taxonID	type	description	language	source
1C598301283DFF91FC885B94FF31FC13.taxon	description	The first mention of Cryphalini as a group name was by Lindemann in 1877. The group-concept was much broader, including Pityophthorus Eichhoff, 1864, Hypoborus Erichson, 1836, Hylocurus Eichhoff, 1872, and Xyloctonus Eichhoff, 1872, all of which are currently placed in separate tribes. Eichhoff described many of the contemporary genera, plus several which are now junior synonyms (Eichhoff 1878 a). Most contemporary Cryphalini genera were classified in ‘ Cryphalidae’ which was a much narrower concept than previously described, containing the known Cryphalini genera (except Eidophelus Eichhoff, 1876) plus Hypoborus and Liparthrum Wollaston, 1854 (Hypoborini) and what is now the genus Pityoborus Blackman, 1922 (Corthylini). Other groups, such as contemporary Micracidini LeConte, 1876 and Xyloctonini Eichhoff, 1878, were explicitly described as different tribes. This approximate group concept of Cryphalini remained until Hopkins presented a major reclassification of Scolytinae in 1915, with a group ‘ Cryphalinae’ in an even broader sense including the contemporary tribes Xyleborini and some genera in the current Dryocoetini Lindemann, 1877 and Corthylini LeConte, 1876. Most of the contemporary Cryphalini are contained in Hopkins ‘ Division I, Subdivision A’, a group described based on the type of vestiture (‘ with scales, rarely with hairs’) and the pattern of asperities along the anterior margin of the pronotum. The characters used to describe and diagnose the genera within the Cryphalini include the presence of distinct elytral strial punctures, the shape of the lateral margins of the pronotum, the degree to which the pronotum was produced, the apex of the elytra, and most importantly, the morphology of the antennae. Eidophelus was considered in a different subdivision with Dendroterus Blandford, 1904 and some other genera now placed in Dryocoetini. Balachowsky (1949) presented a clarified classification of bark beetles which reflects most contemporary tribes. While restricted to the fauna of France, it clearly described Cryphalini as different from Hypoborini. Wood (1954) evaluated the North American Cryphalini genera and species, representing the contemporary scope of the tribe for the region. Two additional genera from East Asia were described by Nobuchi (1981), inferred partly by new sets of characters from the proventriculus (Nobuchi 1969). While these alone enabled identification of most scolytine genera, they were not widely adopted as a diagnostic set of characters. Schedl was a major contributor to Cryphalini systematics. While very prolific in describing new species, his classification of Cryphalini did not settle any of the considerable confusion and his nomenclatural practices were not consistently valid (see Wood and Bright 1992). Schedl described five genera in the contemporary Cryphalini, but four of these are now regarded as junior synonyms. The most recent treatment of Cryphalini was Wood’s (1986 a) review of all scolytine tribes and genera. In this review, Cryphalini was diagnosed based on the combination of the ascending costal margin of the elytra, by the base of the elytra that is unarmed (or rarely armed with a continuous line, not crenulations), by the declivous (steeply sloped) pronotum usually armed with serrations and asperities, by the concealed head when viewed dorsally, by the eye that is ‘ usually’ entire and ‘ less commonly emarginated’, by the antennal funicle with no more than five segments, by the antennal club which is ‘ strongly flattened’ and never truncate, by the lateral margins of the pro- and mesotibiae armed with denticles, and by the metanepisternum which has a posterior margin concealed by the elytron. Wood and Bright brought significant order to the chaos of the classification of Scolytinae as a whole, including Cryphalini. Many genera were synonymized, and a small number was described. Their compilation of the catalogs (Wood and Bright 1987; Wood and Bright, 1992) is a milestone in scolytine systematics and provides a very useful starting point for taxonomic reviews. Allothenemus Bright and Torres, 2006 was subsequently described. During the final stages of this manuscript preparation, four further genera were described in Cryphalini: Atomothenemus Bright, 2019, Microsomus Bright, 2019, Pygmaeoborus Bright, 2019, and Trypolepis Bright, 2019.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012835FF9EFC885F65FADAF8E1.taxon	materials_examined	Type of tribe Coriacephilus Schedl, 1939.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012835FF9EFC885F65FADAF8E1.taxon	diagnosis	Diagnosis Tarsi with third segment cylindrical. Eye deeply emarginated. Antennal funicle with 3 – 5 segments. Antennal club with sutures and a complete septum. Hypomeron with bifurcating setae.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012832FF98FC885E2AFD6CF93B.taxon	type_taxon	Type of genus Stephanoderes coriaceus Eichhoff, 1878. Diagnosis Coriacephilus can be separated from other Cryphalini by the combination of cylindrical third tarsal segments, deeply emarginated eye, antennae with sutures and septum, and bifurcating setae on the hypomeron.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012831FF9AFF285C04FD1FF953.taxon	description	= Cryphalidae Eichhoff, 1878 b: 107.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012831FF9AFF285C04FD1FF953.taxon	materials_examined	Type of tribe Cryphalus Erichson, 1836.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012831FF9AFF285C04FD1FF953.taxon	diagnosis	Diagnosis Tarsi with third segment bilobed. Eye emarginated. Antennal funicle with 3 – 5 segments. Antennal club with sutures and no septum. Hypomeron with bifurcating setae. Postnotum not fused with metanotum. Aedeagus with long penis apodemes, sometimes fused at apex, with a complete tegmen. Tegmen with pair of apodemes.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012831FF9AFC885888FC62FB1F.taxon	type_taxon	Type of genus Bostrichus asperatus Gyllenhal, 1813.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012831FF9AFC885888FC62FB1F.taxon	diagnosis	Diagnosis Cryphalus can be distinguished from similar genera by the combination of emarginated eyes, sutures on the antennae clearly visible but without a septum, by the weakly bilobed third tarsal segments, the proventriculus with a large apical plate, and the aedeagus that has a tegmen sclerotized completing a ring, usually with two tegminal apodemes.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301282AFF83FF285994FCF4FF18.taxon	discussion	Remarks: No formal synonymy has been proposed for this species name. Wood (1969) indicated that they are probably the same, and Wood and Bright (1992) listed the synonyms as above, choosing Cryphalus dorsalis as the valid name with dorsalis getting priority chosen by the first reviser. The nomenclature is being treated as valid, but the subjective synonymy is uncertain (see Mandelshtam and Nikitsky 2005).	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301282AFF83FF285994FCF4FF18.taxon	discussion	Remarks: A replacement name is proposed for Cryphalus confusus Eggers, 1927 b, because the combination is already occupied following the transfer of Margadillius confusus Hopkins, 1915 to Cryphalus. The replacement name is a noun in the genitive case, honoring the original author.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301282AFF83FF285994FCF4FF18.taxon	discussion	Remarks: The original and subsequent spelling is erraticum as an adjective, which should be the masculine erraticus.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301282AFF83FF285994FCF4FF18.taxon	discussion	Remarks: A replacement name is proposed for Cryphalus cylindrus Browne, 1984 d following the transfer of Dacryphalus cylindrus Browne, 1950 from Hypocryphalus. The replacement name is an adjective, meaning either brown or indistinct.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301282AFF83FF285994FCF4FF18.taxon	discussion	Remarks: A replacement name is proposed for Cryphalus laevis Browne, 1984 b because the name is occupied by Hypocryphalus laevis Browne, 1980 a. The replacement name is an adjective meaning slender or graceful.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301282AFF83FF285994FCF4FF18.taxon	discussion	Remarks: A replacement name is proposed for Margadillius fulvus Browne, 1984 b after its transfer to Cryphalus, because the name is already occupied by Cryphalus fulvus Niisima, 1908. The name is an adjective, meaning either brown or yellow, maintaining the original meaning.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301282AFF83FF285994FCF4FF18.taxon	discussion	Remarks: A replacement name is proposed for Hypocryphalus minutus Browne, 1980 c following its transfer to Cryphalus. The name is occupied by Margadillius minutus Hopkins, 1915. The replacement name is an adjective, and maintains the original meaning, referencing the very small size.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012828FF83FC885994FB64F815.taxon	discussion	Remarks: A replacement name is proposed for Cryphalus striatulus Browne, 1981 because the name is occupied, following the revised status of Cryphalomorphus striatulus Browne, 1978. The new is formed from the diminutive of the adjective punctatus meaning punctured, and s striatus meaning with lines or rows.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012828FF83FC885994FB64F815.taxon	discussion	Remarks: A replacement name is proposed for Hypocryphalus formosanus Schedl, 1952 b because the name is occupied by Cryphalus formosanus Schedl, 1942 a. The replacement name is a noun in the genitive case, honoring the original author. The types of both species were unavailable for study.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012829FF82FF285994FEC5F839.taxon	discussion	Remarks: A replacement name is proposed for Margadillius terminaliae Browne, 1984 c because the name is already occupied by Cryphalus terminaliae Browne, 1980 b. The replacement name is a toponym derived from the type locality.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012829FF82FF285994FEC5F839.taxon	discussion	Remarks: A replacement name is proposed for Cryphalus densepilosus Schedl, 1943 because the name is now occupied by Hypocryphalus densepilosus Schedl, 1942 d. The replacement name is formed from the adverb spisse (= densely) and the adjective pilosus (= hairy).	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012829FF82FF285994FEC5F839.taxon	discussion	Remarks: A replacement name is proposed for Cryphalus striatus Browne, 1974 because the name is now occupied by Hypocryphalus striatus Hopkins, 1915. The replacement name is a noun in the genitive case, named after the host plant Storckiella.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012829FF82FF285994FEC5F839.taxon	discussion	Remarks: Cryphalus striatulus (Browne, 1978) is transferred to Cryphalus based on the external diagnostic characters visible. The replacement name is no longer necessary.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012829FF82FC885994FB4DF999.taxon	discussion	Remarks: A replacement name is proposed for Euptilius exiguus Browne, 1984 c because the name is occupied by Cryphalus exiguus Blandford, 1894 b. The replacement name is a noun in the genitive case, honoring the original collector, K. Takahashi.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012829FF8DFC885F5EFE78FF53.taxon	materials_examined	Type of tribe Trypophloeus Fairmaire, 1864.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012829FF8DFC885F5EFE78FF53.taxon	diagnosis	Diagnosis Tarsi with third segment cylindrical. Eye emarginated. Antennal funicle with 3 – 5 segments. Antennal club with or without sutures, and at most a partial septum. Hypomeron with single setae, or rarely mixed with a few bifurcating setae. Postnotum not fused with metanotum. Typically with scale-like interstrial bristles, and sparse interstrial ground vestiture.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012824FF8FFF285D2FFE6DFB46.taxon	type_taxon	Type of genus Atomothenemus unicus Bright, 2019.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012824FF8FFF285D2FFE6DFB46.taxon	diagnosis	Diagnosis Antennae with two funicle segments (including the pedicel), antennal club with horizontal sutures.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012824FF8FFC8858D7FAEFFBFC.taxon	type_taxon	Type of genus Cosmoderes monilicollis Eichhoff, 1878 a.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012824FF8FFC8858D7FAEFFBFC.taxon	diagnosis	Diagnosis Body shape elongated, antennae with short funicle, antennal club large and without sutures, marginal asperities present, mesocoxae contiguous, anterior plate of proventriculus short.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301281AFFB0FF285994FB30F838.taxon	discussion	Remarks: A replacement name is proposed for Periocryphalus sobrinus Wood, 1974 following the generic synonymy of Hypothenemus and Periocryphalus, creating conflict with Stephanoderes sobrinus Schedl, 1965 a. The replacement name is an adjective, referencing the prominent posterolateral margin of the elytra. The holotype of Periocryphalus sobrinus Wood, 1974 at BMNH is lost and erroneously replaced with a Cnesinus sp., which clearly does not match the species description. A paratype at USNM collected from the same location on a different date matches the original and subsequent descriptions of the species. (Labeled ‘ PARATYPE // Periocryphalus // sobrinus // S. L. Wood 1972 //// Periocryphalus // sobrinus //. 72 Wood // S. L. Wood // RS / RGS Exp. Brazil // 12 ′ 31 ′ S 51 ′ 46 ′ W //// R. A. Beaver F 03 // 7. XI. 1968 //// S. L. Wood Collection //// USNMENT 01448187 ’.) Since this species is the type of the genus Periocryphalus, the stability of the name-bearing type is critical. Therefore, the single remaining specimen in the type series at USNM is designated as a neotype of Periocryphalus sobrinus Wood, 1974 a, and subsequently, the neotype of Hypothenemus marginatus Johnson.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301281AFFB0FF285994FB30F838.taxon	discussion	Remarks: Bright (2019) noted that Hypothenemus seriatus (Eichhoff, 1872) was a junior synonym of H. obscurus (Fabricius, 1801), as it was synonymized by Wood (1954), based on a specimen compared with the type by Eggers, but the synonymy was largely ignored (including by Wood). We consider the synonymy originally proposed by Wood to be rejected. Hypothenemus obscurus is a highly damaging pest which is not present in some areas where H. seriatus is present. We found that the species were sister in the phylogeny. The morphological differences have been well documented (Vega et al., 2015), and they are genetically distinct (Mitchell and Maddox 2010, Johnson et al 2018). It is possible that there is much more variation than documented which may make definitive identification by morphology difficult or impossible, but more evidence would be needed to justify grouping the distinct morphotypes as one species, as well as alternative recognition of the pestiferous beetles which devastate the macadamia and other tropical nut industries. The economic and policy implications of the synonymy could hinder the necessary biosecurity measures to prevent the pest species establishing in other areas.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301281AFFB0FF285994FB30F838.taxon	discussion	Remarks: A replacement name is proposed for Ptilopodius squamosus Schedl, 1953 b after transferring this species from Ptilopodius, since the combination is already occupied by Stephanoderes squamosus Hopkins, 1915. The new name is the diminutive of the adjective squamosus (= scaly), referring to the scalelike setae on the pronotum and elytra.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012818FFB3FF285994FDD1FB8D.taxon	diagnosis	Diagnosis The teardrop-shaped antennae, rounded lateral margins of the pronotum, separated mesocoxae, and absence of a sclerotized proventriculus diagnose members of this genus.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012816FFBDFF285CBAFE97F9C0.taxon	diagnosis	Diagnosis This genus can be distinguished from other Trypophloeini by the antennae, with a single, procurved suture, and by the lack of any vestiture on the elytra.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012816FFBDFF285C6FFEE4FA1B.taxon	type_taxon	Type of genus	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012817FFBCFF2859F0FE29FE70.taxon	type_taxon	Type of genus Pygmaeoborus cubensis Bright, 2019.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012817FFBCFF2859F0FE29FE70.taxon	diagnosis	Diagnosis Similar to Hypothenemus except antennae with single suture indicated by dense setae on apical third.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012817FFBCFF285C6BFDE5F90D.taxon	type_taxon	Type of genus Bostrichus binodulus Ratzeburg, 1837.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012817FFBCFF285C6BFDE5F90D.taxon	diagnosis	Diagnosis This genus can be diagnosed by the emarginated eye, by the coneshaped antennal club with slightly recurved sutures, and by the rounded lateral margins of the pronotum.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012815FFBEFF285B6CFDEDFBD5.taxon	description	= Xyloteroideae Lindemann, 1877: 165. = Xyloteridae Eichhoff, 1878 b: 411. = Trypodendrinae Trédl, 1907: 70 (Unavailable name). = Xyloterinae Nüsslin, 1911 a: 432.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012815FFBEFF285B6CFDEDFBD5.taxon	description	= Trypodendrina Nunberg, 1954: 16. = Trypodendrini Browne, 1961: 45. = Trypodendroninae Wood, 1978: 114.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012815FFBEFF285B6CFDEDFBD5.taxon	materials_examined	Type of tribe Xyloterus Erichson, 1836.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012815FFBEFF285B6CFDEDFBD5.taxon	diagnosis	Diagnosis Eye divided. Mycangia present on females.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012813FFB8FF285829FDD7FD58.taxon	type_taxon	Type of genus Dermestes domesticus Linnaeus, 1758.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012813FFB8FF285829FDD7FD58.taxon	diagnosis	Diagnosis Antennal club with a single acutely procurved suture. Mycangia on hypomeron horizontal (antero-posteriorly).	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012813FFB8FC885A37FAE2FBB5.taxon	type_taxon	Type of genus Bostrichus politus Say, 1826.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012813FFB8FC885A37FAE2FBB5.taxon	diagnosis	Diagnosis Antennal club with a single weakly procurved suture. Mycangia on hypomeron horizontal (antero-posteriorly elongate).	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012813FFBBFC885FE1FED1FEB1.taxon	description	= Eidophelinae Murayama, 1954: 200 (Unavailable name). = Eidopherinae Murayama, 1954: 200.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012813FFBBFC885FE1FED1FEB1.taxon	materials_examined	Type of tribe Ernoporus Thomson, 1859.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012813FFBBFC885FE1FED1FEB1.taxon	diagnosis	Diagnosis Eye entire, rarely broadly emarginated. Antennae with three to five funicle segments. Postnotum fused with metanotum. Penis apodemes short and fused.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012810FFBBFF285D66FB20FE0C.taxon	type_taxon	Type of genus Eidophelus imitans Eichhoff, 1876	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012810FFBBFF285D66FB20FE0C.taxon	diagnosis	Diagnosis Eidophelus can be diagnosed from superficially similar genera by the entire eye, by the lateral and basal margins of the pronotum marked by a carina, and by the antennal club which has a partial or absent septum, and by the proventriculus characters.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301280BFFA3FC8859B2FCF4FF18.taxon	discussion	Remarks: A replacement name is proposed for Eidophelus minutissimus Schedl, 1962 a because the name is now occupied by Ptilopodius minutissimus Schedl, 1943. The replacement name is an adjective, and maintains the original meaning, referencing the very small size.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301280BFFA3FC8859B2FCF4FF18.taxon	discussion	Remarks: A replacement name for Ernoporicus ater Nobuchi, 1975 is proposed because the name is already occupied by Negritus ater Eggers, 1923 a. The name is an adjective, following the meaning of the original name, meaning black.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301280BFFA3FC8859B2FCF4FF18.taxon	discussion	Remarks: Cryphalus parvus Browne, 1984 a is transferred to Eidophelus based on the cylindrical third tarsal segment, on the entire eye, and on the near-vertical apex of declivity. A replacement name is subsequently proposed due to the name being occupied by Hypothenoides parvus Hopkins, 1915. The replacement name is the diminutive of the adjective parvus (= small), emphasizing that particularly small body size.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012808FFA2FC885FDBFDE7FECB.taxon	type_taxon	Type of genus Apate tiliae Panzer, 1793.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012808FFA2FC885FDBFDE7FECB.taxon	diagnosis	Diagnosis This genus can be diagnosed by the combination of the entire eye, by the pronotum with concentric rows of asperities, by the distinct summit, and by the proventriculus which has anteriorly pointing spines posterior to the masticatory brush.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012805FFAEFF285993FEA7FE0E.taxon	type_taxon	Type of genus Eidophelus argutus Wood, 1960.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012805FFAEFF285993FEA7FE0E.taxon	diagnosis	Diagnosis Hemicryphalus can be diagnosed by the broadly emarginated, slightly tapered eye shape, by the straight or weakly procurved sutures on the antennal club, by the lateral margin of the pronotum which has a fine carina, and by the proventriculus which has clusters of long, giant spines.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012805FFAEFC885993FAF7FE11.taxon	type_taxon	Type of genus Procryphalus populi Hopkins, 1915.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012805FFAEFC885993FAF7FE11.taxon	diagnosis	Diagnosis Distinguished from other Ernoporini and former Cryphalini by the rounded lateral margins of the pronotum, by the eye oval, sometimes weakly emarginated, with the top half wider than the lower half, and by the proventriculus with clusters of giant spines.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012800FFABFF285B9DFD5BFC71.taxon	type_taxon	Type of genus Acorthylus asperatus Brèthes, 1922.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012800FFABFF285B9DFD5BFC71.taxon	diagnosis	Diagnosis This genus can be diagnosed from all other Scolytinae by the unique second funicle segment which is much larger than the first.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012800FFABFF285992FD49FEF4.taxon	materials_examined	Type of tribe Corthylus Erichson, 1836.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012800FFABFF285992FD49FEF4.taxon	distribution	Distribution Most genera and species are only known from the Americas.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012800FFABFC885864FBEFFCB5.taxon	type_taxon	Type of genus Cryptocarenus diadematus Eggers, 1937.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C5983012800FFABFC885864FBEFFCB5.taxon	diagnosis	Diagnosis Cryptocarenus can be diagnosed from other Scolytinae by the combination of a broadly emarginated eye, metatibiae with spines only on the distal end (as in Hypothenemus), and an oblique groove instead of a spine or callus below the base of the elytra, and by the dwarfed, flightless males.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287FFFD4FF285BE1FDF4FC7D.taxon	type_taxon	Type of genus Neocryphus argentinensis Nunberg, 1956.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287FFFD4FF285BE1FDF4FC7D.taxon	diagnosis	Diagnosis This genus can be diagnosed by the combination of a broadly emarginated eye, antennae with a second funicle segment smaller than the first, and a large antennal club.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287FFFD4FC885B09FB09FC80.taxon	type_taxon	Type of genus Stegomerus vulgaris Wood, 1967.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287FFFD4FC885B09FB09FC80.taxon	diagnosis	Diagnosis Stegomerus can be diagnosed by the short, almost spherical antennal scape, the antennal club, which is mostly symmetrical and with one or two visible procurved sutures, by the rounded lateral margin of the pronotum with bifurcating setae on the hypomeron, and the broadly divided apical plate of the proventriculus.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287CFFD7FF285E22FA83F92E.taxon	type_taxon	Type of genus Trypolepis antillica Bright, 2019.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287CFFD7FF285E22FA83F92E.taxon	diagnosis	Diagnosis This genus can be diagnosed from the similar Acorthylus and Neocryphus by the antenna, which has only two funicle segments, with the first (pedicel) of a similar size to the second, and from the superficially similar Ernoporus by the broadly emarginated eye.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287BFFD0FF2858D5FDBAFBA4.taxon	type_taxon	Type of genus Stephanoderes dispar Eggers, 1936.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287BFFD0FF2858D5FDBAFBA4.taxon	diagnosis	Diagnosis Stephanopodius can be distinguished by the pronotum strongly arched, laterally inflated and rounded, with very rough asperities near the summit, by the antennal club nearly segmented, with two or three distinct sutures or grooves marked by thick, plumose setae, and by the metatibiae with lateral edge angulate and denticles only on apical margin, and by the bilobed third tarsi.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287BFFD0FF285993FE2BFEFE.taxon	materials_examined	Type of tribe Xyloctonus Eichhoff, 1872 a.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
1C598301287BFFD0FF285993FE2BFEFE.taxon	distribution	Distribution Africa, Southeast Asia and Oceania.	en	Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You, Jordal, Bjarte H. (2020): Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae). Insect Systematics and Diversity 4 (3), No. 1: 1-81, DOI: 10.1093/isd/ixaa002, URL: http://dx.doi.org/10.1093/isd/ixaa002
