taxonID	type	description	language	source
231E878BFFD5FFF0D5DA6255EACDF8D4.taxon	discussion	See Herman (2001) for more synonymies and references.	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD5FFF0D5DA6255EACDF8D4.taxon	diagnosis	Diagnosis. Maxillary palpomere 4 aciculate; protrochantin exposed; epipleural ridge of elytra absent; longitudinal row of tibial spines absent; tarsal formula 2 - 2 - 2; basolateral ridges on tergites II – VII present (Herman 1970).	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD5FFF0D5DA6255EACDF8D4.taxon	biology_ecology	Habitats. They occur on banks of rivers, streams, ponds and other freshwater, but a few are found in intertidal mudflats or fine gravels (Frank & Ahn 2011).	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD7FFF2D5DA65FDEA37FE06.taxon	description	(Figs. 1 A, 2 – 3) Description. Length 1.04 – 1.30 mm. Body flattened, surface pubescent; dark brown. Head. Almost as long as wide and slightly narrower than pronotum; widest across eyes; clypeus trapezoidal; eye moderate in size and rounded in lateral aspect, with large eye facets and interfacetal setae; temple well developed, about 0.8 times as long as eye length in dorsal view; gular suture fused and broadly divergent in anterior and posterior part; neck well developed, distinctly transversely reticulate sculptures with some short setae (Figs. 2 B – C); antenna slender, moderately elongate, and pubescent, antennomere 1 longest, 4 and 6 shortest, 1 – 2 and 7 – 11 elongate, 3 and 5 slightly elongate, 4 and 6 transverse (Fig. 2 A). Mouthparts. Labrum transverse, semicircula; mandible triangle, apex bifid, with one blunt subapical tooth, prostheca and molar tooth well developed; maxillary palpus with four palpomeres, palpomere 1 transverse and small, 2 slightly dilated to apex, 3 largest, 2 – 3 with dense setae; apical margin of 3 about 3.0 times as wide as basal margin of 4; labial palpus with three elongate palpus, apex of palpomere 1 with two long setae and one pore, apex of 2 with one pore, 3 with few short sensilla at apex; mentum quadrate with one pair of long macrosetae and many setae. Thorax. Pronotum transverse, width 0.24 – 0.27 mm, widest about anterior third, surface pubescent; prosternum with few setae (Fig. 2 D); scutellum an inverted triangle, anterior margin of scutellum emarginate, posterior part reticulate with many setae (Fig. 2 E); elytron elongate and pubescent, posterior margin with membranous lobe; hind wing present; mesoventral process slightly extended and blunted at apex; metaventral carinae incomplete; metaventrite with two pairs of long macrosetae and dense setae (Fig. 2 F); all tibiae with one long macroseta in middle. Abdomen. Posterior margin of tergite VII with fringe of setae; postero-lateral margin of tergite VIII slightly convex. Secondary sexual characteristics. Male sternite VIII emarginate, female with linear form; male tergite IX with ventral structs, struts as long as remainder, female tergite IX without ventral struts. Male sternite IX elongate, female sternite IX pentagon. Genitalia. Aedeagus oval (Figs. 3 A – B); paramere short; spermatheca as in Fig. 3 C.	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD7FFF2D5DA65FDEA37FE06.taxon	materials_examined	Type material. Holotype, 1 ♂, labeled as follows: KOREA: Jeju Prov., Seogwipo-si, Haye-dong, N 33 ° 13 ' 54.45 " E 126 ° 22 ' 19.81 " 2 m, 19 X 2014, KJ Ahn, IS Yoo, JS Lee, 10 - 20 cm deep under / in gravels on seashore. ‘ Holotype, Thinobius jejuensis Lee and Ahn, Desig. Jae-Seok Lee and Kee-Jeong Ahn 2021 ’ ‘ Deposited in the Chungnam National University Insect Collection, Korea’. Paratypes, 3 exx., labeled as same as Holotype (1 on slide; 1 on CNUIC voucher); 30 exx., same data as former except for ‘ N 33 ° 13 ' 54.20 " E 126 ° 22 ' 18.49 " 2 m, 05 VIII 2020, JS Lee, JY Park, YJ Kim, flotation’. 24 exx., same data as former except for ‘ Yeraehaean-Ro, N 33.232254, E 126.371442, 19 X 2020, KJ Ahn, Under pebbles / gravels by flotation’.	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD7FFF2D5DA65FDEA37FE06.taxon	distribution	Distribution. Korea (Jeju island).	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD7FFF2D5DA65FDEA37FE06.taxon	discussion	Remarks. This species is similar to T. marinus, but can distinguished by the length of temple and the shape of antennomeres. Temple is shorter than the diameter of eye in T. jejuensis Lee & Ahn, sp. n. (Fig. 2 B), but longer than the diameter of eye in T. marinus (Cameron, 1917). In addition, antennomeres 5 and 7 – 8 are elongate in the new species (Fig. 2 A), but 4 – 8 are transverse in T. marinus. All specimens were collected on Jeju-do island, Korea	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD6FFF5D5DA607FEE35F8F9.taxon	description	(Figs. 1 B, 4 – 5) Description. Length 1.60 – 1.80 mm. Body flattened, surface pubescent; light brown to dark brown, head and abdomen dark brown, pronotum, elytra and antenna brown, mouthparts and legs light brown. Head. Almost as long as wide and slightly narrower than pronotum; widest across eyes; clypeus trapezoidal; eye moderate in size and rounded in lateral aspect, with large eye facet and interfacetal setae; temple well developed, diameter of eye slightly longer than temple; gular suture fused and broadly divergent at posterior part; neck well developed, transversely reticulate sculptures distinct, seta absent (Fig. 4 B); antenna slender, moderately elongate, and pubescent; antennomere 1 longest, 3 shortest, 1 – 11 elongate (Fig. 4 A). Mouthparts. Labrum transverse, lateral margin rounded, anterior margin slightly emarginate; mandible triangular, apex bifid with subapical tooth, prostheca and molar tooth well developed; maxillary palpus with four palpomeres, palpomere 1 transverse and small, 2 slightly dilated to apex, 3 largest, 2 – 3 with many setae, 4 short and slender; apical margin of 3 more than 2.0 times as wide as basal margin of 4; labial palpus with three elongate palpi, palpomere 1 with one long, one short seta and two pores, 2 with one long seta and one pore, 3 with few short sensilla at apex; mentum quadrate with two pairs of long macrosetae and many setae. Thorax. Pronotum transverse, width 0.29 – 0.30 mm, widest near half of length, surface pubescent with seven pairs of macrosetae and many setae; prosternum with one pair of long setae and a few short setae, apex of prosternal process sharp and elevated (Figs. 4 C – D); scutellum an inverted triangle, anterior margin of scutellum emarginate, posterior part reticulate with a few setae and inverted triangle impression, impression about 1.6 times as wide as long, elytron elongate and pubescent; posterior margin with membranous lobe; hind wing present; mesoventral process slightly extended and pointed at apex; metaventral carina absent; metaventrite distinctly reticulate with two pairs of long macrosetae and many setae (Fig. 4 E); all tibiae with one long macroseta in middle. Abdomen. Posterior margin of tergite VII with fringe of setae; posterior margin of tergite VIII emarginate; posterior margin of sternite VIII convex. Secondary sexual characteristics. Male tergite IX with ventral structs, struts shorter than other part, female gonocoxites present; female with one ring structure (Fig. 4 F). Genitalia. Aedeagus oval (Fig. 5 A) with serrated pattern in lateral aspect (Fig. 5 B); paramere long and slender; spermatheca as in Fig. 5 C.	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD6FFF5D5DA607FEE35F8F9.taxon	materials_examined	Type material. Holotype, 1 ♂, labeled as follows: ‘ KOREA: Gyeongbuk Prov., Pohang-si, Buk-gu, Cheongha-myeon, Cheongjin-ri, N 36 ° 10 ' 38.73 " E 129 ° 23 ' 37.15 " 17 m, 12 VII 2018, IS Yoo, JS Lee, JG Jung, in coarse sand / gravels on seashore, flotation’ ‘ Holotype, Thinobius koreanus Lee and Yoo, Desig. Jae-Seok Lee and In-Seong Yoo, 2021 ’ ‘ Deposited in the Chungnam National University Insect Collection, Korea’. Paratypes, 7 exx., same data as holotype, 5 exx., same data as holotype except for ‘ N 36 ° 10 ʹ 37.8 ʺE 129 ° 23 ʹ 38.0 ʺ 9 m, 27 VIII 2010, KJ Ahn, TK Kim, YH Kim, IS Yoo, JH Song, SG Lee, JH Jeon, under stones on seashore. ’	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD6FFF5D5DA607FEE35F8F9.taxon	distribution	Distribution. Korea.	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD6FFF5D5DA607FEE35F8F9.taxon	discussion	Remarks. This species is similar to T. marinus, but can be distinguished by the shape of antennomeres. Antennomeres 1 – 11 are elongate in Thinobius koreanus sp. n. (Fig. 4 A), but 4 – 8 transverse in T. marinus. The new species is also similar to T. frizzelli from Canada and USA but can be distinguished by the shape of antennomeres. Antennomeres 4 – 10 are quadrate and 4 – 6 are almost same length in the new species (Fig. 4 A), but 4 – 10 rounded form and 5 larger than 4 or 6 in T. frizzelli (Kincaid, 1961). All specimens were collected in the southern part of the Korean peninsula.	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD0FFF6D5DA6166ED50FE96.taxon	description	(Figs. 1 C, 6 – 7)	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD0FFF6D5DA6166ED50FE96.taxon	description	Redescription. Length 1.02 – 1.32 mm. Body flattened, surface pubescent; light brown. Head. Slightly elongate and slightly narrower than pronotum; widest at about half of head; arrow shaped; clypeus slightly trapezoidal, anterolateral margin rounded; eye absent; temple well developed, gular suture fused and broadly divergent at anterior and posterior part; neck well developed, seta absent and reticulate sculptures distinct (Figs. 6 B – C); antenna slender, moderately elongate and pubescent, antennomere 1 longest, 1 – 2 and 9 – 11 elongate, 3 – 8 slightly elongate (Fig. 6 A). Mouthparts. Labrum transverse, lateral margin rounded, anterior margin slightly emarginate; epipharynx slightly developed; mandible triangle, apex bifid, with one blunt subapical tooth, prostheca and molar tooth well developed; maxillary palpus with four palpomeres, palpomere 1 transverse and small, 2 slightly dilated to apex; 3 largest, 2 – 3 with many setae; apical margin of 3 more than 2.0 times as wide as basal part of palpomere 4; labial palpus with three elongate palpomeres, palpomere 1 with one long and one short setae and two pores, 2 with one long seta and one pore, 3 with few short sensilla at apex; antero-middle part of hypopharynx with two blunted macro spines; mentum subquadrate with one pairs of long macrosetae and many setae. Thorax. Pronotum transverse, width 0.19 – 0.23 mm, surface pubescent with seven pairs of long macro setae; apex of prosternal process sharp and elevated; prosternum with one pair of long setae and few setae (Fig. 6 D); scutellum an inverted triangle, middle of anterior margin emarginate, posterior part weakly reticulate with few setae and diamond shaped impression, impression about 1.7 times as wide as long; elytra elongate and pubescent, posterior margin with membranous lobe; hind wing extremely reduced; mesoventrite with one pair of long setae; mesoventral process slightly extended and pointed at apex; metaventral carina absent; metaventrite reticulate, anterior part without setae (Fig. 6 E); all tibiae with one long macroseta in middle. Abdomen. Posterior margin of tergite VII with fringe of setae; posterior margin of sternite VIII convex. Secondary sexual characteristics. Male tergite IX with ventral struts, struts shorter than remainder, female tergite IX without ventral strut; male sternite IX present and female genital appendages present; female with one ring structure (Fig. 6 F). Genitalia. Aedeagus oval (Fig. 7 A) with serrated pattern in lateral aspect (Fig. 7 B); paramere long and slender; spermatheca as in Fig. 7 C.	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD0FFF6D5DA6166ED50FE96.taxon	materials_examined	Materials examined. KOREA: Gyeongbuk Prov.: 3 exx., Gyeongju-si, Gampo-eup, Najeong-beach, N 35 ° 46 ' 56 " E 129 ° 29 ' 27 ", 21 VI 2018, KJ Ahn, pebble beach in mid-tide, flotation; 16 exx., same data as former except for ‘ Najeong-ri, Najeong-beach, N 35 ° 46 ′ 56.79 ″ E 129 ° 29 ′ 30.95 ″ 0 m, 13 VII 2018, IS Yoo, JS Lee, JG Jung, 10 – 80 cm deep under gravels in mid-high tide, flotation’; 6 exx., Jeju Prov.: Seogwipo-si, Daejeong-eup, Sangmori, N 33 ° 11 ′ 55.6 ″ E 126 ° 16 ′ 30.7 ″ 2 m, 20 IX 2018, IS Yoo, JS Lee, WJ Jeong, 30 – 50 cm deep under fine gravels on gravel beach, flotation; 3 exx., same data as former except for ‘ N 33 ° 12 ' 22.21 " E 126 ° 17 ' 25.76 ", 21 IX 2018, IS Yoo, JS Lee, WJ Jeong, in gravels covered with seaweed, flotation’	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
231E878BFFD0FFF6D5DA6166ED50FE96.taxon	distribution	Distribution. Korea, Japan.	en	Lee, Jae-Seok, Yoo, In-Seong, Ahn, Kee-Jeong (2021): Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea. Zootaxa 4985 (2): 261-268, DOI: 10.11646/zootaxa.4985.2.9
