taxonID	type	description	language	source
2236042DFFAC407A6EE7FD7CFBE7FB10.taxon	type_taxon	TYPE SPECIES. — Pronodens siberlingi Koerner, 1940 by original designation.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFAC407A6EE7FD7CFBE7FB10.taxon	diagnosis	EMENDED DIAGNOSIS (revised from Taylor & Webb 1976; Métais & Vislobokova 2007; Tabrum & Métais 2007). — Larger size than Pseudoparablastomeryx Frick, 1937; dentition more selenodont than in Hendryomeryx Black, 1978, Pipestoneia Tabrum & Metais, 2007, and “ Leptomeryx mammifer (Cope, 1885); p 1 missing unlike in Leptomeryx Leidy, 1853 and Pipestoneia; shorter diastema between c and p 2 than in Leptomeryx and Pseudoparablastomeryx; lower jaw symphysis extends posteriorly to a point below anterior root of p 2 unlike in Leptomeryx and Pseudoparablastomeryx; dentary more robust and dorsoventrally deeper at the diastema than in Pseudoparablastomeryx; mental foramen smaller than in Pseudoparablastomeryx located more posteriorly, ventral or slightly anterior to p 2 unlike in Leptomeryx and Pseudoparablastomeryx; premolars relatively smaller than in Leptomeryx; reduced paraconid size in premolars compared to Leptomeryx; paraconid of p 4 not strongly inflected as in Pipestoneia.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFAC40776EAEFAD9FE5BF816.taxon	description	(Fig. 2; Appendix 1)	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFAC40776EAEFAD9FE5BF816.taxon	materials_examined	TYPE MATERIAL. — Holotype. United States of America • CM 726, partial skull with right and left P 2 - M 3 lacking parts of skull anterior to P 2 and posterior to the frontals. TYPE LOCALITY. — CM 2377 (KU-MT- 31, UMPC MV 6203, UWBM C 1709), United States of America, Montana, East of Drummond. MATERIAL. — From GLAC-PALLOC- 0001: GLAC 26988, partial left dentary bearing p 2 - m 3 and partial right dentary bearing p 3 - m 3.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFAC40776EAEFAD9FE5BF816.taxon	diagnosis	DIAGNOSIS. — Much smaller species than Pronodens silberlingi; dentary shallower than in P. silberlingi; p 2 less mesiodistally elongated than in P. silberlingi; labial cusps of m 3 (protoconid, hypoconid, and hypoconulid) more buccally expanded than in P. silberlingi; protocone of P 3 smaller and more buccolingually elongated than in P. silberlingi.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFAC40776EAEFAD9FE5BF816.taxon	discussion	STRATIGRAPHIC AND GEOGRAPHIC RANGE. — Lower unit of the Cabbage Patch beds (Renova Formation, early Arikareean) and upper member of the Kishenehn Formation, western Montana, United States of America (early Arikareean).	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFAC40776EAEFAD9FE5BF816.taxon	description	DESCRIPTION The Pronodens material from the Kishenehn Formation is composed of partial left and right dentaries. The left dentary preserves a little damaged or worn dentition including p 2 - m 3, whereas the right dentary includes a sheared and worn dentition including p 3 - m 3. The left jaw is only the second specimen including the entire cheektooth row ever discovered for the species after CM 8938, the most complete lower jaw known for the species. It is also one of the bestpreserved dentition, albeit worn, available for description for P. transmontanus (Fig. 2). The dentaries of GLAC 26988 are similar to those of Pronodens transmontanus in their robustness of the anterior portion of the jaw (Fig. 3 A; Table 1). In the posterior portion of the molar row, the dentary of the Kishenehn Formation material is deeper than in the specimens from the Cabbage Patch beds, more closely resembling MOR 951. Overall, both the Kishenehn Formation specimen and Cabbage Patch material are dorsoventrally shallower than the dentary of P. silberlingi (Fig. 3 A). Both the left and right jaws show evidence for the absence of p 1 unlike in Leptomeryx, but like all known Pronodens material. The mental foramen can be observed on the left side (Fig. 1 C); its position far anterior to the anterior edge of p 2 is similar to that observed in three known specimens of P. transmontanus, UMPC 3030, UMPC 1782, and UWBM 97460. Two other specimens, an edentulous jaw assigned to P. transmontanus (KUVP 20724) as well as CM 8938, show a more posteriorly located foramen, immediately ventral to the anterior edge of p 2, as in the type of P. silberlingi, a feature previously considered a characteristic of the genus (Taylor & Webb 1976). The postsymphysis is posterior to the mental foramen in the Kishenehn Formation material (Fig. 1 C), unlike in CM 8938 and the holotype of P. silberlingi, YPM 13952. In both GLAC 26988 and YPM 13952, the postsymphysis terminates posteriorly halfway through p 2; in CM 8938, it is fully anterior to p 2, terminating posteriorly at the anterior edge of the tooth (Fig. 1 C). The dentition of GLAC 26988 is best preserved in the left jaw (Appendix 1). The m 1 and m 2 of the right jaw are damaged and p 2 is missing (Fig. 1 A); only the roots of the tooth are preserved. The size of the p 2 of GLAC 26988 falls within the range of measurements observed in Pronodens transmontanus from the Cabbage Patch beds (the type area) for both length and width (Figs 3 B; 4 A; Table 2). The p 2 of P. silberlingi is much more mesiodistally elongated. The shape of the p 2 in the Kishenehn Formation specimen is very similar to that observed in the Cabbage Patch fossils (Fig. 1 E). The protoconid and entoconid are more robust than in UWBM 97460, but this character varies across specimens in the sample from Cabbage Patch (see UMPC 3030, CM 8938, and UWBM 97460 in particular; Fig. 4 A). The length of the p 3 of P. transmontanus from Cabbage Patch overlaps the length of the p 3 of the Kishenehn Formation material, but the buccolingual width of the latter is larger, more similar in fact to P. silberlingi (Figs 3 B; 4 B; Table 2). In both the left and right jaws of GLAC 26988, the portion of the tooth anterior to the paraconid is heavily worn, much more so than the posterior portion of the tooth and cannot be described in detail (Fig. 1 A, C, E). The paraconid itself is larger in GLAC 26988 than in KUVP 18452, KUVP 18668, and CM 8938. The same observation can be made of the protoconid, which is broader buccolingually in the Kishenehn Formation material than in the Cabbage Patch specimens. The ridge extending lingually from the protoconid is longer and narrower in the Cabbage Patch material than in the Kishenehn Formation specimen where it terminates more anteriorly (Fig. 1 E). The anatomy of the posterior-most portion of the tooth, including the entoconid and hypoconid, varies greatly with wear across specimens within the Cabbage Patch sample; the Kishenehn Formation specimen is most similar to KUVP 18452 and CM 8938, although the ridge extending lingually from the entoconid is more lingually inflated in the Kishenehn Formation specimen than in the Cabbage Patch fossils. The length of p 4 in the Kishenehn Formation specimen overlaps with the range of sizes observed in P. transmontanus from the Cabbage Patch beds (Fig. 3 B; Table 2). GLAC 26988 is comparable in width to the largest measured specimen of P. transmontanus from Cabbage Patch as well as the type specimen of P. silberlingi, but it is proportionately broader (Fig. 4 C). The anterior-most portion of the p 4 of the Kishenehn Formation specimen is similar to that of UWBM 97460, the only well preserved p 4 of P. transmontanus from Cabbage Patch, when accounting for wear. The more worn KUVP 18668 shows a similar paraconid morphology as the Kishenehn Formation specimen as well. The shape of the metaconid is complex, but once again, when accounting for wear, is essentially identical to that of UWBM 97460 and CM 8938 (Fig. 1 E). The advanced wear stage of the Kishenehn Formation specimen leads to an uninterrupted connection between the metaconid and entoconid (Fig. 1 E). The shape of the entoconid is simple, once more as a consequence of wear. It is larger, rounder, and more lingually inflated than in UWBM 97460 or CM 8938. There is, however, little difference in the morphology of the hypoconid between GLAC 26988 and UWBM 97460 when accounting for wear. The m 1 of GLAC 26988 is more heavily worn than in UWBM 97460, UMPC 1482, UMPC 3030, and CM 8938 (Fig. 1 E). The Kishenehn Formation specimen overlaps in size with the Cabbage Patch fossils (Fig. 3 C; Table 3) although it is broader for a given length than the specimens from the type area (Fig. 4 D). All these specimens are much smaller than the type of Pronodens silberlingi. GLAC 26988 is heavily worn making the morphologies of the metaconid and entoconid difficult to compare between the Kishenehn Formation specimen and the Cabbage Patch fossils, but the overall shapes of these lingual cusps are consistent across all specimens. The protoconid and hypoconid are more buccally extended in the Glacier National Park specimen (Fig. 1 E), leading to the greater buccolingual width of the tooth in GLAC 26988 compared to the Cabbage Patch fossils. A similar pattern can be observed in the m 2 with an overlap in size of the Kishenehn Formation specimen with the Cabbage Patch fossils (Fig. 3 C; Table 3). The Kishenehn Formation specimen is once again slightly wider for a given length than the Cabbage Patch sample (Fig. 4 E), but they are very similar, and even resemble a specimen of Pronodens from the Six Mile Creek Formation, MOR 951. The type of P. silberlingi is larger than any other specimen of the genus measured. The shape of the metaconid of GLAC 26988 is similar to the one in CM 8938; the shape of the entoconid is similar to that observed in UMPC 1482. The protoconid and hypoconid are buccally expanded in the Kishenehn Formation specimen compared to the Cabbage Patch sample. There are little to no differences in m 3 size between the Glacier specimen and the sample of fossils from the Cabbage Patch beds (Fig. 3 C; Table 3). Unlike in m 1 and m 2, the anterior lophid of the m 3 of GLAC 26988 is buccolingually narrower than that observed in the sample of P. transmontanus from Cabbage Patch or MOR 951 (Fig. 4 F). The buccal cusps (metaconid, entoconid, and entoconulid) of GLAC 26988 are most similar to those observed in CM 8938 and UMPC 1482 or MOR 951; they are large and round with tapered mesial and distal ends. In the Kishenehn Formation specimen, the lingual surface of the entoconid is not as rounded as in CM 8938 and KUVP 18838. Instead, it resembles more closely the morphology observed in UWBM 97494 with a lingually displaced cusp point. The entoconulid is teardrop shaped with the narrow end at the mesial end of the lophid contacting the distal end of the entoconid as in KUVP 18838 and CM 8938; the shapes of the entoconid and entoconulid in UWBM 97494 differ from those found in the Kishenehn Formation fossil as well as other P. transmontanus specimens. The entoconulid of MOR 951 is more symmetrical than in P. transmontanus from Cabbage Patch or the Kishenehn Formation specimen. The protoconid is not as buccally expanded in the Kishenehn Formation specimen as they are in the Cabbage Patch sample at similar wear stages; this is also true of the other labial cusps, the hypoconid and the hypoconulid (Fig. 1 E). However, these cusps are more buccally expanded than in P. silberlingi or MOR 951. Both the hypoconid and the hypoconulid are more mesiodistally expanded in GLAC 26988 than in the Cabbage Patch specimens, leading to more robust bulbous cusps.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFAC40776EAEFAD9FE5BF816.taxon	discussion	REMARKS The Kishenehn Formation specimen is very similar in size to the sample of Pronodens transmontanus from the type area, the Cabbage Patch beds (Tables 1 - 3). None of the measurements of the Kishenehn Formation specimen fall outside of the range of measurements observed in P. transmontanus ± 10 %; only two measurements of the Kishenehn Formation specimen (Ddm 2 and p 3 W) are more than 5 % larger than the measurements observed in the Cabbage Patch sample; this despite the small size of our sample (two and five specimens respectively) that is unlikely to capture the range of size variation in P. transmontanus. In contrast, P. silberlingi is much larger, with the type specimen (YPM 13952) more than 5 % larger than P. transmontanus from Cabbage Patch in 83 % of measurements taken (10 out of 12). YPM 13952 is also more than 10 % larger than the Kishenehn Formation specimen in nine measurements (out of 12) available for both specimens. On average, the teeth of YPM 13952 are 14.96 % larger than those of the Kishenehn Formation specimen. Differences between the Kishenehn Formation specimen and MOR 951 are reduced; however, the two specimens differ by 11.9 and 20.7 % for two dental measurements (m 2 L, m 3 Wpost). The morphology of the teeth in the Kishenehn Formation specimen more closely resembles P. transmontanus than P. silberlingi or Pronodens sp. However, the dentition of the type specimen of P. silberlingi is worn beyond the possibility of tooth crown morphological comparisons. We conservatively assign the material from Kishenehn Formation to P. transmontanus.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFA140776EE0FCBFFC6DFC0D.taxon	type_taxon	TYPE SPECIES. — Miohippus annectens Marsh, 1874 by original designation.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFA640706C0BF998FCCDF90E.taxon	type_taxon	TYPE SPECIES. — Paciculus insolitus Cope, 1879 by original designation.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFA640736C32F8DBFEB1F816.taxon	description	(Fig. 6; Appendix 2)	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFA640736C32F8DBFEB1F816.taxon	materials_examined	MATERIAL EXAMINED. — From CM Locality 5084: CM 87801, partial right dentary with incisor and m 1 - m 2.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFA640736C32F8DBFEB1F816.taxon	discussion	STRATIGRAPHIC AND GEOGRAPHIC RANGE. — Fort Logan Formation, upper member of the Kishenehn Formation, and middle as well as upper units of the Cabbage Patch beds (Renova Formation), western Montana, United States of America (early Arikareean). From Peterson Creek Local Fauna, eastern Idaho, United States of America (early Arikareean).	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFA640736C32F8DBFEB1F816.taxon	description	DESCRIPTION CM 87801 is a large specimen of Paciculus (Fig. 6; Appendix 1). The m 1 is as long as in P. montanus, P. walshi (Lindsay et al. 2016), P. nebraskensis (Alker 1969), and the sample of Paciculus from the Cabbage Patch beds identified as P. cf. P. montanus (Table 6; Fig. 7 A). It is much longer than any other species of the genus. A similar pattern can be observed in the m 2 (Table 6; Fig. 7 B); the Kishenehn Formation specimen is longer than any other species of the genus with the exceptions of P. nebraskensis and P. montanus. The widths of the teeth also place CM 87801 among the very largest Paciculus specimens. The shape of m 1 is nearly identical to P. montanus and the Cabbage Patch sample as well as one extreme of the shapes found in P. insolitus (Cope 1879; Fig. 8). The shape of the m 2 overlaps the range observed in both P. nebraskensis and P. montanus (Fig. 8). The m 1 of the Kishenehn Formation specimen (Fig. 6) shares a number of features with P. montanus that also differentiate it from P. walshi, including: 1) the presence of a distinct anteroconid; 2) a protoconid arm that is short and does not reach the anterior cingulum; 3) a short protolophid II, which, like half of the specimens of P. montanus observed by Lindsay et al. (2016), does not join the metaconid; and 4) a robust posterior arm of the protoconid that does not bifurcate, as in most specimens of P. montanus analyzed by Lindsay et al. (2016). Additionally, as in P. montanus, the m 2 of the Kishenehn Formation specimen bears a long protolophid II. Unlike in P. walshi or P. cedrus Korth, 2014, but like in P. montanus, the anterior lingual cingulum of the m 1 of CM 87801 extends from the anteroconid to the metaconid. The teeth of P. nebraskensis differ from those of CM 87801 by a shorter mesolophid of m 1, a shorter protolophid II on m 2, and the presence of an anterior cingulum of m 2 (Korth 1992). CM 87801 further differs from smaller species of Paciculus by several discrete morphological characters. The m 1 of P. gloveri MacDonald, 1970 has a smaller protolophid I as well as a narrower, buccolingually straighter mesostylid than the Kishenehn Formation specimen. Additionally, P. gloveri bears a posterior spur on the metaconid of m 2, has an isolated mesostylid, and has a very short protolophid II (Williams & Storer 1998), all characters not observed in the Kishenehn Formation fossil. P. copiosus (Korth 2010) bears an anteroconid on m 2 unlike CM 87801. Unlike the Kishenehn Formation specimen, the m 1 of P. dakotensis Korth, 2010 lacks a protolophid II and the protolophid II of the m 2 does not reach the lingual edge of the tooth (Korth 2010). Additionally, the m 2 of P. dakotensis (Korth 2010) bears an anterior cingulum extending lingually anterior to the metaconid, which is absent from the Kishenehn Formation fossil. Paciculus mcgregori (MacDonald, 1970) and the Kishenehn Formation fossil can be distinguished by the incomplete protolophid II of the former and very different anterior portions of m 1 (Martin 1980). The m 2 of P. cedrus, unlike the fossil from the Kishenehn Formation bears a short protolophid II disconnected from an isolated mesostylid as well as a distinct anterior cingulum (Korth 2014). The anterior portion of m 1 of CM 87801 also differs from that observed in the type specimen (SDSM 56510). The Kishenehn Formation specimen, unlike P. insolitus, but like P. walshi, has a short ectolophid on m 1 (Lindsay et al. 2016). Paciculus insolitus and CM 87801 can be further distinguished by differences in the shape of the mesostylid, the pattern of branching of the mesolophid and entolophid from the posterior mure, the size of the metaconid, and the morphology of the protoconid. The posterior cingula of the m 1 and m 2 of P. woodi are more mesiodistally oriented than in the Kishenehn Formation specimen, in which they are much more bucco-lingually oriented. In P. woodi (MacDonald 1963), the anterior mure is interrupted and therefore does not contact the protoconid in the m 1; this is also the case in P. cedrus and P. insolitus, but not in CM 87801. The anterior cingulum of P. woodi lacks the long labial extension seen in CM 87801.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
2236042DFFA640736C32F8DBFEB1F816.taxon	discussion	REMARKS The size and morphology of CM 87801 support it being assigned to Paciculus montanus (Table 6). The holotype of the species is an upper dentition (Black 1961), so it is not currently possible to directly compare the material from the Kishenehn Formation with the type area specimens, however, there is no identifiable difference between CM 87801 and the known lower dentition of P. montanus. The material from the Cabbage Patch beds (Calede 2020) cannot be assigned with confidence to the species based on the rarity of the material and its worn state. However, the size of the lower teeth as well as the morphology of the upper teeth recovered are consistent with P. montanus (Rasmussen 1977; this paper). Future discoveries of dental remains from the Cabbage Patch beds are expected to confirm the presence of the species in this area of Montana in addition to the type area, the Peterson Creek Local Fauna of Idaho and, now, the Kishenehn Formation.	en	Calede, Jonathan J., Constenius, Kurt N., Famoso, Nicholas A., Kehl, Winifred A. (2024): Discovery of Oligocene-aged mammals in Glacier National Park (Kishenehn Formation), Montana. Geodiversitas 46 (9): 367-386, DOI: 10.5252/geodiversitas2024v46a9, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a9.pdf
