 8). Epiandrum shaped as an inverted U, fused with posterior end of transventral sclerite, tips of epiandrum extending considerably beyond base of genital apparatus. Length of genital apparatus 24 (25 – 24), width at base 13 (14 – 16), aedeagus minute. Setae g situated on base of genital arch. Setae 4 a and posterior pair of genital papillae at level of midlength of genital arch. Anal suckers ovate, their size excluding surrounding membrane: longer diameter 14 (10 – 12), shorter diameter 8 (6 – 7). Adanal shield shaped as an inverted Y, narrow, not completely encircling anal field and not fused posteriorly. Ventral measurements: 4 b- 4 a 66 (64 – 66), 4 a-g 28 (25 – 26), ps 3 - ps 3 24 (24 – 30), ps 3 - h 3 42 (38 – 40). Tarsus III 68 (70 – 76) in length, with acute apical process and 2 small teeth on dorsal longitudinal crest, macroseta r with very thick basal part, setae s represented by macroseta about 1 / 3 of macroseta r, other setae filiform, shorter than segment (Fig. 5 D). FEMALE (1 paratype, Figs. 3 – 4). Idiosoma 555 long, 225 wide. Prodorsal shield: posterior angles fused with scapular shields, posterior margin straight, length of shield 155, width 138, setae se separated by 130. Setae c 2 hairlike, about 20 long, situated on striated tegument. Hysteronotal shield almost rectangular, with anterior margin concave, anterior angles acute and extending laterally, length (from anterior margin to lobar apices) 325, width at anterior margin 150. Area between prodorsal and hysteronotal shields wide, distance along midline between these shields 80. Posterior end of opisthosoma with 1 pair of widely separated opisthosomal lobes bearing bases of setae h 3. Opisthosomal lobes short, attenuate apically, strongly bent medially, lateral margin with blunt-angular membrane. Length of terminal cleft 32 width at lobar bases 58. Setae e 1 and openings gl situated approximately at same transverse level. Dorsal measurements c 2 - d 2 120, d 2 - e 2 125, e 2 - h 3 75, d 2 - gl 135, e 1 - gl 25, h 2 - ps 1 50, h 2 - h 2 170, h 3 - h 3 110. Epigynium approximately semicircular, 50 long, 110 wide. Apodemes of egg-laying opening short, extending to midlevel of trochanters III. Epimerites IVa absent.	en	Constantinescu, Ioana Cristina, Chişamera, Gabriel, Mukhim, Khlur B., Adam, Costică (2014): Two new feather mite species of the family Pteronyssidae (Acarina: Analgoidea) from Meghalaya (Northeast India). Zootaxa 3774 (4): 351-366, DOI: 10.11646/zootaxa.3774.4.4
1F5C2C08FF88F82EFF62FB97FEB925BA.taxon	materials_examined	Type material. Male holotype (ANA 091), 2 male (ANA 093) and 1 female (ANA 092) paratypes from Actinodura cyanouroptera (Hodgson) (Passeriformes: Leiothrichidae); INDIA: Meghalaya, East Jaintia Hills District, the region of Khahnar village, 25 ° 21 ' 57.30 " N, 92 ° 36 ' 51.72 " E; 20 February 2013, coll. C. Adam. Differential diagnosis. Of the five previously known species, Timalinyssus actinodurae sp. nov. is closest to T. formosanus Mironov, 2001 from Actinodura morrisoniana Ogilvie-Grant (Leiothrichidae) (Mironov 2001). Males in both species have tarsus III moderately elongated, with an acute apical process and teeth on the longitudinal dorsal crest, the epiandrum shaped as an inverted U with tips extending considerably beyond the base of the genital apparatus, a similar shape of the genital apparatus, and the adanal shield shaped as an inverted Y. Males of the new species clearly differ from those of T. formosanus due to the following features: the prodorsal shield is free from the scapular shields, the hysteronotal shield has a straight anterior margin and heavily sclerotized median band, the anterior angles of the hysteronotal shield are rounded, the setae ps 1 are situated almost apically, near the inner angle of the lobes, the transventral sclerite is present, the setae s are represented by macrosetae, and the setae d are hair-like. In males of T. formosanus, the prodorsal shield is fused with the scapular shields, the hysteronotal shield has the anterior margin slightly sinuous, its anterior angles are acute and a sclerotised band in median band is absent, the setae ps 1 are situated clearly distant from the inner angle of the lobes, the transventral sclerite is absent, the setae d are represented by macrosetae, and the setae s are filiform. Females in both species have the prodorsal shield fused with the scapular shields and clearly differ from T. formosanus, as well from the other three previously known females of the genus (T. oliferae (Mironov), T. longitarsus Wang and Wang, T. curvilobus Mironov), due to the shape of the opisthosomal lobes that are short, apically attenuate, strongly bent medially, and with blunt-angular lateral membrane. In females of T. formosanus, and also in T. oliferae, the opisthosomal lobes are long and roughly ovoid, while in T. curvilobus the lobes are indistinct and only short cone-like extensions are present, and, in females of T. longitarsus, they do not have lobes. Furthermore, females of T. actinodurae differ from T. formosanus due to the absence of epimerites IVa (rudiments of epimerites IVa are present in T. formosanus).	en	Constantinescu, Ioana Cristina, Chişamera, Gabriel, Mukhim, Khlur B., Adam, Costică (2014): Two new feather mite species of the family Pteronyssidae (Acarina: Analgoidea) from Meghalaya (Northeast India). Zootaxa 3774 (4): 351-366, DOI: 10.11646/zootaxa.3774.4.4
1F5C2C08FF88F82EFF62FB97FEB925BA.taxon	etymology	Etymology. The specific epithet actinodurae refers to the generic name of the host species and is a noun in the genitive case.	en	Constantinescu, Ioana Cristina, Chişamera, Gabriel, Mukhim, Khlur B., Adam, Costică (2014): Two new feather mite species of the family Pteronyssidae (Acarina: Analgoidea) from Meghalaya (Northeast India). Zootaxa 3774 (4): 351-366, DOI: 10.11646/zootaxa.3774.4.4
1F5C2C08FF81F824FF62FF52FA03268E.taxon	description	Description. MALE (holotype, range measurements for 5 paratypes in parentheses, Figs. 6 – 7) Idiosoma 385 long (370 – 390), 190 wide (180 – 200). Prodorsal shield with posterolateral angles not expressed, lateral margins with incisions around bases of setae se, posterior margin straight, 115 long (105 – 120) and 100 (95 – 105) wide, surface with granular ornamentation, distance between setae se – se 85 (80 – 85). Setae c 2 filiform 20 (25 – 30) long, setae c 3 lanceolate 32 (35 – 40) long. Prodorsal and hysteronotal shields separated by large area with transverse striae, distance between this shields along midline 50 (40 – 65). Hysteronotal shield entire, not encompassing bases of setae c 1, with rounded anterior angles, with slightly convex and rough anterior margin, and with shallow incisions on lateral margins at level of humeral shields, greatest length 210 (195 – 215), width at anterior margin 110 (110 – 125). Opisthosomal lobes short and rounded, terminal cleft U – shaped, 25 (25 – 40) long. Supranal concavity opened posteriorly, anterior end extending slightly above level of setae f 2. Distal, inner and lateral margins of lobes with narrow entire membrane, width of opisthosoma at level of setae f 2 85 (80 – 85). Dorsal setae e 1 situated posterior to openings gl. Lengths of dorsal setae: d 2 14 (15 – 20), e 2 26 (26 – 35). Dorsal measurements: c 2 - d 2 70 (62 – 70), d 2 - e 2 95 (90 – 100), d 2 - gl 30 (24 – 30), e 2 - h 2 50 (45 – 55), h 2 - h 2 60 (55 – 70), h 3 - h 3 50 (40 – 50), ps 1 - h 3 15 (10 – 15). Transventral sclerite with nearly straight anterior margins in the middle area, and fused with epiandrum, length along midline 10 (9 – 10). Epiandrum long, completely encompassing genital apparatus, tips of epiandrum thin, extending beyond level of setae 4 a. Genital apparatus very long, 68 (68 – 80) in length, 20 (20 – 25) in width, aedeagus minute, much shorter than genital apparatus. Seatae 4 a and g approximately at same level. Diameter of anal suckers 20 (20 – 30). Adanal shield having a T or cross-shape, poorly sclerotized. Ventral measurements: 3 a- 4 a 55 (55 – 65), g- 4 a 16 (16 – 25), g-ps 3 66 (66 – 75), ps 3 - ps 3 24 (25 – 30), ps 3 - h 3 46 (46 – 55). Tarsus III 56 (65 – 70) long, with narrow bidentate apex, seta r slightly longer than segment (Fig. 10 C). Tarsus IV shorter than tibia IV, setae d and e with apical caps, close to each other, situated near apex of segment (Fig. 10 D). FEMALE (1 paratype, Figs. 8 – 9). Idiosoma 440 long, 80 wide. Prodorsal shield as in male, 135 long, 105 wide, distance se: se 80. Setae c 2 filiform, 20 long; setae c 3 lanceolate, 40 long. Arrangement of hysteronotal shields: unpaired anterior hysteronotal sclerite, central sclerite, pair of lateral opisthosomal sclerites and pair of pygidial sclerites. Anterior hysteronotal sclerite represented by transverse plate narrowed in median part, with all margins irregular, with anterior margin strongly concave, encompassing bases of setae c 1, c 2. Anterior hysteronotal sclerite and central sclerite separated by narrow band of soft tegument with 5 striae. Central sclerite of complicate form: (Fig. 8), attenuated posteriorly, anterior margin straight, posterior end roughly romb-like, length along midline 120 and width at anterior margin 75; setae e 1 off this sclerite. Lateral opisthosomal sclerites represented by plates of irregular form, bearing setae e 2, openings gl situated in antero-median margin of these shields, surface with 5 oblique striae. Pygidial sclerites small, roughly triagular shape, encompassing bases of setae h 2, h 3 and ps 1; seta f 2 inserted laterally, on soft tegument. Subhumeral seta c 3 slightly curved, longer than trochanter III, 32 long. Dorsal measurements: c 2 - d 2 115, d 2 - e 2 90, e 2 - h 3 85, d 2 - gl 75, e 1 - gl 50, h 2 - ps 1 20, h 2 - h 2 75, h 3 - h 3 60.	en	Constantinescu, Ioana Cristina, Chişamera, Gabriel, Mukhim, Khlur B., Adam, Costică (2014): Two new feather mite species of the family Pteronyssidae (Acarina: Analgoidea) from Meghalaya (Northeast India). Zootaxa 3774 (4): 351-366, DOI: 10.11646/zootaxa.3774.4.4
1F5C2C08FF81F824FF62FF52FA03268E.taxon	materials_examined	Type material: Male holotype (ANA 084), 5 male (ANA 086 – ANA 090) and 1 female (ANA 085) paratypes, from Hemixos flavala Blyth (Passeriformes: Pycnonotidae); INDIA: Meghalaya, East Jaintia Hills District, the region of Khahnar village, 25 ° 21 ' 57.30 " N, 92 ° 36 ' 51.72 " E, 6 February 2013, coll. C. Adam. Differential diagnosis. Pteroherpus meghalayensis sp. nov. belongs to the hoplophorus species group (Mironov & Wauthy 2008). Males of this species-group have a prodorsal shield with a short posterior part, and a bluntly rounded posterior margin, the posterolateral angles are not expressed, and the distance from the anterior margin of this shield to the level of the setae se is equal with the distance between the bases of these setae. The opisthosomal lobes are short and wide, the posterior margin is bluntly rounded, the terminal cleft is U-shaped, and the setae ps 2 are filiform. The female has an unpaired anterior hysteronotal sclerite, and the opisthosomal sclerite are whole and separated from the central sclerite. The male of Pteroherpus meghalayensis is distinguished from all other species of the genus due to a very long genital apparatus and the unusual length of the branches of the epiandrum extending beyond the base of the genital apparatus. Among previously described species of the genus, Pteroherpus meghalayensis sp. nov. is closest to P. chlorocichlae Mironov and Wauthy, 2008 because the hysteronotal shield and the opisthosomal lobes in male have a similar shape and there is also a similar arrangement of the hysterosomal shields in females (unpaired anterior hysteronotal sclerite as transverse plate, central sclerite, lateral opisthosomal sclerites and pygidial sclerites). The male of the new species is distinguished from P. chlorocichlae due to the following features: the absence of the dorsal additional small sclerites, the transventral sclerite with a straight anterior margin, the extension of the tips of epiandrum beyond the basis of the genital apparatus, the extreme length of the genital apparatus (68 - 80), and the T or cross-shape of adanal shield. In males of P. chlorocichlae, a pair of very small additional sclerites is present near the posterior end of the prodorsal shield, the transventral sclerite has concave anterior margins, the branches of epiandrum extend to the midlevel of the genital apparatus, the genital apparatus length is 25, and the adanal shield has a distinctly concave anterior margin. Females of P. meghalayensis differ from those of P. chlorocichlae due to the shape of the anterior hysteronotal and central sclerites; the anterior hysteronotal sclerite has an irregular margin and the central sclerite has a rhomboid shape in the posterior half, while in P. chlorocichlae, the anterior hysteronotal sclerite has a straight margin and the central sclerite is almost parallel-sided and only in the posterior half is rounded.	en	Constantinescu, Ioana Cristina, Chişamera, Gabriel, Mukhim, Khlur B., Adam, Costică (2014): Two new feather mite species of the family Pteronyssidae (Acarina: Analgoidea) from Meghalaya (Northeast India). Zootaxa 3774 (4): 351-366, DOI: 10.11646/zootaxa.3774.4.4
1F5C2C08FF81F824FF62FF52FA03268E.taxon	etymology	Etymology. The specific name meghalayensis refers to the state of Meghalaya where the mite was collected.	en	Constantinescu, Ioana Cristina, Chişamera, Gabriel, Mukhim, Khlur B., Adam, Costică (2014): Two new feather mite species of the family Pteronyssidae (Acarina: Analgoidea) from Meghalaya (Northeast India). Zootaxa 3774 (4): 351-366, DOI: 10.11646/zootaxa.3774.4.4
