taxonID	type	description	language	source
1D3687F8A761A668FF62FAAFFBCC75B8.taxon	description	Figs 1 - 4.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A761A668FF62FAAFFBCC75B8.taxon	materials_examined	Type material. Vietnam, Vinh Ha Long Prov. (SW), Dao Bo Hon, Hang Bo Nau Cave, 12. VI. 2003, leg. P. Trontelj & B. Sket, holotype ♂ (MNHN JC 309), paratypes: 1 ♂ (MNHN JC 309), 1 ♀ (SEM).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A761A668FF62FAAFFBCC75B8.taxon	etymology	Name. To emphasize the obvious basal position amongst the volvatory “ doratodesmids ”.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A761A668FF62FAAFFBCC75B8.taxon	diagnosis	Diagnosis. Differs from congeners by the apparently imperfect volvation, due to particularly short paraterga, coupled with an especially simple gonopod structure.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A761A668FF62FAAFFBCC75B8.taxon	description	Description: Length of adults of both sexes ca 4.8 - 5.0 mm, width 0.6 - 0.65 mm, body broadest at segment 3 or 4; ♂♂ a little smaller than ♀♀. Holotype ca 4.8 mm long and 0.6 mm wide. Coloration uniformly pallid or light yellowish. Adults with 20 segments, body subcylindrical (Fig. 1 A), pattern of conglobation typical of “ Doratodesmidae ” as described by Golovatch (2003), but volvation itself obviously somewhat imperfect in barely concealing all (especially hind) legs (Figs 1 A, C, D). Head (Fig. 2 B) slightly transverse, rather densely pilose, microgranular amd microvillose just below antennae and on vertex, with a pair of paramedian, almost contiguous knobs; isthmus between antennal sockets very narrow. Antennae (Fig. 2 C) very short and stout; antennomere 6 longer than 5, dorso-apically with an evident pit containing a tight group of minute bacilliform sensilla; antennomere 8 with the usual four sensory cones apically. Collum rather large, regularly convex, not covering the head from above, entire surface microvillose, with four transverse rows of round tubercles with a pit on top (apparently representing the former place of insertion of bisegmented, tactile setae, which are mostly lost) (Fig. 2 A). Prozona very finely alveolate, collum and metaterga covered with a cerotegumental crust held by abundant microvilli; stricture between pro- and metazona broad and shallow, more finely alveolate-microgranular than prozona; limbus microcrenulate, nearly fully hidden by nearby abundant microvilli (Figs 3 C, D). Metaterga behind collum with three transverse rows of tubercles (Figs 1 A-F), some of which still retain long, bisegmented setae (Fig. 3 B). Paraterga subvertical, very narrow, downwards barely reaching level of venter, clearly trisinuate caudolaterally at base due to two lobulations (Figs 1 C, D; 2 D; 3 A); paraterga 2 strongly enlarged, with a series of lobulations anterolaterally, both schism and hyposchism very small; paraterga 3 and 4 slightly shorter than others (Fig. 1 B), overlap of following paraterga typical, latter broadly rounded and evidently trilobate. Pore formula normal, ozopores located on top of a small porostele-like tubercle on ventrocaudal lobulation (Fig. 3 A). Pleurotergal ridges very small, alveolate-microgranular like entire ventral surface. Epiproct short, also with tubercles, directed ventrocaudad, with the usual four cones (= spinneret setae) just below tip; para- and hypoprocts as in Figs 1 F; 2 E. Sterna usually with a deep, narrow, longitudinal depression between coxae (Fig. 2 F), but sterna between coxae 6, 7 and 9 much wider (Fig. 4 A). Gonopod aperture transverse-oval, relatively small, far from reaching lateral sides of segment 7. Legs very short; tarsal segment longest; claw usual, simple, very slightly curved ventrad; some setae sparsely microdenticulate (Figs 2 F; 3 E). Gonopods (Figs 4 B, C) very simple. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally, with a conspicuous triangular lobe frontolaterally. Telopodite much longer than coxite, slender throughout, setose in its basal half, with a conspicuous, only apically denticulate, lateral, distofemoral process (dp) at about midway, with a hairpad in distal third, but seminal groove unexpectedly terminating subapically.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A761A668FF62FAAFFBCC75B8.taxon	discussion	Remarks. This remarkable species somewhat bridges the former Haplodesmidae and Doratodesmidae. Hence the quotation marks which are used for “ haplodesmids ” and “ doratodesmids ” above and hereafter. In addition to showing the tergal trichome of peculiar, bisegmented setae and the unusually simple gonopods so characteristic of Cylindrodesmus (both features partly shared also with several true “ doratodesmids ”), it also, most importantly, has the body only capable of incomplete volvation, due to the relatively very short paraterga. Moreover, the ozopores are borne on small porostele-like tubercles, a feature characteristic of most “ haplodesmids ”, but only rarely encountered among “ doratodesmids ”. At least the incomplete volvation and the simple biramous gonopods seem to be relatively primitive characters amongst the more advanced “ doratodesmids ” in which volvation is already perfect. The new species especially resembles the similarly tuberculate-setose Dyomerothrix gremialis Hoffman, 1982 or Parapauroplus monodentus Zhang in Zhang & Wang, 1993, which also show rather simple gonopods (Hoffman 1982 a; Zhang and Wang 1993).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A76AA66BFF62FE7CFCFC7249.taxon	description	Figs 5 - 8.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A76AA66BFF62FE7CFCFC7249.taxon	materials_examined	Type material. Vietnam, Thanh Hoa Prov., Pu Luong, Lung Cao, Hang Lang Lua Cave, 11. XII. 2003, leg. L. Deharveng & team (Vn 0312 - 05), holotype ♂ (MNHN JC 310), paratypes: 1 ♂, 1 juv. (MNHN JC 310), 1 ♂ (ZMUM); Ha Nam Ninh Prov., Cuc Phuong National Park, Cave of Prehistoric Man, 10. X. 1998, leg. L. Deharveng (VIET- 537), 1 ♂ (MNHN JC 310); same locality, Water Fairy Cave, 11. X. 1998, leg. L. Deharveng (VIET- 541), 1 ♂, 2 juv. (MNHN JC 310); same locality, cave 4 (Son Cung Cave?), 21. VI. 2003, leg. P. Trontelj & B. Sket, 1 ♂, 1 ♀, 3 juv. (MNHN JC 310), 1 ♂, 1 juv. (ZMUC), 1 ♂, 1 juv. (NMNHS), 1 ♂ (MCSNV), 1 ♂, 1 juv. (OBBFUL), 1 ♀ (SEM); Ha Nam Ninh Prov., outside Cuc Phuong National Park, cave 3, 20. VI. 2003, leg. P. Trontelj & B. Sket, 1 ♂, 1 ♀ (ZMUM).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A76AA66BFF62FE7CFCFC7249.taxon	etymology	Name. To emphasize the obvious dorsal projections on body segments 17 - 19.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A76AA66BFF62FE7CFCFC7249.taxon	diagnosis	Diagnosis. Most similar to Parapauroplus monodentus Zhang in Zhang & Wang, 1993, which also shows evident mid-dorsal projections only on body segments (16) 17 - 19, but differs by the presence of porosteles, the slightly lower metatergal tubercles devoid of microsetae near each of the main setae, and the slightly stouter gonopod telopodite showing a shorter midway process.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A76AA66BFF62FE7CFCFC7249.taxon	description	Description. Length of adults of both sexes ca 10 - 14 mm, width 2.3 - 2.6 mm, body broadest at segment 3 or 4; ♂♂ usually a little smaller than ♀♀. Holotype ca 11 mm long and 2.5 mm wide. Coloration uniformly pallid or light yellowish. Adults with 20 segments, pattern of conglobation typical of “ Doratodesmidae ” (Figs 5 A). Head (Fig. 6 A) and tegument (Figs 5 B, C; 6 F, G) as in preceding species, but antennae somewhat longer (Fig. 6 A), collum slightly flattened mid-dorsally, not covering the head from above (Fig. 6 A). Metaterga behind collum with three transverse, rather irregular rows of tubercles (Figs 5 A-C), most of which still retain short, bisegmented setae (Fig. 6 G), metaterga 17 - 19 each with an evident dorsal outgrowth (Fig. 5 C). Paraterga mostly directed ventrolaterad, rather broad, slightly surpassing level of venter, clearly trisinuate caudolaterally at base, due to two lobulations (Figs 5 B, D; 6 B); paraterga 2 strongly enlarged, subvertical, margin nearly entire, with only very faint traces of a series of small lobulations anterolaterally (Fig. 6 A), both schism and hyposchism very small; paraterga 3 and 4 slightly shorter than others, overlap of following paraterga typical, latter broadly rounded and slightly 3 - or 4 - lobate. Limbus spiculate (Fig. 6 F). Pore formula normal, ozopores located on top of a small porostele, slightly above ventrocaudal lobulation (Figs 6 C-E). Pleurotergal carinae wanting. Epiproct elongated, also with tubercles, directed ventrocaudad, with the usual four cones just below tip; para- and hypoprocts as in Figs 5 E, F. Sterna usually with a deep, narrow depression between coxae (Figs 5 D; 6 B), only sterna between coxae 6, 7 and 9 much wider (Fig. 7 A). Gonopod aperture transverse-oval, relatively small, far from reaching lateral sides of segment 7. Legs relatively long; femoral and tarsal segments longest and equal; claw simple, very slightly curved ventrad; some setae very sparsely microdenticulate (Figs 5 D; 6 B). Gonopods (Figs 7; 8) very simple. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally, with a conspicuous rounded lobe frontolaterally. Telopodite considerably longer than coxite, slightly stouter, setose in its basal half, with an inconspicuous, digitiform, simple, lateral, distofemoral process (dp) at about midway, more distally with a slightly folded and enlarged shaft (= solenomere) with dense pilosity and a hairpad; seminal groove terminating subapically, acropodite short.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A76AA66BFF62FE7CFCFC7249.taxon	discussion	Remarks. This species seems to be close to some of the “ doratodesmids ” showing evident mid-dorsal projections on some of the metaterga, e. g. Ascetophacus macclurei Hoffman, 1977, Cerastelachys cavernicola (Sinclair, 1901), Pocillidorsus dorsiangulatus Zhang in Zhang & Wang, 1993 and Parapauroplus monodentus Zhang in Zhang & Wang, 1993, etc. (Hoffman 1977 a, 1977 b; Zhang and Wang 1993). Cave 4 also supports another millipede, Glyphiulus mediobliteratus Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2007 (Golovatch et al. 2007 b).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A776A672FF62FBA6FC967681.taxon	description	Figs 9 - 13.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A776A672FF62FBA6FC967681.taxon	materials_examined	Type material. Vanuatu, Espirito Santo, Malo Island, Avorani, forest litter, Berlese extraction, 15. IX. 2006, leg. L. Deharveng & A. Bedos (SK 06 - 15 / 06), holotype ♂ (MNHN JC 311), paratypes: 1 ♂, 4 ♀♀ (MNHN JC 311), 1 ♂, 1 ♀ (ZMUM), 1 ♀ (NMNHS), 1 ♀ (MCSNV), 1 ♀ (ZMUC); same place, date and collectors (SK 06 - 15 / 13), 1 ♂, 1 ♀, 1 juv. (MNHN JC 311); Espirito Santo, Fapon doline 3, 5. IX. 2006, leg. L. Deharveng & A. Bedos (SK 06 - 15 / 18), 5 ♀♀, 4 juv. (MNHN JC 311), 1 ♂ (SEM); Espirito Santo, very humid forest at Boutmas Pass, 27. IX. 2006, leg. L. Deharveng & A. Bedos (SK 06 - 27 / 12), 1 ♂ (MNHN JC 311); Espirito Santo, Boutmas, forest above Jourdain River, Berlese extraction of litter, 27. IX. 2006, leg. L. Deharveng & A. Bedos (SK 06 - 27 / 09), 1 ♂ (MNHN JC 311); Espirito Santo, Boutmas, Fapon doline 2, 5. IX. 2006, leg. L. Deharveng & A. Bedos (SK 06 - 05 / 12), 2 ♀♀ (MNHN JC 311); Espirito Santo, Funatus, Katae Cave, Berlese extraction of guano, 18.09.2006, leg. J. Lips (SK 06 - 18 / 07, Katae 4 gu), 1 ♂, 2 juv. (ZMUC).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A776A672FF62FBA6FC967681.taxon	etymology	Name. To emphasize a superficial transition from a still non-volvatory “ haplodesmid ” condition to the typical, fully volvatory “ doratodesmid ” one.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A776A672FF62FBA6FC967681.taxon	diagnosis	Diagnosis. Differs from congeners by the tergal trichome being composed of only very long, bisegmented, tactile setae, coupled with the gonopod telopodite having a peculiar, dentate-microtuberculate, lateral process at midway and a lobiform, slightly coiled distal part.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A776A672FF62FBA6FC967681.taxon	description	Description. Length of adults of both sexes ca 3.5 - 4.2 mm, width 0.6 - 0.75 mm, body broadest at segment 3 or 4; ♂♂ usually a little smaller than ♀♀. Holotype ca 3.8 mm long and 0.65 mm wide. Coloration uniformly pallid, ranging from white to light yellow or pink. Adults with 19 segments, body subcylindrical (Figs 9; 10 A), pattern of conglobation typical of “ Doratodesmidae ”, but volvation itself barely complete, poorly concealing at least legs of several caudal segments. Head (Figs 10 E, F; 11 A, B) slightly transverse, rather densely pilose, microgranular; isthmus between antennal sockets very narrow. Antennae (Figs 11 C, D) very short and stout; antennomere 6 longer than 5, dorsoapically with an evident pit containing minute bacilliform sensilla; antennomere 8 with usual four sensory cones apically. Collum rather large, even and convex, not covering the head from above, very densely and irregularly setose, setae bisegmented, tactile (Fig. 10 F). Prozona very finely alveolate, collum and metaterga covered with a rather even cerotegumental crust held by microvilli, this crust, like on collum, being perforated only by dense, irregularly arranged, tactile, bisegmented setae; stricture between pro- and metazona broad and shallow, even more finely alveolate than prozona; limbus entire, not hidden by sparse microvilli (Figs 11 F; 12 A-C). Paraterga subvertical and rather narrow, weakly set off by 1 - 2 shallow depressions or sulci, slightly bisinuate caudally at base (Figs 10 E; 11 E); paraterga 2 strongly enlarged, schism and hyposchism very small; paraterga 3 and 4 shorter than others (Figs 9; 10 A, E), following paraterga broadly rounded, overlap typical. Pore formula normal, ozopores located near ventral margin, progressively closer to caudal corner of paraterga. Pleurotergal ridges very small, microgranular like entire ventral surface (Fig. 11 E). Epiproct short, slightly uneven, directed ventrocaudad, below tip with the usual four cones; para- and hypoproct as in Figs 10 C, D. Sterna usually with a minute middle knob separating bases of nearly contiguous coxae (Fig. 12 D), only sterna between coxae 6, 7 and 9 much wider (Fig. 12 E). Gonopod aperture transverse-oval, relatively small, far from reaching lateral sides of segment 7, as in Fig. 12 E. Legs relatively short; coxae and prefemora microgranular; tarsal segment longest; claw usual, simple, very slightly curved ventrad; some setae very sparsely microdenticulate (Fig. 12 D). Coxae 1 of ♂ with a distinct distomedial spine, coxae 2 of ♂ perforated by vas deferens (Fig. 11 B). Gonopods (Figs 12 E, F; 13) rather complex. Coxae subquadrate, very large, scaly, with only a few setae ventrolaterally. Telopodite only slightly longer than coxite, its basal part setose and slender, with a conspicuous, densely dentate-tuberculate, distofemoral outgrowth (dp) at about midway, more distally with an expanded, somewhat folded, coiled, broadly trifid lobe, carrying a very short solenomere branchlet laterally.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A776A672FF62FBA6FC967681.taxon	discussion	Remarks. This remarkable species partly bridges the former Haplodesmidae and Doratodesmidae in showing the tergal trichome of peculiar, bisegmented setae so characteristic of Cylindrodesmus (in which, however, it is mixed with usual setation), also shared with several true “ doratodesmids ”, on the one hand, and the body and gonopod structure of a rather typical “ doratodesmid ”, on the other.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A770A67DFF62FD63FC447141.taxon	description	Figs 14 - 17.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A770A67DFF62FD63FC447141.taxon	materials_examined	Type material. China, Guangxi Prov., Yachang Nature Reserve, Yan Wu Dong Cave, 31.05.2007, leg. L. Deharveng & A. Bedos (CHI-GX 07 - 31 / 01), holotype ♂ (IZAS), paratypes: 2 ♂♂, 6 ♀♀ (MNHN JC 312), 1 ♂ (SEM); same locality, Xia Yan Dong Cave, 28. V. 2007, leg. L. Deharveng & A. Bedos (CHI-GX 07 - 28 / 01), 5 ♂♂ (SCAU); same locality, Xiao Shui Dong Cave, 2. VI. 2007, leg. F. Bréhier (CHI-GX 07 - 02 / 07), 4 ♀♀, 1 juv. (IZAS); same locality, She Dong Cave, 30. V. 2007, leg. F. Bréhier (CHI-GX 07 - 30 / 08), 2 ♂♂, 1 ♀, 1 juv. (SCAU), 1 ♂, 1 ♀, 1 juv. (ZMUM), 1 ♂ (ZMUC), 1 ♂ (NMNHS), 1 ♂ (MCSNV).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A770A67DFF62FD63FC447141.taxon	etymology	Name. To emphasize the broad paraterga.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A770A67DFF62FD63FC447141.taxon	diagnosis	Diagnosis. Differs from all congeners except E. similis sp. n. by the very broad and mostly only slightly declivous paraterga, such that volvation is apparently imperfect, from E. similis sp. n. by the collum bearing no row of peculiar teeth along its entire front margin, and the gonopods being less enlarged and simpler distally.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A770A67DFF62FD63FC447141.taxon	description	Description. Length of adults of both sexes ca 11 - 13 mm, width 2.4 - 3.0 mm, body broadest at segments 5 - 16. Holotype ca 12 mm long and 3.0 mm wide. Coloration uniformly yellow-brown to brownish, sometimes slightly marbled. Adults with 20 segments (Fig. 14 A), conglobation pattern typical of “ Doratodesmidae ”, but volvation apparently imperfect because of paraterga being too broad and mostly only slightly declivous, leaving small lacunae laterally even when body maximally enrolled. Head (Figs 14 F; 15 A) and tegument (Figs 15 E, F) basically as in preceding species, but antennae somewhat longer, antennomeres 3 and 6 of subequal length (Fig. 14 F), collum not covering the head from above, slightly elevated frontolaterally, with 4 - 5 irregular rows of mostly flat tubercles / bosses, including a row of more distinct cones at caudal edge (Figs 15 A, B). Metaterga behind collum with three transverse, rather irregular rows of similarly flat tubercles, most of which still retain short, bisegmented setae (Fig. 15 A, B, D-F); limbus microcrenulate. Paraterga with evident shoulders anteriorly, mostly directed laterad, only slightly declivous, very broad, tips about level with venter, very distinctly crenulate caudolaterally (Figs 14 A-E; 15 C, D); paraterga 2 rather strongly enlarged, directed ventrolaterad, lateral margin evidently quadrilobulate, both schism and hyposchism small; paraterga 3 and 4 slightly shorter than others, bilobate laterally (Figs 14 B; 15 B), following paraterga more broadly rounded and quadrilobulate, overlap typical. Pore formula normal, ozopores located at base of ventrocaudal lobulation (Fig. 15 D). Pleurotergal carinae wanting. Epiproct strongly flattened, dorsally also tuberculate, with several deep incisions at lateral edge, directed ventrocaudad, with the usual four cones just below tip; para- and hypoprocts as in Figs 14 C-E. Sterna usually with a rather deep, narrow depression between coxae (Fig. 16 A), only those between coxae 7 and 9 much wider (Fig. 16 B). Gonopod aperture transverse-oval, relatively small, far from reaching lateral sides of segment 7 (Fig. 16 B). Legs very long and slender; femur somewhat longer than tarsus; several basal segments in ♂ microtuberculate; claw simple, very slightly curved ventrad; some setae very sparsely microdenticulate (Figs 14 F; 15 C; 17 A). Gonopods (Figs 16 B-E; 17 B, C) very simple. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally, with a conspicuous triangular lobe frontolaterally. Telopodite considerably longer than coxite, slender, setose not only in its basal half but also at base of a digitiform and conspicuously microtuberculate distofemoral process (dp) situated laterally at about midway of telopodite, more distally with a slightly folded acropodite devoid of a hairpad; seminal groove terminating subapically.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A770A67DFF62FD63FC447141.taxon	discussion	Remarks. This species seems to be especially close to the next species.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77FA679FF62FA23FBB07166.taxon	description	Figs 18 - 21.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77FA679FF62FA23FBB07166.taxon	materials_examined	Type material. China, Guangxi Prov., Mulun Nature Reserve, Gui Dong 2 Cave, 18. V. 2007, leg. F. Bréhier (CHI-GX 07 - 18 / 20), holotype ♂ (IZAS), paratypes: 2 ♂♂, 2 ♀♀, 2 juv. (MNHN JC 313), 1 ♂, 1 ♀ (ZMUM), 1 ♂ (ZMUC), 1 ♂ (SEM); same locality, Shen Long Dong Cave, 22. V. 2007, leg. L. Deharveng & A. Bedos (CHI-GX 07 - 22 / 01); 1 ♂, 3 ♀♀, 4 juv. (SCAU).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77FA679FF62FA23FBB07166.taxon	etymology	Name. To emphasize the obvious similarities with the preceding species.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77FA679FF62FA23FBB07166.taxon	diagnosis	Diagnosis. See diagnosis of the previous species.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77FA679FF62FA23FBB07166.taxon	description	Description. Length of adults of both sexes ca 11 - 13 mm, width 3.0 - 3.3 mm; ♂♂ usually a little smaller than ♀♀. Holotype ca 12 mm long and 3.0 mm wide. Coloration uniformly pallid or light yellowish, only seldom marbled light-brownish. All characters virtually as in E. latus sp. n. (Figs 18 A-C, E, F; 19 B-F; 20 A, B), but collum with a row of very conspicuous teeth at front margin (Figs 18 D; 19 A). Gonopods (Figs 20 C, D; 21) slightly more complex than in E. latus sp. n. Telopodite distally somewhat expanded and folded, with a hairpad subapically.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77FA679FF62FA23FBB07166.taxon	discussion	Remarks. Together with E. latus sp. n., this remarkable species partly bridges the gap between “ doratodesmoid ” Haplodesmidae and the large tropical family Cryptodesmidae Karsch, 1879 in showing the very broad and, at most, only slightly declivous paraterga so characteristic of cryptodesmids, including those occurring in East and Southeast Asia (Hoffman 1980; Simonsen 1990). However, the metaterga in Cryptodesmidae are often densely setose, the collum is always enlarged, flabellate and covering the head from above to an even greater extent than it does in Pyrgodesmidae, whereas the paraterga are often deeply incised caudolaterally or have clear radii, again much as in some Pyrgodesmidae. Therefore, it seems safe for the present to consider the Cryptodesmidae as a distinct family in the superfamily Polydesmoidea (Hoffman 1980, 1982 b; Simonsen 1990). What is more remarkable, however, is that the above two new Chinese species have the body so strongly flattened, and the paraterga apparently too broad to allow a tight volvation. This condition represents the opposite to that observed for E. basalis sp. n., in which the paraterga are obviously too short to permit complete volvation. In other words, within Eutrichodesmus we find species representing virtually the entire spectrum of “ doratodesmid ” evolution of conglobation, including a few exceptional cases of imperfect volvation. Concerning the two new Chinese species that are not capable of complete conglobation, the situation somewhat parallels that observed in Proeilodesmus mecistonyx Hoffman, 1990 (Sphaeriodesmidae Humbert & DeSaussure, 1869, Sphaeriodesmoidea), a cavernicole from Mexico that is apparently incapable of volvation (Hoffman 1990). In contrast to “ doratodesmids ”, however, most of the characters of this millipede seem plesiomorphic and suggest a link between a presumed flat-bodied chelodesmidean (= leptodesmidean) ancestor and the remaining, convex, truly volvatory members of this rather large, Neotropical family (Hoffman 1990; Golovatch 2003).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77BA647FF62FA06FCA0759B.taxon	description	Figs 22 - 26.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77BA647FF62FA06FCA0759B.taxon	materials_examined	Type material. China, Guizhou Prov., Qianxi County, Hong Lin Village, Tiao Shuz Dong Cave, 18. XI. 2003, leg. L. Latella & G. Rossi, holotype ♂ (MNHN JC 314), paratypes: 1 ♂, 1 juv. (MNHN JC 314), 1 ♀ (SEM); same locality and cave, 18. XI. 2003, leg. L. Latella, D. Avesani & G. Rossi, 1 ♂, 5 ♀♀, 1 juv. (MNHN JC 314), 2 ♀♀ (SCAU); same locality, Liao Jing Ling Dong Cave, 16. XI. 2001, leg. L. Latella, 1 ♂, 1 ♀, 1 juv. (MCSNV), 1 ♀ (IZAS), 1 ♀ (ZMUM), 1 ♀ (ZMUC), 1 ♀ (MNHN JC 314), 1 ♂ (SEM); same locality, Shu Jia Yan Dong Cave, 16. XI. 2001, leg. L. Latella & Berzacola, 1 ♀ (MNHN JC 314); same locality, Da Kong Dong Cave, 12. XI. 2003, leg. L. Latella, 1 ♀ (MNHN JC 314); same locality, Luo Sai Dong Cave, 19. XI. 2001, leg. D. Avesani, L. Latella & S. Meggiorini, 1 ♀ (MNHN JC 314).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77BA647FF62FA06FCA0759B.taxon	etymology	Name. To emphasize the deeply incised caudolateral lobulations on most metaterga.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77BA647FF62FA06FCA0759B.taxon	diagnosis	Diagnosis. Differs from all congeners by the unusually deeply incised caudolateral and lateral lobulations on most of the metaterga, as well as the peculiar, biramous, multituberculate, midway, lateral process (dp) of the gonopod telopodite.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77BA647FF62FA06FCA0759B.taxon	description	Description. Length of adults of both sexes ca 7 - 8 mm, width 1.1 - 1.2 mm, body broadest at segments 3 and 4. Holotype ca 7 mm long and 1.2 mm wide. Coloration rather uniformly whitish to light yellow, anterior body parts often a little infuscate, light yellow-brownish. Adults with 20 segments (Fig. 22 A), conglobation pattern typical of “ Doratodesmidae ”, complete. Head (Fig. 23 B) basically as in preceding species, but tegument slightly rougher (Figs 24 A, B, D-F), antennae somewhat shorter (Fig. 23 B), collum not covering head from above, regularly convex, with four rows of high, conical tubercles (Figs 22 B, E). Metaterga behind collum with three transverse, rather regular rows of similarly high tubercles, most of which still retain very short, bisegmented setae (Fig. 22; 24 E, F); limbus microcrenulate. Paraterga mostly directed ventrolaterad, strongly declivous, narrow, tips clearly surpassing the level of venter, unusually distinctly crenulate / lobulate laterally and caudolaterally, usually with characteristic, deep incisions between lobules (Figs 22; 23 E, F); paraterga 2 strongly enlarged, subvertical, anterolateral and lateral margins both evidently trilobate, caudal margin above schism with 4 - 5 similarly evident lobules, both schism and hyposchism small; paraterga 3 and 4 slightly shorter than others, bilobate laterally (Fig. 22 B), following paraterga broadly rounded and mostly evidently trilobate, overlap typical. Pore formula normal, ozopores located near base of ventrocaudal lobulation (Fig. 23 F). Pleurotergal carinae wanting. Epiproct strongly flattened, dorsally also tuberculate, with several deep incisions at lateral edge, directed ventrocaudad, with the usual four cones just below tip; para- and hypoprocts as in Figs 23 A, D. Sterna usually with a rather deep, narrow depression between coxae (Fig. 24 C), only those between coxae 6, 7 and 9 much wider (Fig. 25 A). Gonopod aperture transverse-oval, relatively small, far from reaching lateral sides of segment 7. Legs rather short; femoral and tarsal segments equal and longest; several basal segments microtuberculate; claw simple, slightly curved ventrad; some setae very sparsely microdenticulate (Fig. 24 C). Gonopods (Figs 25 A, B) relatively complex. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally, with a conspicuous triangular lobe frontolaterally. Telopodite considerably longer than coxite, unciform, rather slender, setose in its basal half, with a conspicuous, biramous, microtuberculate, distofemoral process (dp) laterally at about midway, more distally with a slightly folded acropodite bearing several small outgrowths, but devoid of a hairpad; seminal groove terminating subapically.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A77BA647FF62FA06FCA0759B.taxon	discussion	Remarks. Glyphiulus latellai Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2007 is another millipede recorded from Tiao Shuz Dong and Shu Jia Yan caves (Golovatch et al. 2007 a), whereas Liao Jing Ling Dong Cave contains the subcosmopolitan Oxidus gracilis (C. L. Koch, 1847) (Geoffroy and Golovatch 2004).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A745A640FF62FD99FD6B7562.taxon	description	Figs 27 - 30.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A745A640FF62FD99FD6B7562.taxon	materials_examined	Material. China, Yunnan Prov., Mengzi County, pothole no. 2 (Ma Fa Tiao Dong), 6. I. 1989, leg. P. Beron, 2 ♁♁, 1 ♀ (MNHN JC 315), 1 ♁, 3 ♀♀, 9 juv. (NMNHS), 1 ♁ (ZMUM), 1 ♀ (SEM); Yunnan Prov., Mengzi County, Longbaopo Dong Cave, 27. XI. 1995, leg. I. Kos, B. Sket & F. Velkovrh, 1 ♁ & 1 ♀ in copula (OBBFUL), 1 ♁ (ZMUC), 1 ♁ (SEM).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A745A640FF62FD99FD6B7562.taxon	diagnosis	Diagnosis. Differs from congeners by the simultaneous lack of mid-dorsal outgrowths on metaterga and of lateral lobulations on paraterga 2, the presence of deep crenulations at the caudal edge of most metaterga, coupled with a peculiar shape of the gonopod telopodite supplied with a large, multidenticulate, lateral lobe and a long, slightly curved, rather simple acropodite. Short redescription. Length of adults of both sexes ca 10 - 11 mm, width 1.8 - 2.1 mm, body broadest at segments 3 and 4. Coloration rather uniformly whitish to light yellow, anterior body parts often a little infuscate, light yellowish-brown. Adults with 20 segments, conglobation pattern typical of “ Doratodesmidae ”, complete (Golovatch 2003). Superficially, strongly resembling Eutrichodesmus incisus sp. n. All characters fully agreeing with the original description (Zhang and Wang 1993), but augmented here by illustrations showing more details: head (Fig. 28 C), antennae (Fig. 28 C, D), collum (Fig. 27 D), meta- and paraterga (Figs 27 A-C, E, F; 28 B), tegument and tergal setae (Figs 28 F; 29 A, B), ozopores (Fig. 28 E), telson (Fig. 27 F; 28 A), legs (Figs 29 C, D; 30 A). Sterna usually with a rather deep, narrow depression between coxae (Fig. 28 B), only those between coxae 6, 7 and 9 much wider (Fig. 30 A). Gonopod aperture transverse-oval, relatively small, far from reaching lateral sides of segment 7. Legs rather long; femoral and tarsal segments equal and longest; several basal segments microtuberculate; claw simple, slightly curved ventrad; some setae sparsely microdentate (Figs 29 C, D; 30 A). Gonopods (Figs 30 B, C) relatively complex. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally. Telopodite considerably longer than coxite, subunciform, basal half slender and setose, at about midway conspicuously enlarged, with a multidenticulate lateral lobe (lo) and a very short solenomere (sl) with a hairpad on top; acropodite relatively long and simple.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A745A640FF62FD99FD6B7562.taxon	discussion	Remarks. This species has hitherto been described or reported from three caves in Yunnan, China (Wang and Zhang 1993; Golovatch 2003). One of the new samples (from Longbaopo Cave) is strictly topotypic, while Ma Fa Tiao Dong Cave is known to support another millipede, Glyphiulus subgranulatus Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2007 (see Golovatch et al. 2007 a).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74FA64EFF62FB89FC0F75F6.taxon	type_taxon	Type genus: Haplodesmus Cook, 1895 (replacement name for the preoccupied Haplosoma Verhoeff, 1894).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74FA64EFF62FB89FC0F75F6.taxon	diagnosis	Diagnosis. A family of Polydesmoidea with gonopod aperture relatively modest in size, transverse-ovoid; gonocoxae subquadrate, non-globose, enlarged laterally, often densely setose; telopodites usually rather simple, uni-, bi- or triramous, usually with evidence of torsion near end of femorite, rather stout (about as high as coxae) to very slender, always held parallel to main axis, not crossing each other even distally, but hinging into a considerable hollow formed by ventromedian parts of coxae; seminal groove and coxal cannula usually present, solenomere only seldom being a separate branch (occasionally long and flagelliform), usually taking up much to most of telopodite distad of a more or less evident, lateral process, lobe or outgrowth near end of femorite, latter enlarged or slender; acropodite varying from very conspicuous to nearly absent. Body shape ranging from vermiform / subcylindrical or “ pyrgodesmid-like ” (see Key below) (neither capable of volvation) to “ doratodesmid ” (capable of volvation). Paraterga, if present, always declivous, hence the dorsum is invariably very convex. Metaterga clothed with a cerotegumental crust, usually with 2 - 4 transverse rows of tubercles / bosses, sometimes with mid-dorsal projections. Pore formula usually normal, porosteles sometimes present. Tergal pubescence often with bisegmented setae. Walking leg coxae mostly contiguous medially, due to very narrow sterna. Head usually slightly transverse, only rarely elongated; antennae usually stout, antennomere 6 usually longest.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74FA64EFF62FB89FC0F75F6.taxon	discussion	Only six recognizable genera are contained in this small family.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74CA64EFF62FDB6FE1F7078.taxon	type_taxon	Type species: Prosopodesmus jacobsoni Silvestri, 1910, by original designation (type locality: Batavia (= Jakarta), Java, Indonesia).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74CA64EFF62FDB6FE1F7078.taxon	diagnosis	Diagnosis. Body with 20 segments, pyrgodesmid-like, not capable of volvation, with rather well developed and declivous paraterga. Metaterga with three transverse rows of bosses / tubercles. Gonopods strongly resembling those of typical Polydesmidae, being rather simple, unciform; coxae with only a few ventrolateral setae; telopodites sometimes with a lateral outgrowth at midway, always devoid of a separate solenomere branch; seminal groove terminating distally or apically, hairpad often present.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74CA64FFF62FB3CFC4676FD.taxon	type_taxon	Type species: Cylindrodesmus hirsutus Pocock, 1889, by monotypy (type locality: Christmas Island, Australia).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74CA64FFF62FB3CFC4676FD.taxon	diagnosis	Diagnosis. Body with 19 (♂) or 20 (♀) segments, subcylindrical, not capable of volvation, with paraterga 2 rather well developed, but following ones mostly represented by lateral swellings. Collum and metaterga with abundant setation in part mixed with long, bisegmented, tactile setae. Gonopods especially simple; coxae with only a few ventrolateral setae; telopodite (= solenomere) sometimes with a lateral outgrowth at midway; seminal groove terminating subapically and devoid of a hairpad.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74DA648FF62FA06FBB37524.taxon	type_taxon	Type species: Atopogonus baccatus Carl, 1926, by monotypy (type locality: New Caledonia, France).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74DA648FF62FA06FBB37524.taxon	diagnosis	Diagnosis. Body with 20 segments, subcylindrical, not capable of volvation, with paraterga 2 rather well developed, but subsequent ones represented by lateral swellings at most. Collum small; collum and following paraterga microtuberculate or granular. Gonopods with small virtually bare and reduced gonocoxae and peculiar, strongly geniculate telopodite, coupled with no traces of either a coxal cannula or a seminal groove.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74DA64FFF62FCBFFD727166.taxon	type_taxon	Type species: Helodesmus porosus Cook, 1896, by monotypy (type locality: Java, Indonesia).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74DA64FFF62FCBFFD727166.taxon	diagnosis	Diagnosis. Body with 19 segments, subcylindrical, not capable of volvation, with paraterga 2 rather well developed, but subsequent ones mostly represented by lateral swellings. Collum still large, covering the head from above, with 4 - 5 transverse rows of setigerous tubercles or pits. Following paraterga with three rows of similar tubercles or pits. Pore formula: 5, 7 - 17 (18). Gonopods with poorly setose gonocoxae and a considerably shortened prefemoral + femoral (= setose) part, deeply biramous thereafter, with a long and flagelliform solenomere devoid of a hairpad.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74AA648FF62FEC9FB957078.taxon	type_taxon	Type species: Doratonotus armatus Pocock, 1894. Proposed to replace the preoccupied Doratonotus Pocock, 1894 (Cook in Cook & Collins 1895) (type locality: Java, Indonesia).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74AA648FF62FEC9FB957078.taxon	diagnosis	Diagnosis. Body “ doratodesmid ”, with or without mid-dorsal projections; conglobation complete. Gonopod coxae usually abundantly setose ventrolaterally; telopodite usually stout and strongly enlarged laterally towards end of femorite, with or without a short solenomere branch thereafter; acropodite variable, from absent to well-developed.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74AA649FF62FB3CFBDA75F6.taxon	type_taxon	Type species: Eutrichodesmus demangei Silvestri, 1910, by original designation (type locality: Phu-Ly, Vietnam).	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
1D3687F8A74AA649FF62FB3CFBDA75F6.taxon	diagnosis	Diagnosis. Body “ doratodesmid ”, with or without mid-dorsal projections; conglobation usually complete. Gonopod coxae usually abundantly setose ventrolaterally; telopodite usually slender, not enlarged towards end of femorite, but with a more or less distinct process or outgrowth laterally, opposite recurvature point of seminal groove; solenomere thereafter taking up most of telopodite, sometimes elaborate; seminal groove terminating distally to subapically, with or without a hairpad; acropodite small to nearly absent.	en	Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, VandenSpiegel, Didier (2009): Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7 (7): 1-53, DOI: 10.3897/zookeys.7.117
