identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
21518782F826B609F0B2FB629DE38CA4.text	21518782F826B609F0B2FB629DE38CA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Papuamicrus Jałoszyński 2023	<div><p>Papuamicrus gen. n.</p><p>Type species: Papuamicrus globosus sp. n. (here designated).</p><p>Diagnosis. Papuamicrus differs from all remaining members of the ‘ Cephennomicrus group’ in three unique apomorphies: (1) prosternal process in ventral view about as long as broad with concave posterior margin, so short that entirely situated on level of anterior margins of procoxae, which, in natural position, are separated by anterior tip of mesoventral process; (2) mesoventral and anterior metaventral processes fused into massive and broad (only ~3 times as long as broad) composite process with lateral margins slightly diverging posteriorly in posterior half, parallel in median and subapical regions, and anteriorly forming tricuspidate tip with strongly and rapidly narrowed anteriormost process fitting into posterior concavity of prosternal process; and (3) aedeagus with diaphragm shifted onto basal area, perpendicular in relation to long axis of median lobe, with median lentiform sclerotization forming attachment site for internal median longitudinal apophyse or tendon of intrinsic diaphragm retractor muscle. Moreover, the following combination of synapomorphies, separately occurring in other genera, but not all together, differentiates Papuamicrus from other genera of Cephenniini: (1) antenna with rapidly broadened, compact dimerous club; (2) head with frontal glands; (3) maxillary palpomere 4 button-shaped; (4) prothorax lacking internal sclerotized ‘reservoirs’ or cavities; (5) pronotum and elytra lacking pits and foveae; (6) procoxal cavities broadly closed posteriorly by postcoxal process of prosternum fused with postcoxal process of hypomeron; (7) prothoracic hypomeron not divided by transverse carina; (8) metaventral intermetacoxal process slightly narrower than mesoventral process, distance between metacoxae subequal to 1/6 of metaventral width at posterior metaventral margin; and (9) posterior margins of mesoventral rests non-carinate.</p><p>Description. Body (Figs 1–3) suboval, strongly convex, virtually glabrous (but microscopic recumbent setae can be seen at high magnifications), pronotum and elytra lacking macrosetae.</p><p>Head capsule (Figs 4–9) short and broad, in intact beetles strongly tilted downwards, so that plane of frons is perpendicular to coronal plane of body; frons and vertex confluent, together transverse; supraantennal tubercles small and weakly elevated; frons with pair of minute frontal glands (Fig. 5; fg); eyes in male large, coarsely faceted, strongly convex and posteriorly emarginate, in female smaller and situated close to antennal fossae; gular plate (Fig. 0; gp) large, transverse, with sharply marked gular sutures (Fig. 9; gs) and strongly transverse reticulate microsculpture, lacking large, oval punctures; posterior tentorial pits indiscernible; submentum short and strongly transverse, vestigial. Mentum (Fig. 9; mn) rectangular, weakly transverse and slightly narrowing anterad, prelabium (Fig. 9; plb) with six pairs of small suckers arranged in two longitudinal rows, labial palps minute and broadly separated, inserted at sides of prelabium, with palpomere 1 largest, slightly longer than broad, palpomere 2 slightly shorter and distinctly narrower than 1, slightly elongate, palpomere 3 much narrower and much longer than 2, rodlike with narrowed apex, about 3 × as long as broad. Maxillae generalized, maxillary palp (Fig. 9; mxp) with minute palpomere 1, strongly elongate, slightly clavate palpomere 2, strongly broadened palpomere 3 which is less than twice as long as broad and has truncate apex, palpomere 4 broad and very short, button-like and densely setose. Mandibles (Fig. 10; md) symmetrical, subtriangular, short, each with small dorsal mandibulo-labral interlocking projection, setose prostheca absent. Labrum (Fig. 9; lbr) short and strongly transverse, with strongly rounded anterior margin, with membranous marginal velum and symmetrically distributed sparse dorsal setae.</p><p>Antennae (Fig.10)slender,composed of 11antennomeres,scape and pedicel distinctly broader than antennomeres 3–9, club dimerous, oval, abruptly delimited, more than twice as broad as antennomere 9. Antennomeres 1–9 sparsely setose, setae on club denser and longer than those on remaining portion of antenna.</p><p>Pronotum (Figs 1, 3) in dorsal view broadly subtrapezoidal with lateral margins narrowing both posteriorly and anteriorly; anterior pronotal corners not visible in strictly dorsal view, blunt; posterior corners obtuse-angled and blunt; pronotal base lacking pits; lateral pronotal carinae smooth, not serrate, sharply developed on entire length.</p><p>Prosternum (Fig. 9) with basisternal portion (Fig. 9; bst) much shorter than coxal region; prosternal process (Fig. 9; psp) in ventral view conspicuously short, not separating procoxae, about as long as broad and with concave posterior margin to receive anterior tip of mesoventral process, in lateral view only slightly elevated beyond ventral surface of procoxae; notosternal sutures (Fig. 9; nss) complete; procoxal cavities broadly closed by posterolateral lobes of prosternum that are fused with postcoxal lobes of hypomera; hypomeral ridges diffuse and poorly discernible; hypomera (Fig. 9; hy) broad and concave, lacking transverse carinae.</p><p>Mesoscutellar shield (Fig. 1) partly exposed between elytral bases, broadly subtriangular, asetose and lacking pits.</p><p>Elytra (Figs 1–3) oval, lacking humeral denticles and basal foveae; apices of elytra rounded together.</p><p>Mesoventrite (Figs 2, 11) with mesoventral intercoxal process (Fig. 11; msvp) fused with anterior metaventral process and together forming elongate, nearly flat and broad platform between and behind mesocoxal rests; lateral margins of process slightly diverging posteriorly in posterior half, parallel in median and anterior regions, and anteriorly mesoventral process tricuspidate, with pair of small lateral subtriangular projections and abruptly narrowed subtriangular anterior tip fitting to posterior emargination of prosternal process. Mesoventral + anterior metaventral process with broad elevated lateral margins forming carinae tapering posteriorly.</p><p>Hind wings absent.</p><p>Metaventrite (Figs 1, 11) short and strongly transverse, distinctly narrowing posteriorly; posterior margins of mesocoxal rests not carinate; lateral margins weakly rounded, posterior margin sinuate along each metacoxa, with inversely subtrapezoidal metaventral intermetacoxal process (Fig. 11; mtvp), as broad as 1/6 of metaventrite at posterior margin. Metanepisterna and metepimera narrow, partly exposed in intact beetles.</p><p>Legs (Figs 2–3) moderately long and slender; pro- and mesocoxae oval, metacoxae strongly transverse; all trochanters short and subtriangular; all femora distinctly clavate; tibiae broadening distad; tarsi moderately slender.</p><p>Abdominal sternites (Fig. 2) unmodified, first visible (III) slightly longer than each of IV–VII and subequal in width to last visible (VIII), abdomen subtriangular with broadly rounded apex, about as long as metaventrite.</p><p>Aedeagus (Figs 12–15) strongly elongate, with symmetrical median lobe and asymmetrical endophallus with broadly tubular structures, diaphragm present, circular and conspicuously small, situated entirely on basal surface of aedeagus and bearing at center large lentiform sclerotization internally connected with median longitudinal apophyse that provides attachment for diaphragm retractor muscle; parameres slender, with apical setae, parameral base with small lateral lobes.</p><p>Distribution and composition. Papuamicrus is represented by one species known to occur in eastern New Guinea.</p><p>Etymology. The name Papuamicrus combines the prefix Papua - derived from the type locality, with the stem - micrus, often used in generic names for members of the ' Cephennomicrus group' of genera. Gender masculine.</p><p>Remarks. Papuamicrus shows a set of unique apomorphies (listed in diagnosis) not known in any other member of the ‘ Cephennomicrus group’ of genera within Cephenniini . These are ventral thoracic and aedeagal characters, difficult to observe. However, even in dorsal view, Papuamicrus can be easily identified by lack of pits and foveae on the pronotum and elytra, lack of conspicuously long and numerous macrosetae, and oval, compact, abruptly delimited dimerous antennal clubs. Within genera of the ‘ Cephennomicrus group’ which have a dimerous antennal club, it is usually loosely assembled, and not compact, not ‘histerid-like’. Exceptions are only Clavomicrus and Trurlia, but antennal clubs in these genera are composed of antennomeres 10 and 11 fused together, so that the antenna is composed of 10 separate antennomeres, and not 11, as in Papuamicrus . There is at least one, Trurlia -like and yet undescribed genus (mentioned in Jałoszyński (2011b)) with the antennal structure similar to that of Papuamicrus, but this genus seems to differ from Trurlia only in unfused two distal antennomeres, and its distinct dorsal foveal system, strongly elongate prosternal process fully separating procoxae, mesoventral process lacking anterior tip, and posteriorly carinate mesocoxal rests are clearly different from character states found in Papuamicrus .</p><p>Genera of Cephenniini (including the still unnamed ‘genus X1’ known from a single female collected in Sulawesi (characterized in Jałoszyński 2011b)) can be identified using the following key (see remarks in Jałoszyński (2020) for methods of observing some important characters):</p></div>	https://treatment.plazi.org/id/21518782F826B609F0B2FB629DE38CA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2023): Papuamicrus gen. n., a new genus of Cephenniini from New Guinea (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5339 (5): 492-500, DOI: 10.11646/zootaxa.5339.5.6, URL: http://dx.doi.org/10.11646/zootaxa.5339.5.6
21518782F822B60AF0B2FEE69A8D8D8E.text	21518782F822B60AF0B2FEE69A8D8D8E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cephenniini Reitter 1882	<div><p>Updated key to extant genera of Cephenniini</p><p>1 All legs with ‘trochantellus’; Neotropical.................................................................. 2</p><p>- Legs lacking ‘trochantellus’............................................................................. 6</p><p>2 Prosternal process absent.............................................. Pseudocephennium Reitter [Neotropical]</p><p>- Prosternal process present, clearly separating procoxae....................................................... 3</p><p>3 Prosternal process in lateral view strongly bent or curved posterad, with apex directed posterad or posteroventrad......... 4</p><p>- Prosternal process in lateral view nearly straight, with apex directed ventrad...................................... 5</p><p>4 Abdomen modified, with conspicuously shortened second visible sternite, additionally hidden under dense fringe of long setae along posterior margin of first visible sternite; prosternal process in ventral view subtriangular, narrowing posterad to rounded apex; procoxal cavities closed posteriorly.............................. Monstrophennium Jałoszyński [Neotropical]</p><p>- Abdomen unmodified, first visible sternite lacking posterior fringe of long setae; prosternal process in ventral view nearly parallel-sided or rhomboidal with truncate or slightly emarginate apex; procoxal cavities open posteriorly................................................................................... Paracephennium O’Keefe [Neotropical]</p><p>5 Prosternal process in ventral view with deeply bifurcate posterior margin............. Furcodes Jałoszyński [Neotropical]</p><p>- Prosternal process in ventral view with truncate or slightly concave posterior margin, not bifurcate................................................................................................ Shyri Jałoszyński [Neotropical]</p><p>6 Pronotum with pair of longitudinal C-shaped furrows......................... Eutheimorphus Franz &amp; Löbl [Oriental]</p><p>- Pronotum lacking C-shaped furrows...................................................................... 7</p><p>7 Pronotum with pair of mediolateral glandular openings....................................................... 8</p><p>- Pronotum without mediolateral openings................................................................... 9</p><p>8 Antennae 11-segmented, with asymmetrical and loosely assembled dimerous club...... Trichokrater Jałoszyński [Oriental]</p><p>- Antennae 10-segmented, with oval and symmetrical club formed by two fused antennomeres...................................................................................................... Trurlia Jałoszyński [Oriental]</p><p>9 Antennae 10-segmented, with large club formed by two fused antennomeres.......... Clavomicrus Jałoszyński [Oriental]</p><p>- Antennae clearly 11-segmented......................................................................... 10</p><p>10 Prothoracic basisternal region about as long as procoxal rests or longer..................... Etelea Csiki [W Palaearctic]</p><p>- Prothoracic basisternal region clearly shorter than procoxal rests............................................... 11</p><p>11 Prosternal process absent.............................................................................. 12</p><p>- Prosternal process present............................................................................. 13</p><p>12 Antenna with indistinct trimerous club............................... Cephennium Müller &amp; Kunze [W Palaearctic]</p><p>- Antenna with distinct dimerous club................................. Nanophthalmus Motschulsky [W Palaearctic]</p><p>13 Pronotum lacking antebasal pits......................................................................... 14</p><p>- Pronotum with at least one pair of antebasal pits or foveae.................................................... 18</p><p>14 Antennae gradually thickened, lacking abruptly delimited club; prosternal process in ventral view extremely thin, lamellate, in lateral view narrowly subtriangular and strongly projecting ventrad beyond ventral margins of procoxae...................................................................................... Cephazteca Jałoszyński [Neotropical]</p><p>- Antennae with dimerous or trimerous clubs; prosternal process in ventral view subquadrate, subtriangular or subrectangular with rounded or truncate tip, in lateral view not strongly projecting ventrally..................................... 15</p><p>15 Antennae with trimerous clubs; prothorax with two pairs of internal ‘cavities’ (visible as translucent oval areas on sides in posterior half)........................................................................ genus X1 (Sulawesi)</p><p>- Antennae with clearly dimerous clubs; prothorax with at most one pair of internal ‘cavities’......................... 16</p><p>16 Prosternal process in ventral view about as long as broad; posterior margins of mesocoxal rests not carinate; aedeagus with internal lentiform sclerotization at base........................................ Papuamicrus gen. n. [New Guinea]</p><p>- Prosternal process clearly elongate; posterior margins of mesocoxal rests carinate; aedeagus lacking internal basal lentiform sclerotization........................................................................................ 17</p><p>17 Antennal club composed of two unilaterally expanded antennomeres; each prothoracic hypomeron partly divided into anterior and posterior region by transverse carina; notosternal sutures partly obliterated; elytra lacking macrosetae................................................................................... Cephennococcus Jałoszyński [Oriental]</p><p>- Antennal club composed of two symmetrical antennomeres; prothoracic hypomera undivided; notosternal sutures developed on entire length; elytra with numerous long macrosetae............................... Indomicrus Jałoszyński [Oriental]</p><p>18 Basal elytral foveae large and filled with dense setae........................................................ 19</p><p>- Basal elytral foveae, if discernible, asetose................................................................ 20</p><p>19 Prosternal process short, not projecting ventrally beyond procoxae and not bent................................................................................................... Hlavaciellus Jałoszyński [Oriental, Himalaya]</p><p>- Prosternal process long, projecting ventrally beyond procoxae and bent posteriorly at obtuse angle........................................................... Cephennodes Reitter [Afrotropical, Palaearctic, Oriental, Nearctic, Australian]</p><p>20 Prosternal and mesoventral processes inversely T-shaped in cross-section, so that mesal portions of pro- and mesocoxae are hidden under expanded ventral surface of each process...................................................... 21</p><p>- Prosternal and mesoventral processes subtriangular or trapezoidal in cross-section, mesal portions of pro- and mesocoxae visible in ventral view................................................................................ 22</p><p>21 Pronotum with sublateral carinae; elytra with subhumeral carinae................... Cephennula Jałoszyński [Oriental]</p><p>- Pronotum lacking sublateral carinae; elytra lacking subhumeral carinae.............. Lathomicrus Jałoszyński [Oriental]</p><p>22 Each elytron with humeral carina; pronotum with more than two pairs of small but deep lateral foveae.......................................................................................... Foveomicrus Jałoszyński [Oriental]</p><p>- Humeral carinae absent; pronotum with at most two pairs of lateral pits......................................... 23</p><p>23 Prothorax with posterolateral internal ‘cavities’ (in dried specimens visible as translucent areas in posterior half).............................................................................. Pomphopsilla Jałoszyński [Afrotropical]</p><p>- Prothorax lacking internal ‘cavities’............. Cephennomicrus Reitter [Afrotropical, Palaearctic, Oriental, Australian]</p></div>	https://treatment.plazi.org/id/21518782F822B60AF0B2FEE69A8D8D8E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2023): Papuamicrus gen. n., a new genus of Cephenniini from New Guinea (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5339 (5): 492-500, DOI: 10.11646/zootaxa.5339.5.6, URL: http://dx.doi.org/10.11646/zootaxa.5339.5.6
21518782F821B60AF0B2FD889DCB8BE6.text	21518782F821B60AF0B2FD889DCB8BE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Papuamicrus globosus Jałoszyński 2023	<div><p>Papuamicrus globosus sp. n.</p><p>(Figs 1–15)</p><p>Type material. Holotype: PAPUA NEW GUINEA (Morobe Province): ♁, three labels: “ N. Guinea / Bíró 1899” [yellowish, printed], “Sattelberg / Huon-Golf.” [yellowish, printed], “ PAPUAMICRUS / globosus m. / P. Jałoszyński, 2023 / HOLOTYPUS” (MHNG) . Paratypes: 1 ♁, 1 ♀, “PAPUA NG: Morobe / above Wau / 1450m, 21.V.1992 / G. Cuccodoro #5A” [white, printed], and yellow “PARATYPUS” label (MHNG, cPJ) .</p><p>Diagnosis. As for genus; aedeagus broadest near proximal third, apex broadly subtriangular and rounded, endophallus with asymmetrical broad tube in distal 2/3 flanked by weakly sclerotized longitudinal structures, in subapical region with pair of lateral elongate and poorly sclerotized components flanking distal C-shaped portion of flagellum.</p><p>Description. Body of male (Figs 1–3) stout, strongly convex, light to moderately dark reddish brown, setae barely discernible; BL 0.83–0.86 mm.</p><p>Head (Figs 4–9) broadest at large, strongly convex eyes, HL 0.10–0.13 mm, HW 0.23–0.25 mm; frons and vertex confluent and weakly convex, virtually impunctate and sparsely covered with short recumbent setae. Frontal glands (Fig. 5) small and distinct. Each eye composed of 24 coarse facets. Antennae (Fig. 10) slender, AnL 0.33 mm; scape and pedicel elongate, antennomere 3 indistinctly elongate, 4–9 each distinctly elongate (9 longest, more than twice as long as broad), 10 and 11 each about as long as broad, 11 subconical with rounded apex.</p><p>Pronotum (Figs 1, 3) broadest near middle; PL 0.25–0.26 mm, PW 0.43–0.44 mm. Anterior margin in dorsal view broadly rounded; anterior corners (visible in anterodorsal view) subtriangular and blunt, weakly obtuse-angled, lateral margins strongly rounded in anterior half and weakly rounded in posterior half, where they are distinctly convergent posteriorly, smooth, not serrate; posterior corners strongly obtuse-angled and blunt; base indistinctly biarcuate and with short and indistinct shallow emargination in front of mesoscutellar shield. Disc virtually impunctate, covered sparsely with microscopic recumbent setae.</p><p>Elytra (Figs 1–2) together oval, broadest near anterior third; EL 0.48 mm, EW 0.45 mm, EI 1.06; humeral calli indistinct, not delimited by carinae or step-wise impressions. Punctures and setae on elytra similar to those on pronotum. Hind wings absent.</p><p>Legs moderately long and slender; unmodified.</p><p>Aedeagus (Figs 12–15) conspicuously elongate; AeL 0.38; median lobe in ventral view cigar-shaped, broadest in sub-basal third, in subapical area slightly but rapidly narrowing toward subtriangular apex of dorsal wall of aedeagus, ventrally ostium closed by membranous folds; endophallus with asymmetrical broad tubular sclerotized structure and with several weakly sclerotized structures flanking broad flagellum; each paramere long and slender, with three apical setae.</p><p>Female. Similar to male but with clearly smaller eyes, each composed of nine facets. BL 0.85 mm; HL 0.11 mm, HW 0.24 mm, AnL 0.30 mm; PL 0.26 mm, PW 0.43 mm; EL 0.48 mm, EW 0.45 mm, EI 1.06.</p><p>Distribution. Papua New Guinea.</p><p>Etymology. The Latin adjective globosus refers to the subglobose body form.</p><p>Remarks. The specimen designated here as the holotype was collected by the famous Hungarian zoologist Lajos Bíró, who explored Papua New Guinea at the end of the 20th century. Sattelberg, the type locality, is a village on the Huon Peninsula, eastern Papua New Guinea.</p></div>	https://treatment.plazi.org/id/21518782F821B60AF0B2FD889DCB8BE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2023): Papuamicrus gen. n., a new genus of Cephenniini from New Guinea (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5339 (5): 492-500, DOI: 10.11646/zootaxa.5339.5.6, URL: http://dx.doi.org/10.11646/zootaxa.5339.5.6
