taxonID	type	description	language	source
8545C70565AD5BE6AB0C91E0A73FC442.taxon	description	Figs 2, 3 鹅凰嶂耳草 (e Huang Zhang er Căo)	en	Xu, Yi-Da, Zhang, Ying, Wang, Rui-Jiang (2023): Hedyotis longiramulis (Rubiaceae), a new species from south China. PhytoKeys 230: 271-287, DOI: http://dx.doi.org/10.3897/phytokeys.230.87675, URL: http://dx.doi.org/10.3897/phytokeys.230.87675
8545C70565AD5BE6AB0C91E0A73FC442.taxon	materials_examined	Type. China. Guangdong Province: Yangchun City, Bajia Town, Guangdong Ehuangzhang Provincial Nature Reserve, roadsides, 21 ° 52 ' N, 111 ° 25 ' E, elev. 643 m. April 9, 2021, Y. D. Xu & R. J. Wang 6540 (holotype: IBSC [IBSC 0865777!]; isotype: IBSC [IBSC 0865778!]).	en	Xu, Yi-Da, Zhang, Ying, Wang, Rui-Jiang (2023): Hedyotis longiramulis (Rubiaceae), a new species from south China. PhytoKeys 230: 271-287, DOI: http://dx.doi.org/10.3897/phytokeys.230.87675, URL: http://dx.doi.org/10.3897/phytokeys.230.87675
8545C70565AD5BE6AB0C91E0A73FC442.taxon	diagnosis	Diagnosis. The species is similar to H. caudatifolia in having a subshrubby habit, ovate to lanceolate leaves, and long lateral branches with several terminal and axillary inflorescences, but differs from it by having puberulent, more or less tetragonal and decussately sulcate juvenile stems (versus glabrous and terete in H. caudatifolia), waxy leaf surface (versus non-waxy in H. caudatifolia), shorter peduncles (0.5 - 2.0 cm versus 2.0 - 10.0 cm in H. caudatifolia), a higher length ratio of corolla lobe to tube (0.9 - 1.0 versus approximately 0.8 in H. caudatifolia), and subglobose capsules (versus ellipsoid-oblong or ellipsoid in H. caudatifolia).	en	Xu, Yi-Da, Zhang, Ying, Wang, Rui-Jiang (2023): Hedyotis longiramulis (Rubiaceae), a new species from south China. PhytoKeys 230: 271-287, DOI: http://dx.doi.org/10.3897/phytokeys.230.87675, URL: http://dx.doi.org/10.3897/phytokeys.230.87675
8545C70565AD5BE6AB0C91E0A73FC442.taxon	description	Description. Perennial woody subshrubs, 40 - 120 cm tall. Stem more or less tetragonal and decussately sulcate at juvenile internodes, becoming terete with age, purplish, puberulent, branched at upper part. Leaves opposite, 5 - 16 x 1.5 - 4 cm on main stem and 1.0 - 6.5 x 0.3 - 1.5 cm on lateral branches, ovate to lanceolate, coriaceous, dark green adaxially, greyish-green or sometimes purplish abaxially, both surfaces waxy, apex acute or subacute, base cuneate or shortly decurrent; petiole 5 - 15 mm long on main stem and 2 - 5 mm long on lateral branches, waxy or puberulent; midrib depressed adaxially and prominent abaxially, secondary veins usually 5 - 6 on each side, sometimes indistinct adaxially; stipules 4 - 10 x 3 - 6 mm, triangular, apex acute to acuminate, margin sparsely glandular serrate, puberulent abaxially. Inflorescences growing on long lateral branches, terminal and axillary in the upper nodes, 1.5 - 3.5 cm long, cymose or paniculate-cymose; inflorescence axes tetragonal, sulcate; peduncles 0.5 - 2.0 cm long; bracts ca. 1 mm long, subulate. Flowers heterostylous, pedicels 0.9 - 2.0 mm long. Hypanthium ca. 1 mm long, obconic to subglobose; lobes 4, ca. 0.9 x 0.9 mm, ovate-triangular, blunt or rounded at apex. Corolla white or purplish, tube 3.5 - 3.8 mm long, glabrous abaxially and densely or sparsely pubescent adaxially; lobes 4, 3.5 - 3.8 x 1.8 - 2.2 mm, ovate-triangular; stamens 4, anthers ca. 0.9 mm long; stigma bilobed, ca. 0.5 mm long, subglobose, papillate. Long-styled flowers: stamens included, filaments ca. 0.6 mm long, adnate to the middle part of corolla tube; style ca. 7.6 mm long, exserted, glabrous. Short-styled flowers: stamens exserted, filaments ca. 2.8 mm long, adnate to the throat of corolla tube; style ca. 2.7 mm long, included, glabrous. Fruits capsular, ca. 2.0 mm in diameter, subglobose, glabrous, dehiscent diplophragmously; seeds several, ca. 1 mm long, cymbiform, with reticulate surface. (Fig. 3 A-C.)	en	Xu, Yi-Da, Zhang, Ying, Wang, Rui-Jiang (2023): Hedyotis longiramulis (Rubiaceae), a new species from south China. PhytoKeys 230: 271-287, DOI: http://dx.doi.org/10.3897/phytokeys.230.87675, URL: http://dx.doi.org/10.3897/phytokeys.230.87675
8545C70565AD5BE6AB0C91E0A73FC442.taxon	distribution	Distribution and habitat. Hedyotis longiramulis is only known from Yangchun City of Guangdong Province, China. It grows mainly in damp places under broad-leaved forests, sometimes on roadsides at the elevation of 500 - 700 m. The associated species are mainly (Hance ex Benth.) Krass. (Melastomataceae), Melastoma sanguineum Sims (Melastomataceae), Dunnia sinensis Tutcher (Rubiaceae), Dicranopteris ampla Ching & P. S. Chiu (Gleicheniaceae) and Selaginella doederleinii Hieron. (Selaginellaceae).	en	Xu, Yi-Da, Zhang, Ying, Wang, Rui-Jiang (2023): Hedyotis longiramulis (Rubiaceae), a new species from south China. PhytoKeys 230: 271-287, DOI: http://dx.doi.org/10.3897/phytokeys.230.87675, URL: http://dx.doi.org/10.3897/phytokeys.230.87675
8545C70565AD5BE6AB0C91E0A73FC442.taxon	etymology	Etymology. The specific epithet " longiramulis " of the new species refers to its long lateral branches bearing many inflorescences.	en	Xu, Yi-Da, Zhang, Ying, Wang, Rui-Jiang (2023): Hedyotis longiramulis (Rubiaceae), a new species from south China. PhytoKeys 230: 271-287, DOI: http://dx.doi.org/10.3897/phytokeys.230.87675, URL: http://dx.doi.org/10.3897/phytokeys.230.87675
8545C70565AD5BE6AB0C91E0A73FC442.taxon	materials_examined	Additional specimens examined (paratypes). China. Guangdong Province: Yangchun City, Guigang Town, Baichong Provincial Nature Reserve, roadside, 13 Sept. 1990, Nian Liu et al. 424 (IBSC); ibid., 18 May 1991, Nian Liu et al. 1735 (IBSC). Yangchun City, Bajia Town, Guangdong Ehuangzhang Provincial Nature Reserve, mountain land and valley, 24 Oct. 1957, Kui Liang 69692 (CANT); ibid., 23 Oct. 1957, Bao-Han Liang 89654 (CANT); ibid., 11 Oct. 1990, Nian Liu et al. 866 and 899 (IBSC); ibid., 11 May 2001, Hua-Gu Ye et al. 5629 (IBSC); ibid., 7 Apr. 2019, Xin-Xin Zhou et al. ZXX 0026 (IBSC); ibid., 12 Aug. 2020, Dan Liang et al. WP 1366 (IBSC); ibid., 10 Sept. 2020, Yi-Da Xu & Fan Su AP 0138 (IBSC); ibid., 9 Apr. 2021, Rui-Jiang Wang & Yi-Da Xu 6541 (IBSC).	en	Xu, Yi-Da, Zhang, Ying, Wang, Rui-Jiang (2023): Hedyotis longiramulis (Rubiaceae), a new species from south China. PhytoKeys 230: 271-287, DOI: http://dx.doi.org/10.3897/phytokeys.230.87675, URL: http://dx.doi.org/10.3897/phytokeys.230.87675
