identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
1C598301283DFF91FC885B94FF31FC13.text	1C598301283DFF91FC885B94FF31FC13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryphalini Classification	<div><p>History of Cryphalini Classification</p> <p>The first mention of Cryphalini as a group name was by Lindemann in 1877. The group-concept was much broader, including Pityophthorus Eichhoff, 1864, Hypoborus Erichson, 1836, Hylocurus Eichhoff, 1872, and Xyloctonus Eichhoff, 1872, all of which are currently placed in separate tribes.</p> <p>Eichhoff described many of the contemporary genera, plus several which are now junior synonyms (Eichhoff 1878a). Most contemporary Cryphalini genera were classified in ‘Cryphalidae’ which was a much narrower concept than previously described, containing the known Cryphalini genera (except Eidophelus Eichhoff, 1876) plus Hypoborus and Liparthrum Wollaston, 1854 (Hypoborini) and what is now the genus Pityoborus Blackman, 1922 (Corthylini). Other groups, such as contemporary Micracidini LeConte, 1876 and Xyloctonini Eichhoff, 1878, were explicitly described as different tribes.</p> <p>This approximate group concept of Cryphalini remained until Hopkins presented a major reclassification of Scolytinae in 1915, with a group ‘Cryphalinae’ in an even broader sense including the contemporary tribes Xyleborini and some genera in the current Dryocoetini Lindemann, 1877 and Corthylini LeConte, 1876. Most of the contemporary Cryphalini are contained in Hopkins ‘Division I, Subdivision A’, a group described based on the type of vestiture (‘with scales, rarely with hairs’) and the pattern of asperities along the anterior margin of the pronotum. The characters used to describe and diagnose the genera within the Cryphalini include the presence of distinct elytral strial punctures, the shape of the lateral margins of the pronotum, the degree to which the pronotum was produced, the apex of the elytra, and most importantly, the morphology of the antennae. Eidophelus was considered in a different subdivision with Dendroterus Blandford, 1904 and some other genera now placed in Dryocoetini.</p> <p>Balachowsky (1949) presented a clarified classification of bark beetles which reflects most contemporary tribes. While restricted to the fauna of France, it clearly described Cryphalini as different from Hypoborini. Wood (1954) evaluated the North American Cryphalini genera and species, representing the contemporary scope of the tribe for the region. Two additional genera from East Asia were described by Nobuchi (1981), inferred partly by new sets of characters from the proventriculus (Nobuchi 1969). While these alone enabled identification of most scolytine genera, they were not widely adopted as a diagnostic set of characters.</p> <p>Schedl was a major contributor to Cryphalini systematics. While very prolific in describing new species, his classification of Cryphalini did not settle any of the considerable confusion and his nomenclatural practices were not consistently valid (see Wood and Bright 1992). Schedl described five genera in the contemporary Cryphalini, but four of these are now regarded as junior synonyms.</p> <p>The most recent treatment of Cryphalini was Wood’s (1986a) review of all scolytine tribes and genera. In this review, Cryphalini was diagnosed based on the combination of the ascending costal margin of the elytra, by the base of the elytra that is unarmed (or rarely armed with a continuous line, not crenulations), by the declivous (steeply sloped) pronotum usually armed with serrations and asperities, by the concealed head when viewed dorsally, by the eye that is ‘usually’ entire and ‘less commonly emarginated’, by the antennal funicle with no more than five segments, by the antennal club which is ‘strongly flattened’ and never truncate, by the lateral margins of the pro- and mesotibiae armed with denticles, and by the metanepisternum which has a posterior margin concealed by the elytron.</p> <p>Wood and Bright brought significant order to the chaos of the classification of Scolytinae as a whole, including Cryphalini. Many genera were synonymized, and a small number was described. Their compilation of the catalogs (Wood and Bright 1987; Wood and Bright, 1992) is a milestone in scolytine systematics and provides a very useful starting point for taxonomic reviews. Allothenemus Bright and Torres, 2006 was subsequently described. During the final stages of this manuscript preparation, four further genera were described in Cryphalini: Atomothenemus Bright, 2019, Microsomus Bright, 2019, Pygmaeoborus Bright, 2019, and Trypolepis Bright, 2019.</p> </div>	https://treatment.plazi.org/id/1C598301283DFF91FC885B94FF31FC13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012835FF9EFC885F65FADAF8E1.text	1C5983012835FF9EFC885F65FADAF8E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Johnson Johnson & Hulcr & Knížek & Atkinson & Mandelshtam & Smith & Cognato & Park & Li & Jordal 2020	<div><p>Coriacephilini Johnson trib. nov.</p> <p>Type of tribe</p> <p>Coriacephilus Schedl, 1939.</p> <p>Diagnosis</p> <p>Tarsi with third segment cylindrical. Eye deeply emarginated. Antennal funicle with 3–5 segments. Antennal club with sutures and a complete septum. Hypomeron with bifurcating setae.</p></div> 	https://treatment.plazi.org/id/1C5983012835FF9EFC885F65FADAF8E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012832FF98FC885E2AFD6CF93B.text	1C5983012832FF98FC885E2AFD6CF93B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coriacephilus Schedl 1939	<div><p>Coriacephilus Schedl, 1939b: 339 (Fig. 9)</p> <p>Type of genus</p> <p>Stephanoderes coriaceus Eichhoff, 1878. Diagnosis</p> <p>Coriacephilus can be separated from other Cryphalini by the combination of cylindrical third tarsal segments, deeply emarginated eye, antennae with sutures and septum, and bifurcating setae on the hypomeron.</p> </div>	https://treatment.plazi.org/id/1C5983012832FF98FC885E2AFD6CF93B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012830FF9BFC885FE2FA7FF8AA.text	1C5983012830FF9BFC885FE2FA7FF8AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coriacephilus coriaceus (Eichhoff 1878) Stephanoderes	<div><p>Coriacephilus coriaceus (Eichhoff, 1878b: 494) (Stephanoderes).</p> <p>= Cryphalus birmanus Eggers, 1925: 153 (syn: Schedl, 1940c).</p> </div>	https://treatment.plazi.org/id/1C5983012830FF9BFC885FE2FA7FF8AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012831FF9AFF285C04FD1FF953.text	1C5983012831FF9AFF285C04FD1FF953.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryphalini Lindemann 1877	<div><p>Cryphalini Lindemann, 1877</p> <p>=Cryphaloideae Lindemann, 1877: 165.</p> <p>=Cryphalidae Eichhoff, 1878b: 107.</p> <p>= Cryphalini Weise, 1883: 181.</p> <p>=Cryphali Blandford, 1898: 185.</p> <p>= Cryphalinae Trédl, 1907: 40.</p> <p>=Cryphalina Balachowsky, 1949: 200.</p> <p>Type of tribe</p> <p>Cryphalus Erichson, 1836.</p> <p>Diagnosis</p> <p>Tarsi with third segment bilobed. Eye emarginated. Antennal funicle with 3–5 segments. Antennal club with sutures and no septum. Hypomeron with bifurcating setae. Postnotum not fused with metanotum. Aedeagus with long penis apodemes, sometimes fused at apex, with a complete tegmen. Tegmen with pair of apodemes.</p></div> 	https://treatment.plazi.org/id/1C5983012831FF9AFF285C04FD1FF953	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012831FF9AFC885888FC62FB1F.text	1C5983012831FF9AFC885888FC62FB1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryphalus Erichson 1836	<div><p>Cryphalus Erichson, 1836: 61 (Figs. 10–12)</p> <p>Synonymy</p> <p>= Pseudocryphalus Ferrari, 1869: 252.</p> <p>= Taenioglyptes Bedel, 1888: 398.</p> <p>= Cryptarthrum Blandford, 1896a: 200.</p> <p>= Allarthrum Hagedorn, 1912c: 355.</p> <p>= Dacryphalus Hopkins, 1915: 42 syn. nov.</p> <p>= Ericryphalus Hopkins, 1915: 38.</p> <p>= Hypocryphalus Hopkins, 1915: 41 syn. nov.</p> <p>= Margadillius Hopkins, 1915: 37 syn. nov.</p> <p>= Piperius Hopkins, 1915: 39.</p> <p>= Toenioglyptes Balachowsky, 1949: 205 (unavailable name). = Ernocryphalus Murayama, 1958: 934.</p> <p>= Acryphalus Tsai and Li, 1963: 604 (unavailable name). = Gugocryphelus Tsai and Li, 1963: 603 (unavailable name). = Jugocryphalus Tsai and Li, 1963: 604.</p> <p>Type of genus</p> <p>Bostrichus asperatus Gyllenhal, 1813.</p> <p>Diagnosis</p> <p>Cryphalus can be distinguished from similar genera by the combination of emarginated eyes, sutures on the antennae clearly visible but without a septum, by the weakly bilobed third tarsal segments, the proventriculus with a large apical plate, and the aedeagus that has a tegmen sclerotized completing a ring, usually with two tegminal apodemes.</p> </div>	https://treatment.plazi.org/id/1C5983012831FF9AFC885888FC62FB1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301282AFF83FF285994FCF4FF18.text	1C598301282AFF83FF285994FCF4FF18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryphalus araucariae Schedl 1970	<div><p>Cryphalus araucariae Schedl, 1970a: 214.</p> <p>Cryphalus armatus Schedl, 1974: 459.</p> <p>Cryphalus artestriatus (Browne, 1970: 553) (Taenioglyptes).</p> <p>Cryphalus artocarpus (Schedl, 1939b: 432) (Ericryphalus).</p> <p>= Cryphalus artocarpus Schedl, 1958d: 498 (syn: Schedl, 1975d). = Cryphalus brownei Wood, 1992b: 79 (unnecessary replacement name).</p> <p>Cryphalus asper (Broun, 1881: 742) (Tomicus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus asperatus (Gyllenhal, 1813: 368) (Bostrichus).</p> <p>= Bostrichus abietis Ratzeburg, 1837: 163 (syn: Wood, 1972b).</p> <p>Cryphalus asperulus Schedl, 1948b: 26.</p> <p>Cryphalus ater Browne, 1984d: 91.</p> <p>Cryphalus babai Murayama, 1961: 29.</p> <p>Cryphalus bakeri (Eggers, 1927c: 77) (Stephanoderes) comb. nov. [Hypocryphalus].</p> <p>Cryphalus balanopselaphus Eggers, 1920: 21.</p> <p>Cryphalus basihirtus Beeson, 1929: 227 comb. nov. [Hypocryphalus]. Cryphalus bellus Schedl, 1957b: 47.</p> <p>Cryphalus bicarinatus (Nobuchi, 1975: 53) (Taenioglyptes).</p> <p>Cryphalus bicolor (Browne, 1983b: 69) (Cryphalomorphus).</p> <p>Cryphalus bidentatus (Browne, 1980b: 383) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus boettcheri Schedl, 1943: 38.</p> <p>Cryphalus brasiliensis Schedl, 1976: 65.</p> <p>Cryphalus brevior (Schedl, 1943: 40) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus brevipilosus Schedl, 1942c: 173.</p> <p>Cryphalus brimblecombei Schedl, 1948b: 26.</p> <p>Cryphalus brunneus Browne, 1984d: 93.</p> <p>Cryphalus buloloensis Browne, 1984c: 67.</p> <p>Cryphalus capucinicollis Schedl, 1950b: 47.</p> <p>Cryphalus capucinoides Eggers, 1939b: 4.</p> <p>Cryphalus capucinomorphus Schedl, 1950b: 48.</p> <p>Cryphalus capucinus Schedl, 1938c: 425.</p> <p>Cryphalus carinatus (Browne, 1980b: 382) (Margadillius) comb. nov. [Margadillius].</p> <p>Cryphalus carpini Berger, 1917: 234.</p> <p>Cryphalus carpinivorus Murayama, 1930: 14.</p> <p>Cryphalus chamaecipariae Niisima, 1910: 10.</p> <p>Cryphalus chinlingensis Tsai and Li, 1963: 612.</p> <p>Cryphalus ciliatipes Blandford, 1896b: 242.</p> <p>Cryphalus cinereotestaceus (Motschulsky, 1866: 403) (Hypoborus). Cryphalus compactus Lea, 1910: 139.</p> <p>Cryphalus confusus (Hopkins, 1915: 38) (Margadillius) comb. nov. [Margadillius].</p> <p>Cryphalus constrictus Schedl, 1942c: 174.</p> <p>= Cryphalus perminimus Schedl, 1942d: 13 (syn: Schedl, 1958e). = Hypocryphalus constrictus Schedl, 1942d: 22.</p> <p>= Hypocryphalus froggatti Nunberg, 1961: 611 (syn: Schedl, 1962c).</p> <p>Cryphalus corpulentus (Schedl, 1942d: 22) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus coryli Stark, 1936: 144.</p> <p>Cryphalus cryptomeriae Niisima, 1908: 91.</p> <p>Cryphalus cylindricus Browne, 1980b: 384.</p> <p>Cryphalus cylindripennis (Schedl, 1959a: 483) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus cylindrus (Browne, 1950: 647) (Dacryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus densepilosus (Schedl, 1942d: 21) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus dilutus Eichhoff, 1878a: 384 comb. res. [Hypocryphalus]. = Cryphalus dilutus Eichhoff, 1878b: 490 (syn: Johnson et al., 2017).</p> <p>Cryphalus dipterocarpi Wood, 1989: 179.</p> <p>Cryphalus discrepans (Schedl, 1965a: 58) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalusdiscretus Eichhoff,1878a: 385 comb. res. [Hypocryphalus]. = Cryphalus discretus Eichhoff, 1878b: 490 (syn: Johnson et al., 2017).</p> <p>= Cryphalus scabricollis Eichhoff, 1878b: 491 (syn: Wood, 1989). = Cryphalus brevisetosus Schedl, 1943: 36 (syn: Wood, 1989).</p> <p>Cryphalus dissimilis (Nobuchi, 1975: 57) (Taenioglyptes).</p> <p>Cryphalus diversicolor Browne, 1984d: 90.</p> <p>Cryphalus dorsalis (Motschulsky, 1866: 403) (Hypoborus).</p> <p>= Hylesinus sericeus Motschulsky, 1866: 402 (syn: Wood, 1969). = Hypoborus nebulosus Motschulsky, 1866: 403 (syn: Wood, 1969).</p> <p>= Cryphalus indicus Eichhoff, 1878a: 384 syn. nov.</p> <p>= Cryphalus indicus Eichhoff, 1878b: 489.</p> <p>Remarks: No formal synonymy has been proposed for this species name. Wood (1969) indicated that they are probably the same, and Wood and Bright (1992) listed the synonyms as above, choosing Cryphalus dorsalis as the valid name with dorsalis getting priority chosen by the first reviser. The nomenclature is being treated as valid, but the subjective synonymy is uncertain (see Mandelshtam and Nikitsky 2005).</p> <p>Cryphalus dubiosus Schedl, 1963c: 66.</p> <p>Cryphalus duplosquamosus Schedl, 1942d: 15.</p> <p>Cryphalus eggersi Johnson nom. nov.</p> <p>= Cryphalus confusus Eggers, 1927b: 395 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Cryphalus confusus Eggers, 1927b, because the combination is already occupied following the transfer of Margadillius confusus Hopkins, 1915 to Cryphalus. The replacement name is a noun in the genitive case, honoring the original author.</p> <p>Cryphalus elaboratus Schedl, 1950b: 51.</p> <p>Cryphalus elongatus Schedl, 1962f: 105.</p> <p>Cryphalus eriobotryae Johnson, 2019: 3.</p> <p>Cryphalus erraticus Schedl, 1979c: 96.</p> <p>Remarks: The original and subsequent spelling is erraticum as an adjective, which should be the masculine erraticus.</p> <p>Cryphalus erythrinae (Hopkins, 1915: 38) (Margadillius) comb. nov. [Margadillius].</p> <p>Cryphalus exiguus Blandford, 1894b: 82.</p> <p>Cryphalus felis Wood, 1989: 180.</p> <p>Cryphalus fici (Browne, 1986: 90) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus ficivorus Murayama, 1958: 933.</p> <p>Cryphalus formosanus Schedl, 1942a: 175.</p> <p>Cryphalus fugax Schedl, 1973b: 87.</p> <p>Cryphalus fuliginosus Blandford, 1895: 319.</p> <p>Cryphalus fulmineus Wood, 1989: 180.</p> <p>Cryphalus fulvus Niisima, 1908: 92.</p> <p>= Cryphalus pini Eggers, 1921: 39 (syn: Eggers, 1923b).</p> <p>Cryphalus furukawai Murayama, 1934a: 59.</p> <p>Cryphalus fuscus Johnson nom. nov.</p> <p>= Cryphalus cylindrus Browne, 1984d: 92 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Cryphalus cylindrus Browne, 1984d following the transfer of Dacryphalus cylindrus Browne, 1950 from Hypocryphalus. The replacement name is an adjective, meaning either brown or indistinct.</p> <p>Cryphalus garambaensis Nunberg, 1965: 25.</p> <p>Cryphalus garciniae Nobuchi, 1959: 24.</p> <p>Cryphalus giganteus Schedl, 1950a: 22.</p> <p>Cryphalus gigas Schedl, 1975b: 218.</p> <p>Cryphalus glabratus (Schedl, 1959a: 484) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus gracilis Johnson nom. nov.</p> <p>= Cryphalus laevis Browne, 1984b: 288 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Cryphalus laevis Browne, 1984b because the name is occupied by Hypocryphalus laevis Browne, 1980a. The replacement name is an adjective meaning slender or graceful.</p> <p>Cryphalus granulatus (Schedl, 1942a: 176) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus helopioides Schedl, 1953b: 295.</p> <p>Cryphalus hirsutus (Nobuchi, 1975: 52) (Taenioglyptes).</p> <p>Cryphalus horridus Eichhoff, 1878a: 384.</p> <p>= Cryphalus horridus Eichhoff, 1878b: 488 syn. nov.</p> <p>Cryphalus imitans (Schedl, 1951d: 148) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus infimus Schedl, 1972c: 287.</p> <p>Cryphalus intermedius Ferrari, 1867: 79.</p> <p>Cryphalus interponens (Schedl, 1953a: 126) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus jeholensis Murayama, 1939: 143.</p> <p>Cryphalus jezoensis Inouye and Nobuchi, 1957: 49.</p> <p>Cryphalus juglansi Niisima, 1913: 3.</p> <p>Cryphalus kagoshimensis (Nobuchi, 1975: 56) (Taenioglyptes).</p> <p>Cryphalus kalambanganus (Schedl, 1943: 39) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus kesiyae Browne, 1975: 288.</p> <p>Cryphalus kivuensis Schedl, 1957b: 48.</p> <p>Cryphalus kolbei (Hagedorn, 1912c: 355) (Allarthrum).</p> <p>Cryphalus kurenzovi Stark, 1936: 150.</p> <p>= Cryphalus punctulatus Eggers, 1942: 29 (syn: Pfeffer, 1944).</p> <p>= Cryphalus ussuriensis Eggers, 1942: 29 (syn: Schedl, 1952e).</p> <p>Cryphalus kyotoensis Nobuchi, 1966a: 53.</p> <p>Cryphalus laevis (Browne, 1980a: 374) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus laricis Niisima, 1909: 142.</p> <p>Cryphalus laticollis (Browne, 1974: 67) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus latus Eggers, 1929a: 10.</p> <p>= Cryphalus premayaensis Murayama, 1943: 97 (syn: Mandelshtam et al., 2007).</p> <p>Cryphalus lepocrinus Tsai and Li, 1963: 619.</p> <p>Cryphalus lipingensis Tsai and Li, 1959: 90.</p> <p>Cryphalus longior Browne, 1984d: 91.</p> <p>Cryphalus longipennis (Browne, 1970: 552) (Taenioglyptes) comb. nov. [Hypocryphalus].</p> <p>Cryphalus longipilis (Browne, 1981: 128) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus longipilus Schedl, 1943: 34.</p> <p>Cryphalus longisetosus (Nobuchi, 1975: 57) (Taenioglyptes).</p> <p>Cryphalus longus (Eggers, 1926b: 136) (Ernoporus).</p> <p>= Cryphalus alni Krivolutskaya, 1958: 145 (syn: Mandelshtam, 2002).</p> <p>Cryphalus luteus Johnson nom. nov. [Margadillius].</p> <p>= Margadillius fulvus Browne, 1984b: 289 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Margadillius fulvus Browne, 1984b after its transfer to Cryphalus, because the name is already occupied by Cryphalus fulvus Niisima, 1908. The name is an adjective, meaning either brown or yellow, maintaining the original meaning.</p> <p>Cryphalus magnus (Browne, 1984d: 88) (Margadillius) comb. nov. [Margadillius].</p> <p>Cryphalus major Stebbing, 1903: 270.</p> <p>= Cryphalus morinda Stebbing, 1903: 265 (syn: Wood, 1989).</p> <p>Cryphalus malayensis (Schedl, 1942a: 176) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus malloti Schedl, 1943: 37.</p> <p>Cryphalus malus Niisima, 1909: 144.</p> <p>= Cryphalus padi Krivolutskaya, 1958: 144 (syn: Mandelshtam, 2002).</p> <p>Cryphalus mandschuricus Eggers, 1929b: 10.</p> <p>Cryphalus mangiferae Stebbing, 1914: 542 comb. res. [Hypocryphalus] (nomen protectum).</p> <p>= Cryphalus inops Eichhoff, 1872a: 131 (syn: Wood, 1982) (nomen oblitum).</p> <p>= Hypothenemus griseus Blackburn, 1885: 194 (syn: Wood, 1982) (nomen oblitum).</p> <p>= Hypocryphalus mangiferae Eggers, 1928: 85 (syn: Eggers, 1930).</p> <p>= Cryphalus subcylindricus Schedl, 1942d: 16 (syn: Schedl, 1958e).</p> <p>= Cryphalus mimicus Schedl, 1942d: 17 (syn: Kalshoven, 1958).</p> <p>= Hypocryphalus opacus Schedl, 1942d: 20.</p> <p>Cryphalus margadilaonis (Hopkins, 1915: 38) (Margadillius) comb. nov. [Margadillius].</p> <p>Cryphalus markangensis Tsai and Li, 1963: 618.</p> <p>Cryphalus massonianus Tsai and Li, 1963: 613.</p> <p>Cryphalus meridionalis (Nobuchi, 1975: 55) (Taenioglyptes).</p> <p>Cryphalus mindoroensis (Schedl, 1943: 39) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus minimus Eggers, 1927c: 76.</p> <p>Cryphalus minor (Schedl, 1943: 40) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus minusculus Johnson nom. nov. [Hypocryphalus].</p> <p>= Hypocryphalus minutus Browne, 1980c: 495 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Hypocryphalus minutus Browne, 1980c following its transfer to Cryphalus. The name is occupied by Margadillius minutus Hopkins, 1915. The replacement name is an adjective, and maintains the original meaning, referencing the very small size.</p> <p>Cryphalus minutus (Hopkins, 1915: 37) (Margadillius) comb. nov. [Margadillius].</p> <p>Cryphalus miyalopiceus Tsai and Li, 1963: 608.</p> <p>Cryphalus mollis Schedl, 1955a: 288 comb. res. [Hypocryphalus].</p> <p>= Hypocryphalus tongaensis Schedl, 1979c: 104 (syn: Beaver, 1987). Cryphalus montanus Nobuchi, 1964: 129.</p> <p>Cryphalus moorei (Schedl, 1964b: 247) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus neglectus Schedl, 1962d: 106.</p> <p>Cryphalus negrosensis Browne, 1979: 85.</p> <p>Cryphalus niger Schedl, 1942c: 172.</p> <p>Cryphalus nigericus Browne, 1973: 289.</p> <p>Cryphalus nigricans Schedl, 1943: 35.</p> <p>Cryphalus nigrosetosus (Schedl, 1948b: 27) (Hypocryphalus) comb.</p> </div>	https://treatment.plazi.org/id/1C598301282AFF83FF285994FCF4FF18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012828FF83FC885994FB64F815.text	1C5983012828FF83FC885994FB64F815.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryphalus pubescens Hopkins 1915	<div><p>Cryphalus pubescens Hopkins, 1915: 39.</p> <p>= Cryphalus subconcentralis Hopkins, 1915: 40 (syn: Wood, 1954).</p> <p>Cryphalus pulchellus (Nobuchi, 1975: 54) (Taenioglyptes).</p> <p>Cryphalus punctatostriatus Schedl, 1942a: 175.</p> <p>Cryphalus punctatus Browne, 1984d: 66.</p> <p>Cryphalus punctipennis Schedl, 1942c: 169.</p> <p>Cryphalus punctistriatulus Johnson nom. nov.</p> <p>= Cryphalus striatulus Browne, 1981: 127 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Cryphalus striatulus Browne, 1981 because the name is occupied, following the revised status of Cryphalomorphus striatulus Browne, 1978. The new is formed from the diminutive of the adjective punctatus meaning punctured, and s striatus meaning with lines or rows.</p> <p>Cryphalus pusillimus Schedl, 1942c: 171.</p> <p>Cryphalus pusillus Schedl, 1943: 38.</p> <p>Cryphalus quadrituberculatus (Schedl, 1963c: 64) (Margadillius) comb. nov. [Margadillius].</p> <p>Cryphalus redikorzevi Berger, 1917: 232.</p> <p>Cryphalus reflexus (Browne, 1980b: 383) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus resiniferi Schedl, 1943: 36.</p> <p>Cryphalus rhusii Niisima, 1909: 145.</p> <p>= Cryphalus kurilensis Krivolutskaya, 1968: 53 (syn: Mandelshtam, 2006).</p> <p>Cryphalus robustus Eichhoff,1872a: 131 comb. res. [Hypocryphalus]. Cryphalus rotundus (Hopkins, 1915: 41) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus rubentis Hopkins, 1915: 40.</p> <p>Cryphalus ruficollis Hopkins, 1915: 40.</p> <p>= Cryphalus fraseri Hopkins, 1915: 40 (syn: Wood, 1989).</p> <p>= Cryphalus approximatus Hopkins, 1915: 41 (syn: Wood, 1954). = Cryphalus balsameus Hopkins, 1915: 41 (syn: Wood, 1954).</p> <p>= Cryphalus amabilis Chamberlin, 1917: 321 (syn: Wood, 1954). = Cryphalus grandis Chamberlin, 1917: 323 (syn: Wood, 1954).</p> <p>= Cryphalus canadensis Chamberlin, 1918: 88 (syn: Wood, 1954). = Cryphalus mainensis Blackman, 1922b: 126 (syn: Wood, 1954). = Cryphalus ruficollis coloradensis Wood, 1954: 1008 (syn: Wood, 1977c).</p> <p>Cryphalus rufopilosus Schedl, 1942a: 174.</p> <p>Cryphalus rugosus (Schedl, 1958b: 102) (Ericryphalus).</p> <p>Cryphalus saltuarius Weise, 1891: 336 (nomen protectum).</p> <p>= Bostrichus asperatus Ratzeburg, 1837: 163 (unavailable name). = Cryphalus scriba Gozis, 1886: 31 (nomen oblitum).</p> <p>Cryphalus samoensis Beeson, 1929: 224.</p> <p>Cryphalus sandakanensis Schedl, 1937b: 548 comb. res. [Hypocryphalus].</p> <p>= Hypocryphalus maculatus Browne, 1962a: 303 (syn: Wood, 1989).</p> <p>Cryphalus sarawakensis Schedl, 1972h: 199.</p> <p>Cryphalus sawadai Nobuchi and Takahashi, 1965: 3.</p> <p>Cryphalus scabripennis Schedl, 1972h: 199.</p> <p>Cryphalus schedli Johnson nom. nov. [Hypocryphalus].</p> <p>= Hypocryphalus formosanus Schedl, 1952b: 62 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Hypocryphalus formosanus Schedl, 1952b because the name is occupied by Cryphalus formosanus Schedl, 1942a. The replacement name is a noun in the genitive case, honoring the original author. The types of both species were unavailable for study.</p> <p>Cryphalus scopiger Berger, 1917: 228.</p> </div>	https://treatment.plazi.org/id/1C5983012828FF83FC885994FB64F815	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012829FF82FF285994FEC5F839.text	1C5983012829FF82FF285994FEC5F839.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryphalus securus (Schedl 1940) Ericryphalus	<div><p>Cryphalus securus (Schedl, 1940a: 436) (Ericryphalus).</p> <p>Cryphalus sejugatus Schedl, 1965a: 54.</p> <p>Cryphalus sichotensis Kurentsov, 1941: 152.</p> <p>Cryphalus sidneyanus (Nördlinger, 1856: 75) (Bostrichus).</p> <p>Cryphalus silvanus Schedl, 1951d: 145.</p> <p>Cryphalus similis (Eggers, 1922: 168) (Stephanoderes).</p> <p>Cryphalus simplex (Schedl, 1939b: 341) (Ericryphalus).</p> <p>Cryphalus sinoabietis Tsai and Li, 1963: 606.</p> <p>⊇ Cryphalus sinoabietis opienensis Tsai and Li, 1963: 607.</p> <p>Cryphalus solomonensis Johnson nom. nov.</p> <p>= Margadillius terminaliae Browne, 1984c: 450 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Margadillius terminaliae Browne, 1984c because the name is already occupied by Cryphalus terminaliae Browne, 1980b. The replacement name is a toponym derived from the type locality.</p> <p>Cryphalus sordidus (Nobuchi, 1975: 51) (Taenioglyptes).</p> <p>Cryphalus sparsepilosus Schedl, 1942c: 172.</p> <p>Cryphalus spathulatus (Schedl, 1938b: 49) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus spissepilosus Johnson nom. nov.</p> <p>= Cryphalus densepilosus Schedl, 1943: 36 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Cryphalus densepilosus Schedl, 1943 because the name is now occupied by Hypocryphalus densepilosus Schedl, 1942d. The replacement name is formed from the adverb spisse (= densely) and the adjective pilosus (= hairy).</p> <p>Cryphalus squameus Schedl, 1943: 38.</p> <p>Cryphalus squamulosus Strohmeyer, 1911: 20.</p> <p>Cryphalus storckiellae Johnson nom. nov.</p> <p>= Cryphalus striatus Browne, 1974: 66 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Cryphalus striatus Browne, 1974 because the name is now occupied by Hypocryphalus striatus Hopkins, 1915. The replacement name is a noun in the genitive case, named after the host plant Storckiella.</p> <p>Cryphalus striatulus (Browne, 1978: 576) (Cryphalomorphus) comb. nov. [Hypothenemus].</p> <p>= Hypothenemus brownei Beaver, 1991: 53 syn. nov. (unnecessary replacement name).</p> <p>Remarks: Cryphalus striatulus (Browne, 1978) is transferred to Cryphalus based on the external diagnostic characters visible. The replacement name is no longer necessary.</p> <p>Cryphalus striatus (Hopkins, 1915: 41) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus strigilatus Wichmann, 1914: 137.</p> <p>Cryphalus strigipennis Schedl, 1950b: 50.</p> <p>Cryphalus strohmeyeri Stebbing, 1914: 538.</p> <p>= Cryphalus indicus Stebbing, 1902: 403 (syn: Stebbing, 1914) (Permanently invalid name due to primary homonymy).</p> <p>Cryphalus subcompactus Lea, 1910: 140.</p> <p>Cryphalus subgranulatus Schedl, 1943: 37.</p> <p>Cryphalus submuricatus Eichhoff, 1878a: 385.</p> <p>= Cryphalus submuricatus Eichhoff, 1878b: 492 syn. nov.</p> <p>= Cryphalus tuberculatus Schedl, 1943: 37 syn. nov.</p> <p>Cryphalus substriatus Schedl, 1942a: 174.</p> <p>Cryphalus subtuberculatus Schedl, 1942c: 168.</p> <p>Cryphalus subvestitus Schedl, 1959a: 482.</p> <p>Cryphalus sumatranus (Schedl, 1939c: 38) (Dacryphalus) comb. nov. [Hypocryphalus].</p> </div>	https://treatment.plazi.org/id/1C5983012829FF82FF285994FEC5F839	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012829FF82FC885994FB4DF999.text	1C5983012829FF82FC885994FB4DF999.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryphalus sundaensis Schedl 1942	<div><p>Cryphalus sundaensis Schedl, 1942d: 14.</p> <p>Cryphalus sylvicola (Perkins, 1900: 181) (Hypothenemus). = Ericryphalus henshawi Hopkins, 1915: 38 (syn: Schedl, 1941b). = Piperius pini Hopkins, 1915: 39 (syn: Schedl, 1963c). = Cryphalus swezeyi Schedl, 1942b: 147 (syn: Wood, 1960). = Cryphalus dimorphus Schedl, 1950b: 38 (syn: Beaver, 1991). = Cryphalus sylvicola obliquus Schedl, 1950b: 48.</p> <p>Cryphalus szechuanensis Tsai and Li, 1963: 614.</p> <p>⊇ Cryphalus szechuanensis tehchangensis Tsai and Li, 1963: 615. Cryphalus tabulaeformis Tsai and Li, 1963: 609.</p> <p>⊇ Cryphalus tabulaeformis chienzhuangensis Tsai and Li, 1963: 610.</p> <p>Cryphalus taiwanus Schedl, 1970c: 359.</p> <p>Cryphalus takahashii Johnson nom. nov. [Ernoporus]</p> <p>= Euptilius exiguus Browne, 1984c: 451 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Euptilius exiguus Browne, 1984c because the name is occupied by Cryphalus exiguus Blandford, 1894b. The replacement name is a noun in the genitive case, honoring the original collector, K. Takahashi.</p> <p>Cryphalus tenuis Schedl, 1942d: 16.</p> <p>Cryphalus terminaliae Browne, 1980b: 384.</p> <p>Cryphalus tetricus (Schedl, 1940a: 437) (Ericryphalus).</p> <p>= Cryphalus papuanus Schedl, 1942c: 170 (syn: Schedl, 1979c). = Cryphalus grayi Schedl, 1968: 265 (syn: Schedl, 1979c). = Ptilopodius brevis Browne, 1970: 556 (syn: Schedl, 1972j). Cryphalus theobromae Nunberg, 1956b: 208 (replacement name). = Cryphalus horridus Graham, 1908: 113 (syn: Nunberg, 1956b) (permanently invalid name due to primary homonymy).</p> <p>Cryphalus triangularis (Schedl, 1975a: 351) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus trypanoides Beeson, 1935a: 106.</p> <p>Cryphalus trypanus Sampson, 1914: 383.</p> <p>Cryphalus tutuilaensis (Schedl, 1951d: 147) (Hypocryphalus) comb. nov. [Hypocryphalus].</p> <p>Cryphalus uapouensis (Beeson, 1935a: 107) (Ericryphalus). Cryphalus upoluensis Schedl, 1951d: 144.</p> <p>Cryphalus variolosus Schedl, 1950b: 49.</p> <p>Cryphalus vestitus Blandford, 1895: 318.</p> <p>Cryphalus viburni Stark, 1936: 151.</p> <p>= Cryphalus viburni Eggers, 1942: 30 (syn: Pfeffer, 1944). Cryphalus vitiensis Browne, 1974: 67.</p> <p>Cryphalus walkeri (Blandford, 1896a: 200) (Cryptarthrum). = Coccotrypes hagedorni Eggers, 1908: 216 (syn: Schedl, 1963c). = Cryphalus javanus Schedl, 1954a: 148 (syn: Schedl, 1958e). Cryphalus wapleri Eichhoff, 1872a: 131.</p> <p>= Cryphalus mekeoi Schedl, 1972e: 61 (syn: Schedl, 1979d). Cryphalus yamaguchii Inouye and Nobuchi, 1957: 55.</p> <p>Cryphalus zimmermani Schedl, 1950b: 51.</p> </div>	https://treatment.plazi.org/id/1C5983012829FF82FC885994FB4DF999	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012829FF8DFC885F5EFE78FF53.text	1C5983012829FF8DFC885F5EFE78FF53.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trypophloeini Nusslin 1911	<div><p>Trypophloeini Nüsslin, 1911 stat. res.</p> <p>=Trypophloeinae Nüsslin, 1911b: 375.</p> <p>Type of tribe</p> <p>Trypophloeus Fairmaire, 1864.</p> <p>Diagnosis</p> <p>Tarsi with third segment cylindrical. Eye emarginated. Antennal funicle with 3–5 segments. Antennal club with or without sutures, and at most a partial septum. Hypomeron with single setae, or rarely mixed with a few bifurcating setae. Postnotum not fused with metanotum. Typically with scale-like interstrial bristles, and sparse interstrial ground vestiture.</p></div> 	https://treatment.plazi.org/id/1C5983012829FF8DFC885F5EFE78FF53	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012824FF8FFF285D2FFE6DFB46.text	1C5983012824FF8FFF285D2FFE6DFB46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atomothenemus Bright 2019	<div><p>Atomothenemus Bright, 2019: 106</p> <p>Type of genus</p> <p>Atomothenemus unicus Bright, 2019.</p> <p>Diagnosis</p> <p>Antennae with two funicle segments (including the pedicel), antennal club with horizontal sutures.</p></div> 	https://treatment.plazi.org/id/1C5983012824FF8FFF285D2FFE6DFB46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012824FF8FFC8858D7FAEFFBFC.text	1C5983012824FF8FFC8858D7FAEFFBFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cosmoderes Eichhoff 1878	<div><p>Cosmoderes Eichhoff, 1878a: 387 (Figs. 14–15)</p> <p>Synonymy</p> <p>= Cosmoderes Eichhoff, 1878b: 495.</p> <p>= Erioschidias Schedl, 1938b: 42 (unavailable name).</p> <p>= Cosmoderus Chamberlin, 1939: 274 (unavailable name). = Allernoporus Kurentsov, 1941: 159 syn. nov.</p> <p>= Vitideres Beeson, 1941: 391 (unavailable name).</p> <p>= Erischidias Schedl, 1942d: 9 (unavailable name).</p> <p>= Dendriops Schedl, 1953a: 125.</p> <p>= Erioschidias Wood, 1960: 21.</p> <p>= Vitideres Beeson, 1961: 301 (unavailable name). = Pseudocosmoderes Nobuchi, 1981: 16.</p> <p>= Vitaderes Wood and Bright, 1992: 901 (unavailable name).</p> <p>Type of genus</p> <p>Cosmoderes monilicollis Eichhoff, 1878a.</p> <p>Diagnosis</p> <p>Body shape elongated, antennae with short funicle, antennal club large and without sutures, marginal asperities present, mesocoxae contiguous, anterior plate of proventriculus short.</p></div> 	https://treatment.plazi.org/id/1C5983012824FF8FFC8858D7FAEFFBFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301281AFFB0FF285994FB30F838.text	1C598301281AFFB0FF285994FB30F838.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypothenemus japonicus (Niisima 1910) Cryphalus	<div><p>Hypothenemus japonicus (Niisima, 1910: 10) (Cryphalus).</p> <p>Hypothenemus javanus (Eggers, 1908: 215) (Stephanoderes).</p> <p>= Stephanoderes obesus Hopkins, 1915: 30 (syn: Wood, 1957b).</p> <p>= Stephanoderes brunneus Hopkins, 1915: 31 (syn: Bright, 2019.</p> <p>= Stephanoderes frontalis Hopkins, 1915: 31 (syn: Wood, 1954).</p> <p>= Stephanoderes philippinensis Hopkins, 1915: 31 (syn: Wood, 1975b).</p> <p>= Stephanoderes bananensis Eggers, 1922: 167 (syn: Wood, 1972b).</p> <p>= Stephanoderes kalshoveni Schedl, 1939c: 35 (syn: Wood, 1972b).</p> <p>= Stephanoderes cryphalomorphus Schedl, 1939e: 14 (syn: Wood, 1977a).</p> <p>= Stephanoderes bituberculatus Eggers, 1940a: 126 (syn: Wood, 1977a).</p> <p>= Stephanoderes subagnatus Eggers, 1940b: 101 (syn: Wood, 1972b).</p> <p>= Stephanoderes pistor Schedl, 1951e: 102 (syn: Wood, 1976).</p> <p>= Stephanoderes prosper Schedl, 1951e: 103 (syn: Wood, 1976).</p> <p>Hypothenemus kamathi Beaver, 1995: 16.</p> <p>Hypothenemus kraunhiae Murayama, 1950: 50.</p> <p>Hypothenemus lefevrei (Schedl, 1952d: 8) (Stephanoderes).</p> <p>Hypothenemus leprieuri (Perris, 1866: 194) (Dryocoetes).</p> <p>= Stephanoderes albipilis Reitter, 1887: 195 (syn: Eggers, 1940e).</p> <p>= Dryocoetes kraussei Wichmann, 1911: 210 (syn: Eggers, 1921). Hypothenemus leptosquamus Bright, 2019: 152.</p> <p>Hypothenemus liberiensis (Hopkins, 1915: 31) (Stephanoderes).</p> <p>= Stephanoderes theobromae Eggers, 1932: 32 (syn: Wood, 1968). Hypothenemus liliputianus Bright, 2019: 152.</p> <p>Hypothenemus lineatus (Eggers, 1927a: 175) (Stephanoderes).</p> <p>Hypothenemus longipennis (Eggers, 1935: 305) (Stephanoderes).</p> <p>Hypothenemus longipilis Schedl, 1952a: 451.</p> <p>Hypothenemus loranthus (Schedl, 1942d: 8) (Margadillius) comb. nov. [Margadillius].</p> <p>Hypothenemus macrolobii (Eggers, 1940c: 234) (Stephanoderes).</p> <p>Hypothenemus madagascariensis Schedl, 1953c: 82.</p> <p>Hypothenemus magnus (Eggers, 1924: 102) (Stephanoderes).</p> <p>Hypothenemus major Browne, 1970: 555.</p> <p>Hypothenemus malayensis (Schedl, 1977c: 499) (Lepiceriodes).</p> <p>Hypothenemus mallyi (Hopkins, 1915: 32) (Stephanoderes).</p> <p>= Stephanoderes soussouensis Eggers, 1943b: 74 (syn: Wood, 1972b).</p> <p>Hypothenemus malus (Schedl, 1957b: 56) (Stephanoderes).</p> <p>Hypothenemus mangovorus Schedl, 1961b: 134.</p> <p>Hypothenemus marginatus Johnson nom. nov.</p> <p>= Periocryphalus sobrinus Wood, 1974a: 22 syn. nov.</p> <p>Remarks: A replacement name is proposed for Periocryphalus sobrinus Wood, 1974 following the generic synonymy of Hypothenemus and Periocryphalus, creating conflict with Stephanoderes sobrinus Schedl, 1965a. The replacement name is an adjective, referencing the prominent posterolateral margin of the elytra. The holotype of Periocryphalus sobrinus Wood, 1974 at BMNH is lost and erroneously replaced with a Cnesinus sp., which clearly does not match the species description. A paratype at USNM collected from the same location on a different date matches the original and subsequent descriptions of the species. (Labeled ‘ PARATYPE // Periocryphalus // sobrinus // S. L. Wood 1972 //// Periocryphalus //sobrinus//.72 Wood // S. L. Wood // RS/RGS Exp. Brazil // 12′31′S 51′46′W //// R. A. Beaver F03 // 7.XI.1968 //// S. L. Wood Collection //// USNMENT 01448187’.) Since this species is the type of the genus Periocryphalus, the stability of the name-bearing type is critical. Therefore, the single remaining specimen in the type series at USNM is designated as a neotype of Periocryphalus sobrinus</p> <p>Wood, 1974a, and subsequently, the neotype of Hypothenemus marginatus Johnson.</p> <p>Hypothenemus marshalli (Eggers, 1936b: 36) (Stephanoderes).</p> <p>Hypothenemus mateui (Schedl, 1965g: 198) (Stephanoderes).</p> <p>Hypothenemus melanarius (Schedl, 1953c: 80) (Cosmoderes).</p> <p>Hypothenemus melasomus (Lea, 1910: 140) (Cryphalus).</p> <p>Hypothenemus meridensis Wood, 2007: 506.</p> <p>Hypothenemus miles (LeConte, 1878: 433) (Cryphalus).</p> <p>Hypothenemus minor (Eggers, 1927a: 178) (Adiaeretus).</p> <p>Hypothenemus modestus (Murayama, 1940: 236) (Cryphalus).</p> <p>Hypothenemus morigerus (Schedl, 1957b: 57) (Stephanoderes).</p> <p>Hypothenemus morio (Eggers, 1940b: 101) (Stephanoderes).</p> <p>Hypothenemus morosus Schedl, 1965a: 57.</p> <p>Hypothenemus mozambiquensis Eggers, 1943b: 75.</p> <p>Hypothenemus mulongensis (Eggers, 1940c: 235) (Stephanoderes).</p> <p>Hypothenemus multidentatulus (Schedl, 1962f: 95) (Stephanoderes). Hypothenemus multidentatus (Hopkins, 1915: 28) (Stephanoderes). = Stephanoderes ferrugineus Hopkins, 1915: 29 (syn: Wood, 1972b). = Stephanoderes nitidifrons Hopkins, 1915: 31 (syn: Wood, 1972b). = Hypothenemus hopkinsi Browne, 1963b: 53 (syn: Wood, 1972b). Hypothenemus multipunctatus (Schedl, 1939c: 36) (Stephanoderes). Hypothenemus muticus (Schedl, 1961b: 136) (Stephanoderes).</p> <p>Hypothenemus namosianus Browne, 1983a: 78.</p> <p>Hypothenemus nanellus Wood, 1971: 34.</p> <p>Hypothenemus nanoparvus Bright, 2019: 154.</p> <p>Hypothenemus natalensis (Schedl, 1941a: 392) (Archeophalus).</p> <p>Hypothenemus nesiotus Bright, 2019: 154.</p> <p>Hypothenemus nigropiceus (Schedl, 1951c: 20) (Stephanoderes).</p> <p>Hypothenemus novateutonicus (Schedl, 1951e: 105) (Ptilopodius) comb. nov. [Ptilopodius].</p> <p>Hypothenemus obscurifrons Bright, 2019: 155.</p> <p>Hypothenemus obscurus (Fabricius, 1801: 395) (Hylesinus).</p> <p>= Stephanoderes asperulus Eichhoff, 1872a: 133.</p> <p>= Stephanoderes cassiae Eichhoff, 1878b: 152.</p> <p>= Hypothenemus kuennemanni Reitter, 1902: 140 (syn: Wood, 1972b).</p> <p>= Stephanoderes moschatae Schaufuss, 1905: 8 (syn: Wood, 1972b). = Stephanoderes rufescens Hopkins, 1915: 29 (syn: Wood, 1972b).</p> <p>= Stephanoderes buscki Hopkins, 1915: 30 (syn: Wood, 1972b).</p> <p>= Stephanoderes amazonicus Eggers, 1934: 78 (syn: Wood, 1972b).</p> <p>= Hypothenemus emarginatus Schedl, 1942d: 11 (syn: Wood, 1972b).</p> <p>Remarks: Bright (2019) noted that Hypothenemus seriatus (Eichhoff, 1872) was a junior synonym of H. obscurus (Fabricius, 1801), as it was synonymized by Wood (1954), based on a specimen compared with the type by Eggers, but the synonymy was largely ignored (including by Wood). We consider the synonymy originally proposed by Wood to be rejected. Hypothenemus obscurus is a highly damaging pest which is not present in some areas where H. seriatus is present. We found that the species were sister in the phylogeny. The morphological differences have been well documented (Vega et al., 2015), and they are genetically distinct (Mitchell and Maddox 2010, Johnson et al 2018). It is possible that there is much more variation than documented which may make definitive identification by morphology difficult or impossible, but more evidence would be needed to justify grouping the distinct morphotypes as one species, as well as alternative recognition of the pestiferous beetles which devastate the macadamia and other tropical nut industries. The economic and policy implications of the synonymy could hinder the necessary biosecurity measures to prevent the pest species establishing in other areas.</p> <p>Hypothenemus opacus (Eichhoff, 1872a: 132) (Stephanoderes).</p> <p>= Hypothenemus dolosus Wood, 1974a: 21 (syn: Bright, 2019). Hypothenemus paradoxus Schedl, 1964c: 45. Hypothenemus parallelus (Hopkins, 1915: 25) (Stephanoderes). Hypothenemus parasquamosus Bright, 2019: 160. Hypothenemus parvulosus Bright, 2019: 160. Hypothenemus parvulus Browne, 1984d: 93. Hypothenemus paulus Bright, 2019: 162. Hypothenemus perappositus (Schedl, 1934: 91) (Stephanoderes). Hypothenemus perexiguus Bright, 2019: 162. Hypothenemus perhispidus (Eggers, 1927a: 177) (Stephanoderes). Hypothenemus perpunctatus (Eggers, 1940c: 233) (Stephanoderes). Hypothenemus piaparolinae Johnson, Atkinson and Hulcr, 2016: 418. Hypothenemus pilosus Hopkins, 1915: 20. Hypothenemus ponticus Bright, 2019: 164. Hypothenemus praecellens (Schedl, 1972c: 288) (Stephanoderes). Hypothenemus pubescens Hopkins, 1915: 19.</p> <p>= Hypothenemus subelongatus Hopkins, 1915: 19 (syn: Wood, 1972b).</p> <p>= Stephanoderes opacifrons Hopkins, 1915: 25 (syn: Wood, 1972b).</p> <p>= Hypothenemus minutissimus Schedl, 1952a: 450 (syn: Wood, 1989). Hypothenemus pubipennis (Eggers, 1935: 305) (Stephanoderes). Hypothenemus puertoricensis (Bright and Torres, 2006: 409) (Trischidias). Hypothenemus pullus (Wood, 1971: 33) (Periocryphalus) comb. nov. [Periocryphalus]. Hypothenemus pygmaeomorphus Bright, 2019: 166. Hypothenemus rotundicollis (Eichhoff,1878a: 385) (Stephanoderes).</p> <p>= Stephanoderes sculpturatus Eichhoff, 1878a: 385 syn. nov. (syn: Eichhoff, 1896).</p> <p>= Stephanoderes rotundicollis Eichhoff, 1878b: 45 syn. nov.</p> <p>= Stephanoderes sculpturatus Eichhoff, 1878b: 45 syn. nov.</p> <p>= Stephanoderes quercus Hopkins, 1915: 32 syn. nov. Hypothenemus rubrithorax Bright, 2019: 168. Hypothenemus ruficeps Perkins, 1900: 181. Hypothenemus rugifer (Schedl, 1965d: 9) (Pachynoderes). Hypothenemus ruginosus Wood, 1989: 178.</p> <p>= Pachynoderes rugifer Schedl, 1977a: 395 (syn: Wood, 1989) (Permanently invalid name due to primary homonymy). Hypothenemus rugosipes Wood, 2007: 515. Hypothenemus sambesianus Eggers, 1943b: 74. Hypothenemus sapporoensis (Niisima, 1910: 3) (Cryphalus). Hypothenemus sassaensis (Eggers, 1924: 102) (Stephanoderes). Hypothenemus schedli Browne, 1963b: 54.</p> <p>= Stephanoderes pubescens Schedl, 1942d: 18 (syn: Browne, 1963b). Hypothenemus scutiae (Schedl, 1959c: 708) (Stephanoderes). Hypothenemus seoulensis Choo and Woo, 1989: 58. Hypothenemus seriatus (Eichhoff, 1872a: 133) (Stephanoderes) stat. res.</p> <p>= Stephanoderes pulverulentus Eichhoff, 1872a: 133 (syn: Wood, 1973).</p> <p>= Stephanoderes vulgaris Schaufuss, 1897: 209 (syn: Wood, 1972b).</p> <p>= Stephanoderes georgiae Hopkins, 1915: 26 (syn: Wood, 1972b).</p> <p>= Stephanoderes minutus Hopkins, 1915: 26 (syn: Wood, 1972b).</p> <p>= Stephanoderes texanus Hopkins, 1915: 26 (syn: Wood, 1954).</p> <p>= Stephanoderes ficus Hopkins, 1915: 27 (syn: Wood, 1954).</p> <p>= Stephanoderes fiebrigi Hopkins, 1915: 27 (syn: Wood, 1954).</p> <p>= Stephanoderes floridensis Hopkins, 1915: 27 (syn: Wood, 1954).</p> <p>= Stephanoderes pini Hopkins, 1915: 27 (syn: Wood, 1954).</p> <p>= Stephanoderes salicis Hopkins, 1915: 27 (syn: Wood, 1954).</p> <p>= Stephanoderes tamarindi Hopkins, 1915: 27 (syn: Wood, 1972b).</p> <p>= Stephanoderes lucasi Hopkins, 1915: 28 (syn: Wood, 1954).</p> <p>= Stephanoderes soltaui Hopkins, 1915: 28 (syn: Wood, 1954).</p> <p>= Stephanoderes virentis Hopkins, 1915: 28 (syn: Wood, 1954).</p> <p>= Stephanoderes nitidipennis Hopkins, 1915: 29 (syn: Wood, 1972b).</p> <p>= Stephanoderes nitidulus Hopkins, 1915: 29 (syn: Wood, 1977c).</p> <p>= Stephanoderes pecanis Hopkins, 1915: 29 (syn: Wood, 1954).</p> <p>= Stephanoderes subopacicollis Hopkins, 1915: 30 (syn: Wood, 1977c).</p> <p>= Stephanoderes niger Hopkins, 1915: 31 (syn: Wood, 1972b).</p> <p>= Hypothenemus robustus Blackman, 1922a: 88 (syn: Wood, 1954).</p> <p>= Hypothenemus cassavaensis Schedl, 1938a: 453 (syn: Wood, 1989).</p> <p>= Stephanoderes hawaiiensis Schedl, 1941b: 112 (syn: Wood, 1989).</p> <p>= Stephanoderes darwinensis Schedl, 1942c: 178 (syn: Wood, 1972b).</p> <p>= Hypothenemus striatulus Schedl, 1942d: 12 (syn: Wood, 1989).</p> <p>= Hypothenemus marovoayi Schedl, 1953c: 81 (syn: Wood, 1989).</p> <p>= Stephanoderes andersoni Wood, 1954: 1045 (syn: Wood, 1972b).</p> <p>= Stephanoderes liquidambarae Wood, 1954: 1046 (syn: Wood, 1972b).</p> <p>= Stephanoderes asperatus Schedl, 1967c: 226 (syn: Wood, 1989). Hypothenemus setiferous Bright, 2019: 169.</p> <p>Hypothenemus setosus (Eichhoff, 1868: 391) (Hypoborus).</p> <p>= Stephanoderes congonus Hagedorn, 1912a: 337 (syn: Wood, 1975b).</p> <p>Hypothenemus simoni (Reitter, 1887: 194) (Stephanoderes).</p> <p>Hypothenemus sobrinus (Schedl, 1965a: 58) (Stephanoderes).</p> <p>Hypothenemus socialis (Schedl, 1957b: 57) (Stephanoderes).</p> <p>Hypothenemus solitarius (Schedl, 1950a: 24) (Stephanoderes).</p> <p>Hypothenemus solivagus Bright, 2019: 171.</p> <p>Hypothenemus solocis Wood, 1974a: 21.</p> <p>Hypothenemus sparsedentatus (Schedl, 1942c: 178) (Stephanoderes). Hypothenemus sparsus Hopkins, 1915: 20.</p> <p>= Hypothenemus similis Hopkins, 1915: 20 (syn: Wood, 1954).</p> <p>= Stephanoderes tridentatus Hopkins, 1915: 31 (syn: Wood, 1954). Hypothenemus spinatus (Schedl, 1977b: 282) (Miocryphalus).</p> <p>Hypothenemus spinicollis (Schedl, 1965a: 59) (Stephanoderes).</p> <p>Hypothenemus spinosus (Schedl, 1979c: 98) (Lepiceroides).</p> <p>Hypothenemus squamosulus Johnson nom. nov. [Ptilopodius].</p> <p>= Ptilopodius squamosus Schedl, 1953b: 294 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name is proposed for Ptilopodius squamosus Schedl, 1953b after transferring this species from Ptilopodius, since the combination is already occupied by Stephanoderes squamosus Hopkins, 1915. The new name is the diminutive of the adjective squamosus (= scaly), referring to the scalelike setae on the pronotum and elytra.</p> <p>Hypothenemus squamosus (Hopkins, 1915: 26) (Stephanoderes).</p> <p>Hypothenemus stigmosus (Schedl, 1951e: 101) (Stephanoderes).</p> <p>= Stephanoderes garciae Schedl, 1958c: 42 (syn: Wood, 1989).</p> <p>Hypothenemus striatus (Atkinson, 1993: 422) (Trischidias).</p> <p>Hypothenemus styrax (Schedl, 1942a: 177) (Stephanoderes).</p> <p>Hypothenemus subacuminatus (Schedl,1942a: 177) (Stephanoderes). Hypothenemus subterrestris Johnson, Atkinson and Hulcr, 2016: 421.</p> <p>Hypothenemus suspectus Wood, 1974a: 22.</p> </div>	https://treatment.plazi.org/id/1C598301281AFFB0FF285994FB30F838	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012818FFB3FF285994FDD1FB8D.text	1C5983012818FFB3FF285994FDD1FB8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypothenemus tectus Bright 2019	<div><p>Hypothenemus tectus Bright, 2019: 173.</p> <p>Hypothenemus teretis Wood, 1971: 35. Hypothenemus teteforti (Menier, 1971: 141) (Stephanoderes). Hypothenemus tredli (Reitter, 1908: 55) (Cryphalus). Hypothenemus tristis (Eichhoff, 1876: 200) (Stephanoderes).</p> <p>= Stephanoderes taihokuensis Schedl, 1952b: 63. (syn: Beaver, 2011a).</p> <p>= Hypothenemus cosmoderoides Murayama, 1961: 30 (syn: Beaver and Liu, 2010). Hypothenemus trivialis Wood, 1974a: 20. Hypothenemus tuberosus (Schedl, 1942d: 19) (Stephanoderes). Hypothenemus turnbowi Bright, 2019: 173. Hypothenemus ustulatus Bright, 2019: 174. Hypothenemus vernaculus Bright, 2019: 174. Hypothenemus versicolor Bright, 2019: 176. Hypothenemus vesculus Wood, 1974a: 21. Hypothenemus villosus Bright, 2019: 176. Hypothenemus virolae Wood, 2007: 509. Hypothenemus vitis Browne, 1970: 554. Hypothenemus winkleri (Reitter, 1907: 192) (Stephanoderes). Hypothenemus woodi Bright, 2019: 153 (replacement name).</p> <p>= Trischidias minutissimus Wood, 1954: 1069 (syn: Bright, 2019). Hypothenemus xanthophloeae (Schedl, 1957b: 58) (Stephanoderes).</p> <p>Macrocryphalus Nobuchi, 1981: 14 stat. res. (Fig. 19) Type of genus</p> <p>Macrocryphalus oblongus Schedl, 1965.</p> <p>Diagnosis</p> <p>The teardrop-shaped antennae, rounded lateral margins of the pronotum, separated mesocoxae, and absence of a sclerotized proventriculus diagnose members of this genus.</p></div> 	https://treatment.plazi.org/id/1C5983012818FFB3FF285994FDD1FB8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012816FFBDFF285CBAFE97F9C0.text	1C5983012816FFBDFF285CBAFE97F9C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microsomus atomus Bright 2019	<div><p>Microsomus atomus Bright, 2019.</p> <p>Diagnosis</p> <p>This genus can be distinguished from other Trypophloeini by the antennae, with a single, procurved suture, and by the lack of any vestiture on the elytra.</p> </div>	https://treatment.plazi.org/id/1C5983012816FFBDFF285CBAFE97F9C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012816FFBDFF285C6FFEE4FA1B.text	1C5983012816FFBDFF285C6FFEE4FA1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microsomus Bright 2019	<div><p>Microsomus Bright, 2019: 178</p> <p>Type of genus</p></div> 	https://treatment.plazi.org/id/1C5983012816FFBDFF285C6FFEE4FA1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012817FFBCFF2859F0FE29FE70.text	1C5983012817FFBCFF2859F0FE29FE70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pygmaeoborus Bright 2019	<div><p>Pygmaeoborus Bright, 2019: 178</p> <p>Type of genus</p> <p>Pygmaeoborus cubensis Bright, 2019.</p> <p>Diagnosis</p> <p>Similar to Hypothenemus except antennae with single suture indicated by dense setae on apical third.</p> </div>	https://treatment.plazi.org/id/1C5983012817FFBCFF2859F0FE29FE70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012817FFBCFF285C6BFDE5F90D.text	1C5983012817FFBCFF285C6BFDE5F90D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trypophloeus Fairmaire 1864	<div><p>Trypophloeus Fairmaire, 1864: 105 (Fig. 21)</p> <p>Synonymy</p> <p>= Glyptoderes Eichhoff, 1878b: 34.</p> <p>Type of genus</p> <p>Bostrichus binodulus Ratzeburg, 1837.</p> <p>Diagnosis</p> <p>This genus can be diagnosed by the emarginated eye, by the coneshaped antennal club with slightly recurved sutures, and by the rounded lateral margins of the pronotum.</p></div> 	https://treatment.plazi.org/id/1C5983012817FFBCFF285C6BFDE5F90D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012815FFBEFF285994FDF9FDF3.text	1C5983012815FFBEFF285994FDF9FDF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trypophloeus niger Stark 1936	<div><p>Trypophloeus niger Stark, 1936: 152.</p> <p>Trypophloeus populi Hopkins, 1915: 37.</p> <p>Trypophloeus rybinskii Reitter, 1895: 72.</p> <p>⊇ Trypophloeus rybinskii corsicus Eggers, 1912: 113 (syn: Balachowsky, 1949).</p> <p>= Trypophloeus rybinskii salicis Stark, 1952: 283 (Permanently invalid name due to primary homonymy).</p> <p>Trypophloeus salicis Hopkins, 1915: 36.</p> <p>= Trypophloeus concentralis Hopkins, 1915: 36 (syn: Wood, 1954).</p> <p>Trypophloeus striatulus (Mannerheim, 1853: 235) (Cryphalus).</p> <p>= Cryphalus nitidus Swaine, 1912: 349 (syn: Wood, 1969).</p> <p>= Cryphalus punctipennis Hopkins, 1915: 37 (syn: Wood, 1954). Trypophloeus thatcheri (Wood, 1954: 994) (Cryphalus).</p> <p>Trypophloeus tremulae Stark, 1952: 287.</p> </div>	https://treatment.plazi.org/id/1C5983012815FFBEFF285994FDF9FDF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012815FFBEFF285B6CFDEDFBD5.text	1C5983012815FFBEFF285B6CFDEDFBD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xyloterini LeConte 1876	<div><p>Xyloterini LeConte, 1876</p> <p>=Xyloteri LeConte, 1876: 356.</p> <p>=Xyloteroideae Lindemann, 1877: 165.</p> <p>=Xyloteridae Eichhoff, 1878b: 411.</p> <p>=Trypodendrinae Trédl, 1907: 70 (Unavailable name).</p> <p>=Xyloterinae Nüsslin, 1911a: 432.</p> <p>= Xyloterini Reitter, 1913: 28.</p> <p>=Xyloterina Balachowsky, 1949: 196.</p> <p>=Trypodendrina Nunberg, 1954: 16.</p> <p>=Trypodendrini Browne, 1961: 45.</p> <p>=Trypodendroninae Wood, 1978: 114.</p> <p>Type of tribe</p> <p>Xyloterus Erichson, 1836.</p> <p>Diagnosis</p> <p>Eye divided. Mycangia present on females.</p></div> 	https://treatment.plazi.org/id/1C5983012815FFBEFF285B6CFDEDFBD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012813FFB8FF285829FDD7FD58.text	1C5983012813FFB8FF285829FDD7FD58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trypodendron Stephens 1830	<div><p>Trypodendron Stephens, 1830: 353 (Fig. 22 D)</p> <p>Synonymy</p> <p>= Xyloterus Erichson, 1836: 60.</p> <p>= Tripodendron Redtenbacher, 1845: 151 (unavailable name). = Trypodendrum Agassiz, 1846: 380.</p> <p>= Xyloteres Redtenbacher, 1847: 36 (unavailable name). = Xylotrophus Gistel, 1848: 4.</p> <p>= Trypodendrum Gistel, 1856: 368 (unavailable name).</p> <p>Type of genus</p> <p>Dermestes domesticus Linnaeus, 1758.</p> <p>Diagnosis</p> <p>Antennal club with a single acutely procurved suture. Mycangia on hypomeron horizontal (antero-posteriorly).</p></div> 	https://treatment.plazi.org/id/1C5983012813FFB8FF285829FDD7FD58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012813FFB8FC885A37FAE2FBB5.text	1C5983012813FFB8FC885A37FAE2FBB5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xyloterinus Swaine 1918	<div><p>Xyloterinus Swaine, 1918: 44 (Fig. 22 C)</p> <p>Type of genus</p> <p>Bostrichus politus Say, 1826.</p> <p>Diagnosis</p> <p>Antennal club with a single weakly procurved suture. Mycangia on hypomeron horizontal (antero-posteriorly elongate).</p></div> 	https://treatment.plazi.org/id/1C5983012813FFB8FC885A37FAE2FBB5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012813FFBBFC885FE1FED1FEB1.text	1C5983012813FFBBFC885FE1FED1FEB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ernoporini Nusslin 1911	<div><p>Ernoporini Nüsslin, 1911 stat. res.</p> <p>=Ernoporinae Nüsslin, 1911b: 375.</p> <p>=Eidophelinae Murayama, 1954: 200 (Unavailable name). =Eidopherinae Murayama, 1954: 200.</p> <p>Type of tribe</p> <p>Ernoporus Thomson, 1859.</p> <p>Diagnosis</p> <p>Eye entire, rarely broadly emarginated. Antennae with three to five funicle segments. Postnotum fused with metanotum. Penis apodemes short and fused.</p></div> 	https://treatment.plazi.org/id/1C5983012813FFBBFC885FE1FED1FEB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012810FFBBFF285D66FB20FE0C.text	1C5983012810FFBBFF285D66FB20FE0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eidophelus Eichhoff 1876	<div><p>Eidophelus Eichhoff, 1876: 200 (Figs. 23–26)</p> <p>Synonymy</p> <p>= Idophelus Rye, 1877: 362 (unavailable name).</p> <p>= Lepicerus Eichhoff, 1878a: 388 syn. nov.</p> <p>= Scolytogenes Eichhoff, 1878a: 387 syn. nov.</p> <p>= Lepicerus Eichhoff, 1878b: 501 syn. nov.</p> <p>= Scolytogenes Eichhoff, 1878b: 497 syn. nov.</p> <p>= Lepidocerus Rye, 1880: 103 syn. nov.</p> <p>= Cryphalomorphus Schaufuss, 1891: 12 syn. nov.</p> <p>= Letznerella Reitter, 1913: 68 syn. nov.</p> <p>= Ernoporides Hopkins, 1915: 34 syn. nov.</p> <p>= Hypothenoides Hopkins, 1915: 11 syn. nov.</p> <p>= Ptilopodius Hopkins, 1915: 11 syn. nov.</p> <p>= Ernoporicus Berger, 1917: 242 syn. nov.</p> <p>= Neocryphalus Eggers, 1922: 169 syn. nov.</p> <p>= Negritus Eggers, 1923a: 141 syn. nov.</p> <p>= Lepicerinus Hinton, 1936: 473 syn. nov.</p> <p>= Cylindrotomicus Eggers, 1936c: 633 syn. nov. (unavailable name). = Eocryphalus Kurentsov, 1941: 161 syn. nov.</p> <p>= Ernopocerus Balachowsky, 1949: 211 syn. nov. (unavailable name). = Ernoporoides Balachowsky, 1949: 201 syn. nov. (unavailable name).</p> <p>= Ernopocerus Wood, 1954: 986 syn. nov.</p> <p>= Phellodendrophagus Krivolutskaya, 1958: 107.</p> <p>= Cryphalophilus Schedl, 1970c: 358 syn. nov.</p> <p>= Xylocryptus Schedl, 1975a: 352 syn. nov.</p> <p>= Cryphalogenes Wood, 1980: 91 syn. nov.</p> <p>= Ernpocerus Wood, 1982: 858 syn. nov. (unavailable name).</p> <p>= Nigritus Wood and Bright, 1992: 858 syn. nov. (unavailable name).</p> <p>Type of genus</p> <p>Eidophelus imitans Eichhoff, 1876</p> <p>Diagnosis</p> <p>Eidophelus can be diagnosed from superficially similar genera by the entire eye, by the lateral and basal margins of the pronotum marked by a carina, and by the antennal club which has a partial or absent septum, and by the proventriculus characters.</p> </div>	https://treatment.plazi.org/id/1C5983012810FFBBFF285D66FB20FE0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301280BFFA3FC8859B2FCF4FF18.text	1C598301280BFFA3FC8859B2FCF4FF18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytogenes Eichhoff 1878	<div><p>[Scolytogenes].</p> <p>Eidophelus minusculus Johnson nom. nov.</p> <p>= Eidophelus minutissimus Schedl, 1962a: 191 syn. nov. (secondary homonym)</p> <p>Remarks: A replacement name is proposed for Eidophelus minutissimus Schedl, 1962a because the name is now occupied by Ptilopodius minutissimus Schedl, 1943. The replacement name is an adjective, and maintains the original meaning, referencing the very small size.</p> <p>Eidophelus minutissimus (Schedl, 1943: 34) (Ptilopodius) comb. nov. [Ptilopodius].</p> <p>Eidophelus minutus Blandford, 1894b: 88.</p> <p>Eidophelus mus (Schedl, 1975a: 349) (Cryphalomorphus) comb. nov. [Scolytogenes].</p> <p>Eidophelus nanulus (Wood, 1960: 29) (Cryphalomorphus) comb. nov. [Scolytogenes].</p> <p>Eidophelus nigellatus (Schedl, 1950b: 44) (Lepicerinus) comb. nov. [Scolytogenes].</p> <p>Eidophelus niger Johnson nom. nov. [Ernoporicus].</p> <p>= Ernoporicus ater Nobuchi, 1975: 43 syn. nov. (secondary homonym).</p> <p>Remarks: A replacement name for Ernoporicus ater Nobuchi, 1975 is proposed because the name is already occupied by Negritus ater Eggers, 1923a. The name is an adjective, following the meaning of the original name, meaning black.</p> <p>Eidophelus nitidus Schedl, 1965b: 26.</p> <p>Eidophelus nubilus (Wood, 1960: 30) (Cryphalomorphus) comb. nov. [Scolytogenes].</p> <p>Eidophelus ocularis (Schedl, 1965c: 368) (Cryphalomorphus) comb. nov. [Scolytogenes].</p> <p>Eidophelus onyanganus (Schedl, 1941a: 391) (Letznerella) comb. nov. [Scolytogenes].</p> <p>= Cryphalomorphus similaris Schedl, 1965d: 8 (syn: Browne, 1973). Eidophelus opacus (Schedl, 1959a: 477) (Cryphalomorphus) comb. nov. [Scolytogenes].</p> <p>Eidophelus pacificus (Schedl, 1941b: 111) (Ptilopodius) comb. nov. [Ptilopodius].</p> <p>Eidophelus papuanus (Schedl, 1974: 459) (Cryphalomorphus) comb. nov. [Scolytogenes].</p> <p>Eidophelus papuensis (Wood, 1989: 179) (Scolytogenes) comb. nov. [Scolytogenes].</p> <p>= Xylocryptus papuanus Schedl, 1975a: 352 (secondary homonym).</p> <p>Eidophelus paradoxus (Wood, 1992b: 80) (Scolytogenes) comb. nov. [Scolytogenes].</p> <p>= Cryphalomorphus papuanus Schedl, 1979c: 97 (secondary homonym).</p> <p>Eidophelus parvulus Johnson nom. nov. [Cryphalus].</p> <p>= Cryphalus parvus Browne, 1984a: 152 syn. nov. (secondary homonym).</p> <p>Remarks: Cryphalus parvus Browne, 1984a is transferred to Eidophelus based on the cylindrical third tarsal segment, on the entire eye, and on the near-vertical apex of declivity. A replacement name is subsequently proposed due to the name being occupied by Hypothenoides parvus Hopkins, 1915. The replacement name is the diminutive of the adjective parvus (=small), emphasizing that particularly small body size.</p> <p>Eidophelus parvus (Hopkins, 1915: 11) (Hypothenoides) comb. nov. [Scolytogenes].</p> <p>Eidophelus philippinensis (Schedl, 1967a: 126) (Erioschidias) comb.</p> </div>	https://treatment.plazi.org/id/1C598301280BFFA3FC8859B2FCF4FF18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012808FFA2FC885FDBFDE7FECB.text	1C5983012808FFA2FC885FDBFDE7FECB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ernoporus Thomson 1859	<div><p>Ernoporus Thomson, 1859: 147 (Figs. 27–29)</p> <p>Synonymy</p> <p>= Cryphalops Reitter, 1889: 94.</p> <p>= Stephanorhopalus Hopkins, 1915: 35.</p> <p>= Euptilius Schedl, 1940c: 589.</p> <p>= Ernocladius Wood, 1980: 93 syn. nov.</p> <p>= Allothenemus Bright and Torres, 2006: 400 syn. nov.</p> <p>Type of genus</p> <p>Apate tiliae Panzer, 1793.</p> <p>Diagnosis</p> <p>This genus can be diagnosed by the combination of the entire eye, by the pronotum with concentric rows of asperities, by the distinct summit, and by the proventriculus which has anteriorly pointing spines posterior to the masticatory brush.</p></div> 	https://treatment.plazi.org/id/1C5983012808FFA2FC885FDBFDE7FECB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012805FFAEFF285993FEA7FE0E.text	1C5983012805FFAEFF285993FEA7FE0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemicryphalus Schedl 1963	<div><p>Hemicryphalus Schedl, 1963b: 264 (Fig. 30)</p> <p>Type of genus</p> <p>Eidophelus argutus Wood, 1960.</p> <p>Diagnosis</p> <p>Hemicryphalus can be diagnosed by the broadly emarginated, slightly tapered eye shape, by the straight or weakly procurved sutures on the antennal club, by the lateral margin of the pronotum which has a fine carina, and by the proventriculus which has clusters of long, giant spines.</p> </div>	https://treatment.plazi.org/id/1C5983012805FFAEFF285993FEA7FE0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012805FFAEFC885993FAF7FE11.text	1C5983012805FFAEFC885993FAF7FE11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Procryphalus Hopkins 1915	<div><p>Procryphalus Hopkins, 1915: 33 (Fig. 31)</p> <p>Type of genus</p> <p>Procryphalus populi Hopkins, 1915.</p> <p>Diagnosis</p> <p>Distinguished from other Ernoporini and former Cryphalini by the rounded lateral margins of the pronotum, by the eye oval, sometimes weakly emarginated, with the top half wider than the lower half, and by the proventriculus with clusters of giant spines.</p> </div>	https://treatment.plazi.org/id/1C5983012805FFAEFC885993FAF7FE11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012800FFABFF285B9DFD5BFC71.text	1C5983012800FFABFF285B9DFD5BFC71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acorthylus Brethes 1922	<div><p>Acorthylus Brèthes, 1922: 304 (Fig. 32)</p> <p>Synonymy</p> <p>= Phacrylus Schedl, 1938d: 24.</p> <p>Type of genus</p> <p>Acorthylus asperatus Brèthes, 1922.</p> <p>Diagnosis</p> <p>This genus can be diagnosed from all other Scolytinae by the unique second funicle segment which is much larger than the first.</p> </div>	https://treatment.plazi.org/id/1C5983012800FFABFF285B9DFD5BFC71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012800FFABFF285992FD49FEF4.text	1C5983012800FFABFF285992FD49FEF4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Corthylini LeConte 1876	<div><p>Corthylini LeConte, 1876</p> <p>Type of tribe</p> <p>Corthylus Erichson, 1836.</p> <p>Distribution</p> <p>Most genera and species are only known from the Americas.</p></div> 	https://treatment.plazi.org/id/1C5983012800FFABFF285992FD49FEF4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012800FFABFC885864FBEFFCB5.text	1C5983012800FFABFC885864FBEFFCB5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptocarenus Eggers 1937	<div><p>Cryptocarenus Eggers, 1937: 79 (Fig. 33)</p> <p>Synonymy</p> <p>= Cryptocarenus Eggers, 1933a: 10 (unavailable name). = Tachyderes Blackman, 1943: 35.</p> <p>Type of genus</p> <p>Cryptocarenus diadematus Eggers, 1937.</p> <p>Diagnosis</p> <p>Cryptocarenus can be diagnosed from other Scolytinae by the combination of a broadly emarginated eye, metatibiae with spines only on the distal end (as in Hypothenemus), and an oblique groove instead of a spine or callus below the base of the elytra, and by the dwarfed, flightless males.</p> </div>	https://treatment.plazi.org/id/1C5983012800FFABFC885864FBEFFCB5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287FFFD4FF285BE1FDF4FC7D.text	1C598301287FFFD4FF285BE1FDF4FC7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neocryphus Nunberg 1956	<div><p>Neocryphus Nunberg, 1956a: 139 (Fig. 34)</p> <p>Type of genus</p> <p>Neocryphus argentinensis Nunberg, 1956.</p> <p>Diagnosis</p> <p>This genus can be diagnosed by the combination of a broadly emarginated eye, antennae with a second funicle segment smaller than the first, and a large antennal club.</p></div> 	https://treatment.plazi.org/id/1C598301287FFFD4FF285BE1FDF4FC7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287FFFD4FC885B09FB09FC80.text	1C598301287FFFD4FC885B09FB09FC80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stegomerus Wood 1967	<div><p>Stegomerus Wood, 1967: 129 (Fig. 35)</p> <p>Type of genus</p> <p>Stegomerus vulgaris Wood, 1967.</p> <p>Diagnosis</p> <p>Stegomerus can be diagnosed by the short, almost spherical antennal scape, the antennal club, which is mostly symmetrical and with one or two visible procurved sutures, by the rounded lateral margin of the pronotum with bifurcating setae on the hypomeron, and the broadly divided apical plate of the proventriculus.</p> </div>	https://treatment.plazi.org/id/1C598301287FFFD4FC885B09FB09FC80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287CFFD7FF285F15FE00F946.text	1C598301287CFFD7FF285F15FE00F946.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stegomerus longipennis Wood 2007	<div><p>Stegomerus longipennis Wood, 2007: 482.</p> <p>Stegomerus mexicanus Wood, 1967 133.</p> <p>Stegomerus mirandus Wood, 1971: 32.</p> <p>Stegomerus montanus Wood, 1967: 132.</p> <p>Stegomerus parvatis (Wood, 1974a: 17) (Cryphalomorphus) comb. nov. [Scolytogenes].</p> <p>Stegomerus pygmaeus Wood, 1967: 130.</p> <p>Stegomerus vulgaris Wood, 1967: 134.</p> </div>	https://treatment.plazi.org/id/1C598301287CFFD7FF285F15FE00F946	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287CFFD7FF285E22FA83F92E.text	1C598301287CFFD7FF285E22FA83F92E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trypolepis Bright 2019	<div><p>Trypolepis Bright, 2019: 182</p> <p>Type of genus</p> <p>Trypolepis antillica Bright, 2019.</p> <p>Diagnosis</p> <p>This genus can be diagnosed from the similar Acorthylus and Neocryphus by the antenna, which has only two funicle segments, with the first (pedicel) of a similar size to the second, and from the superficially similar Ernoporus by the broadly emarginated eye.</p> </div>	https://treatment.plazi.org/id/1C598301287CFFD7FF285E22FA83F92E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287BFFD0FF2858D5FDBAFBA4.text	1C598301287BFFD0FF2858D5FDBAFBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stephanopodius Schedl 1963	<div><p>Stephanopodius Schedl, 1963a: 633 (Fig. 36)</p> <p>Synonymy</p> <p>= Stephanopodius Schedl, 1941a: 396 (unavailable name). = Cryphalomimus Browne, 1962b: 75 (permanently invalid primary homonym).</p> <p>= Cryphalomimetes Browne, 1963a: 242.</p> <p>= Cryphalmomimetes Wood, 1980: 95 (unavailable name).</p> <p>Type of genus</p> <p>Stephanoderes dispar Eggers, 1936.</p> <p>Diagnosis</p> <p>Stephanopodius can be distinguished by the pronotum strongly arched, laterally inflated and rounded, with very rough asperities near the summit, by the antennal club nearly segmented, with two or three distinct sutures or grooves marked by thick, plumose setae, and by the metatibiae with lateral edge angulate and denticles only on apical margin, and by the bilobed third tarsi.</p> </div>	https://treatment.plazi.org/id/1C598301287BFFD0FF2858D5FDBAFBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287BFFD0FF285993FE2BFEFE.text	1C598301287BFFD0FF285993FE2BFEFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xyloctonini Eichhoff 1878	<div><p>Xyloctonini Eichhoff, 1878b</p> <p>Type of tribe</p> <p>Xyloctonus Eichhoff, 1872a.</p> <p>Distribution</p> <p>Africa, Southeast Asia and Oceania.</p></div> 	https://treatment.plazi.org/id/1C598301287BFFD0FF285993FE2BFEFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287BFFD3FC885B55FA57FB56.text	1C598301287BFFD3FC885B55FA57FB56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryphalini , sensu Wood 1986	<div><p>Key to Genera of Former Cryphalini and Xyloterini</p> <p>This key is made with the diagnostic characters to enable identification of the tribes Coriacephilini, Cryphalini, Trypophloini, Xyloteini, or Ernoporini. Corthylini genera are not included since a thorough review of the tribe would be needed to accurately place the genera in a key. Atomothenemus Bright, 2019, Microsomus Bright, 2019, and Pygmaeoborus Bright, 2019, all from the West Indies, were described during the final preparation of the manuscript and not included in this key.</p> <p>1a. Eye emarginated (with exceptions for some specimens &lt;1.0 mm) or divided. Lower part of elytral declivity usually sloping. Mesocoxae usually narrowly separated similar to metacoxae, or contiguous. Postnotum separated from metathorax by a continuous membrane. Aedeagus with separated (rarely weakly fused) penis apodemes, typically as long as or longer than penis body....................................................... 2</p> <p>1b. Eye oval (with exceptions of specimens with proportionally large eyes, and Procryphalus and Hemicryphalus, have long, broadly emarginated eyes). Lower part of elytral declivity usually vertical. Mesocoxae usually more widely separated than metacoxae. Postnotum fused to metathorax. Aedeagus with fused penis apodemes that are much shorter than penis body. (Ernoporini)............................................................. 12</p> <p>2a. Eye completely divided. Females with a visible mycangium on the hypomeron. Males sometimes with a barely declivous pronotum or flattened pronotal margin. (Xyloterini)......... 10</p> <p>2b. Eye emarginated (rarely oval). No mycangium on hypomeron of females............................................................................ 3</p> <p>3a. Third tarsal segment bilobed. Lateral margins of pronotum always with carina along dorsolateral margin. Split setae on hypomeron usually present. Interstrial ground vestiture usually dense. Antennae always with sutures (which can be recurved or procurved). Body generally rounded. (Cryphalini)............................................................................. Cryphalus</p> <p>3b. Third tarsal segment cylindrical. Lateral margin of pronotum with or without carina along dorsolateral margin.Hypomeron without, or rarely with very few bifurcating setae. Interstrial ground vestiture usually sparse, especially on elytral disc. Antennae with or without sutures. Body usually more elongated.................................................................................... 4</p> <p>4a. Eye deeply emarginated. Antennal club with procurved sutures and a complete septum along the first suture. Lateral margins of pronotum rounded or with weak carina and hypomeron with bifurcating setae. Interstrial ground vestiture hair-like. Distributed only in Asia (rarely collected). (Coriacephilini)............................................... Coriacephilus</p> <p>4b. Not in combination as described above. (Trypophloeini)..... 5</p> <p>5a. Lateral margins of pronotum always rounded. Antennal club cone-like with horizontal sutures and elongate, with a pointed apex. Distributed in temperate climates in hosts including Salix, Populus or Alnus. Proventriculus with short apical plate, less than one fifth of the sclerotized part.................................................................... Trypophloeus</p> <p>5b. Lateral margins of pronotum rounded or with carina. Antennal club flattened, with or without sutures, usually transverse or procurved, apex rounded. Various hosts, widespread distribution, in tropical to temperate areas. Proventriculus with apical plate much more than one fifth of total length (except not sclerotized in Macrocryphalus)....... 6</p> <p>6a. Lateral margins of pronotum rounded. Antennal funicle always with five segments. Antennal club shape tapered proximally, with a weakly visible procurved suture. Elytra covered in hair-like interstrial ground vestiture. Striae not apparent. Proventriculus not sclerotized...................... Macrocryphalus</p> <p>6b. Lateral margins of pronotum with carina. Antennal funicle with 2−5 segments. Antennal club shape and sutures variable. Elytra interstrial ground vestiture variable. Striae usually visible (except some Afrocosmoderes). Proventriculus fully sclerotized.................................................................................. 7</p> <p>7a. Mesocoxae contiguous. Body shape elongate, more than 2.3x as long as wide. Antennae always with 2−3 funicular segments, with the first segment longer than the subsequent funicle segments combined. Antennal club flat and without sutures............................................................... Cosmoderes</p> <p>7b. Mesocoxae narrowly separated. Body shape stout to elongate. Antennae with 3−5 funicle segments. Antennal club flat, with or without sutures or septum............................................... 8</p> <p>8a. Antennal club without sutures or weakly visible procurved sutures. Metatibia with denticles covering at least apical quarter, pointing outwards and distally....... Afrocosmoderes</p> <p>8b. Antennal club usually with straight or slightly procurved sutures and a partial septum (rare exceptions are known). Metatibia with denticles restricted to apex, usually pointed distally................................................................................. 9</p> <p>9a. Antennal club with three sutures and usually a partial septum. Pronotum with marginal asperities. Males smaller, with vestigial wings, and with eyes much smaller than females...................................................................... Hypothenemus</p> <p>9b. Antennal club with one transverse suture and one barely visible procurved suture near apex. Pronotum with no marginal asperities. Males of a similar size, with fully developed wings and with eyes similar to females............... Microcosmoderes</p> <p>10a. Antennal suture (indicated by differences in vestiture) acutely procurved. Males with distinctly flattened frons and dorsally subquadrate to quadrate pronotum. Tegmen with median apodeme........................................................ Trypodendron</p> <p>10b. Antennal suture (indicated by differences in vestiture) broadly procurved. Males with flat or convex frons, and pronotum dorsally rounded. Tegmen without median apodeme......... 11</p> <p>11a. Antennal club uniformly pubescent to base. Male and female of a similar size. Females with longest axis of mycangia either vertical (dorso-ventrally) or horizontal (antero-posteriorly)....................................................................... Indocryphalus</p> <p>11b. Antennal club with basal area corneous. Male distinctly smaller than female. Females with longest axis of mycangia horizontal (antero-posteriorly)............................ Xyloterinus</p> <p>12a. Stout, less than 2.2x as long as wide. Pronotum with distinct summit, asperities in irregular concentric rows. Lateral margins of pronotum rounded, at most with a small carina along the posterior edge. Setae on hypomeron divided in some species. Proventriculus crop with thin spines, no enlarged crop spines or multidentate spines. Proventriculus posterior to the masticatory brush with a tuft of spatulate setae... Ernoporus</p> <p>12b. Proportions variable. Pronotal summit variable, asperities may occur in irregular concentric rows or scattered. Pronotum with carina along dorsolateral edge (except Procryphalus, which has longer body proportions). Setae on hypomeron always hair-like (if present). Proventriculus crop with enlarged clusters of crop spines or multidentate spines. Proventriculus posterior to the masticatory brush smooth, without a tuft of spatulate setae................................................................... 13</p> <p>13a. Base of pronotum constricted, narrower than the base of the elytra. Antennae with straight sutures, the first suture being completely septate. Dorsolateral margin of pronotum rounded. Proventriculus simple, crop with clusters of very large crop spines............................................... Procryphalus</p> <p>13b. Base of pronotum as broad as the base of elytra, not constricted. Antennal sutures variable, sometimes with partial septum. Dorsolateral margins of pronotum marked with carina. Proventriculus and crop variable............................ 14</p> <p>14a. Eye long and tapered ventrally. Antennal club with three transverse sutures. Symetrical incomplete septum along first suture. Crop spines include several large long sclerotized spines. Distributed only in the Pacific islands....................... Hemicryphalus</p> <p>14b. Eye usually elongate-oval shaped. Antennal club flattened, sutures often procurved or absent, oblique partial septum sometimes present. Proventricular crop spines well-developed as short, socketed spines. Widespread in Europe, Africa, Asia and Oceania, rare in tropical and subtropical Americas............................................................................ Eidophelus</p></div> 	https://treatment.plazi.org/id/1C598301287BFFD3FC885B55FA57FB56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johnson, Andrew J.;Hulcr, Jiri;Knížek, Miloš;Atkinson, Thomas H.;Mandelshtam, Michail Yu.;Smith, Sarah M.;Cognato, Anthony I.;Park, Sangwook;Li, You;Jordal, Bjarte H.	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
